Lower Cretaceous Anacoracidae? (Lamniformes: Neoselachii); vertebrae and associated dermal scales from Australia

Partially disarticulated shark vertebrae from the Lower Cretaceous Toolebuc Formation in central Queensland and the Bathurst Island Formation in the Northern Territory provide probable evidence of the Anacoracidae in Australia, and are possibly referable to Pseudocorax. Associated with large shark vertebrae from Canary Station, near Boulia, Queensland, are numerous placoid scales of four primary types which indicate a large pelagic shark. The Canary specimen is one of the few Mesozoic sharks known where scales have been found associated with vertebrae. Problems in referring the new shark material to the Anacoracidae and Pseudocorax are discussed. The significance of vertebral structure and scale morphology in Mesozoic shark evolution and ecology is examined. ‘Lamna daviesii’ Etheridge 1888 is considered a nomen dubium as vertebrae of this kind also occur in other genera in the Lamniformes, Orectolobiformes, and Carcharhinidae.