Permian corals of Timor (Rugosa and Tabulate): history of collection and study

The modern history of collection and study of corals in the Permian strata of Timor began in 1911,with a German expedition (J.Wanner, leader) and a Dutch expedition (H. Molengraaff, leader) to collect Permian and Triassic fossils in the colony of Netherlands Timor, and with a survey by the Swiss geologist F. Weber the same year in Portuguese Timor, the eastern portion of the island. Later expeditions led by Jonker (1916) and Brouwer (1937), both of the Netherlands, greatly increased already huge collections of fossils and additionally, understanding of the island's geology. Monographic studies of these coral collections by Gerth (1921), Koker (1924), Schindewolf (1942), Hehenwarter (1951) and Schouppé & Stacul (1955, 1959) have much enhanced the systematic value of these fossil corals, both Rugosa and Tabulata. Locality information and maps containing collecting localities are somewhat scattered (Wanner 1931, Burck 1923, Marez Oyens 1940, Van Bemmelen 1949), but are summarized here. A list of all valid species names(109 Rugosa, 25 Tabulata) is appended to this paper, with type localities and horizon. Serious problems of nomenclature are avoided in this paper by accepting genus names used in Hill (1981), modified by later systematic studies, such as that of Fedorowski (1986), but some unknown number of names in the list of species are to be synonymized, especially since 31 of them are based on a single specimen (e.g. Niermann 1975). The biostratigraphy of these faunas is uncertain, in great part due to the greatest number of corals having been collected from a tectonic mélange sequence in the Baun to Basleo structural region, and additionally because of the purchase of huge numbers of fossils from the indigenous people of Timor, with accompanying uncertainties regarding locality and horizon data. The coral fauna of Permian age from Timor needs serious restudy to insure its stratigraphic and palaeontologic value, but future study will require new field collection of specimens from the relatively complete stratigraphic sequences in the northern ‘Fatu’ belt of outcrops. The huge numbers of individuals of some coral species provide great opportunities for understanding population structure in these faunas.