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Presence of the genus Cuon in upper Pleistocene and initial Holocene sites of the Iberian Peninsula: new remains identified in archaeological contexts of the Mediterranean region
Authors:Manuel Pérez Ripoll  Juan V. Morales Pérez  Alfred Sanchis Serra  J. Emili Aura Tortosa  Inocencio Sarrión Montañana
Affiliation:1. Departament de Prehistòria i Arqueologia, Facultat de Geografia i Història, Universitat de València, Blasco Ibáñez 28, 46010 València, Spain;2. Museu de Prehistòria de València, SIP (Servei d''Investigació Prehistòrica), Diputació de València, Corona 36, 46003 València, Spain;3. Sociedades Cazadoras–Recolectoras Prehistóricas Research Group, Universidad de La Laguna, Tenerife, Spain;1. Sobolev Institute of Geology & Mineralogy, Siberian Branch of the Russian Academy of Sciences, Novosibirsk, 630090, Russia;2. Laboratory of Mesozoic and Cenozoic Continental Ecosystems, Tomsk State University, Tomsk, 634050, Russia;3. Institute of Plant & Animal Ecology, Urals Branch of the Russian Academy of Sciences, Yekaterinburg, 620144, Russia;4. Institute of Archaeology & Ethnography, Siberian Branch of the Russian Academy of Sciences, Novosibirsk, 630090, Russia;5. Mining Institute, Urals Branch Russian Academy of Sciences, Perm, 614007, Russia;6. Arizona AMS Laboratory, University of Arizona, Tucson, AZ, 85721-0081, USA;1. Institut Català, de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA-ICP, c/ Columnes s/n, Campus de la UAB, 08193, Cerdanyola del Vallès, Barcelona, Spain;2. Departamento de Prehistoria, Historia Antigua y Arqueología, Universidad Complutense de Madrid, Facultad de Geografía e Historia, C/ Prof. Aranguren s/n, 28040, Madrid, Spain;3. Institut Català de Paleoecologia Humana i Evolució Social (IPHES-CERCA), Zona Educacional 4, Campus Sescelades URV (Edifici W3), Tarragona, 43007, Spain;4. Instituto de Evolución en África (IDEA, Madrid), C/ Covarrubias 36, 28010, Madrid, Spain;5. Université Mohamed Premier, Faculté des Sciences, Département de Géologie, BV Mohammed VI, Hay Al Qods, 60000, Oujda, Morocco;6. Dipartimento di Scienze della Terra, Paleo[Fab]Lab, Università degli Studi di Firenze, Via G. La Pira 4, 50121, Firenze, Italy;7. Institut National des Sciences de l’Archéologie et du Patrimoine (INSAP), Av. Allal El-Fassi, Hay Riad, 6828, Rabat, Morocco;8. Universitat Rovira i Virgili, Departament d’Història i Història de l’Art, Avinguda de Catalunya 35, 43002, Tarragona, Spain;9. Unidad Asociada al CSIC. Departamento de Paleobiología. Museo Nacional de Ciencias Naturales, C/ José Gutiérrez Abascal, 2, 28006, Madrid, Spain;10. Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, SE-104 05, Stockholm, Sweden;11. UMR7194-HNHP (CNRS-MNHN-UPVD-Sorbonne Universités), Musée de l’Homme, Palais Chaillot, Place du Trocadéro 17, 75016, Paris, France;1. Institute of Evolution in Africa (IDEA), University of Alcalá de Henares, Covarrubias 36, 28010 Madrid, Spain;2. Real Complutense College at Harvard, 26 Trowbridge Street, Cambridge, MA 02138, USA;3. Department of Prehistory, Complutense University, 28040 Madrid, Spain;4. IPHES, Institut Catalá de Paleoecologia Humana i Evolució Social, Universitat Rovira I Virgili, 43007, Tarragona, Spain;5. Àrea de Prehistòria, Universitat Rovira i Virgili (URV), Avinguda de Catalunya 35, 43002, Tarragona, Spain;6. Unit associated to CSIC. Departamento de Paleobiologia. Museo Nacional de Ciencias Naturales, C/ José Gutierrez Abascal, 2, 28006, Madrid, Spain;7. GQP-CG, Grupo Quaternário e Pré-História do Centro de Geociências (uI&D 73 e FCT), Portugal;8. Museum National d''Histoire Naturelle, Institut de Paleontologie Humaine, 1 Rue René Panhard 75013 Paris, France;1. Dipartimento di Beni Culturali, Università di Bologna, Via degli Ariani 1, 48121 Ravenna, Italy;2. Università degli Studi di Ferrara, Dipartimento di Studi Umanistici, Sezione di Scienze Preistoriche e Antropologiche, Corso Ercole I d’Este, 32, Ferrara, Italy;3. Area de Prehistoria, Universitat Rovira i Virgili (URV), Avinguda de Catalunya 35, 43002 Tarragona, Spain;4. MuSe - Museo delle Scienze, Corso del Lavoro e della Scienza 3, IT 38123, Trento, Italy;5. Polo Museale del Lazio, Museo Nazionale Preistorico Etnografico “L. Pigorini”, Sezione di Bioarcheologia, Piazzale G. Marconi 14, I-00144 Rome, Italy;1. Institut International de Paléoprimatologie et de Paléontologie Humaine: Evolution et Paléoenvironnements, UMR CNRS 7262, Université de Poitiers, 6 rue Michel Brunet, 86022 Poitiers, France;2. Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand;3. Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand;4. Department of Mineral Resources, Rama VI Road, Bangkok 10400, Thailand
Abstract:The dhole (Cuon alpinus) has been documented on the Iberian Peninsula throughout the Middle and Upper Pleistocene in archaeological and palaeontological contexts. However, this is an uncommon species among the Iberian quaternary faunae as a whole. This paper focuses on the remains of the genus Cuon recovered in three different sites of the Iberian Mediterranean region and their morphological and osteometrical study. By comparing these with fourteen complete skeletons of Canis lupus, as well as with other remains of C. alpinus, we intend to highlight some anatomical and metrical differences between Cuon and Canis that, added to those already published, could be a functional base-line for the identification of this genus. The identification of this species is complex, especially in archaeological contexts, and in some cases the dhole has been identified as Canis or Canidae. Furthermore, this paper explores the chronology of the presence of the dhole in the Iberian Peninsula and the different contexts where these animals have been recovered from.
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