Ivane Tarkhnishvili (Tarchanoff): A Major Georgian Figure from the Russian Physiological School

This article is dedicated to one of the outstanding scientists of the nineteenth century: Ivane Tarkhnishvili (Tarchanoff), a Russian physiologist of Georgian origin who graduated from the St. Petersburg Medico-Surgical Academy and worked under the supervision of the founder of Russian physiology, Ivan Sechenov. Among his numerous contributions was the discovery of the skin galvanic reflex; however, Tarkhnishvili's most significant contribution was the discovery of the influence of X-rays on the central nervous system, animal behavior, the heart and circulation, and embryonic development. Indeed, these works have given rise to a new field in science (radiobiology).