Permian (Kungurian) Foraminifera from Western Australia described by Walter Parr in 1942: reassessment and additions

Haig, D.W., October 2017. Permian (Kungurian) Foraminifera from Western Australia described by Walter Parr in 1942: reassessment and additions. Alcheringa 42, 37–66. ISSN 0311-5518.

Exceptionally well-preserved siliceous agglutinated Foraminifera originally recorded by Walter Parr in 1942 are redescribed and illustrated by rendered multifocal reflected-light images. Significant new observations are made on wall texture and apertural morphology. The specimens are from the Quinnanie Shale and lower Wandagee Formation in the Merlinleigh Sub-basin of the Southern Carnarvon Basin, a marginal rift that splayed from the East Gondwana interior rift. During the Early Permian, a restricted shallow sea inundated the rift. The formations are part of sequence III of the Byro Group and belong within the Kungurian Stage (Cisuralian, Lower Permian). Of the 14 agglutinated species described by Parr, six are retained under their original names, viz., Hyperammina coleyi Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], H. rudis Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], Ammodiscus nitidus Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], A. wandageeensis Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], Tolypammina undulata Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar] and Reophax tricameratus Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]; one is transferred to a different species, viz., Thurammina texana Cushman &; Waters, 1928a Cushman, J.A. &; Waters, J.A., 1928a. Some Foraminifera from the Pennsylvanian and Permian of Texas. Contributions from the Cushman Laboratory for Foraminiferal Research 4, 31–55. [Google Scholar]; six are placed with other genera, viz., Thuramminoides pusilla (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Teichertina teicherti (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Sansabaina acicula (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Tolypammina? adhaerens (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Kunklerina subasper (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Trochamminopsis subobtusa (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]); and a species of Ammobaculites Cushman, 1910 Cushman, J.A., 1910. A monograph of the Foraminifera of the North Pacific Ocean. Part 1. Astrorhizidae and Lituolidae. United States National Museum, Bulletin 71(1), 134 pp. [Google Scholar] identified by Parr is now left in open nomenclature. From Parr's material, eight additional species are described: two new species, viz., Hyperammina parri sp. nov. and Gaudryinopsis raggatti sp. nov.; rare representatives of Aaptotoichus quinnaniensis Haig, 2003 Haig, D.W., 2003. Palaeobathymetric zonation of foraminifera from lower Permian shale deposits of a high-latitude southern interior sea. Marine Micropaleontology 49, 317–334. 10.1016/S0377-8398(03)00051-3[Crossref], [Web of Science ®] , [Google Scholar]; and very rare species of Lagenammina Rhumbler, 1911 Rhumbler, L., 1911. Die Foraminiferen (Thalamophoren) der Plankton-Expedition, Erster Teil, Die allgemeinen Organizationsverhaltnisse der Foraminiferen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, Kiel u. Leipzig, 3L.c. (1909), 1–331. [Google Scholar], Giraliarella Crespin, 1958 Crespin, I., 1958. Permian foraminifera of Australia. Bureau Mineral Resources, Geology and Geophysics, Bulletin 48, 1–207. [Google Scholar], Glomospira Rzehak, 1885 Rzehak, A., 1885. Bemerkungen über einige Foraminiferen der Oligocän Formation. Verhandlungen des Naturforschenden Vereins in Brünn 1884(23), 123–129. [Google Scholar], Hormosinella Shchedrina, 1969 Shchedrina, Z.G., 1969. O nekotorykh izmeneniyakh v sisteme semeystv Astrorhizidae i Reophacidae (Foraminifera). Voprosy Mikropaleontologii 11, 157–170. [Google Scholar], and Reophax Denys de Montfort, 1808 Denys de Montfort, P., 1808. Conchyliologie Systématique et Classification Méthodique des Coquilles, Volume 1. F. Schoell, Paris, 409. 10.5962/bhl.title.10571[Crossref] , [Google Scholar], all of which are left in open nomenclature. Hyperammina rudis is the type species of Hyperamminita Crespin, 1958 Crespin, I., 1958. Permian foraminifera of Australia. Bureau Mineral Resources, Geology and Geophysics, Bulletin 48, 1–207. [Google Scholar], a genus now considered a junior subjective synonym of Hyperammina Brady, 1878 Brady, H.B., 1878. On the reticularian and radiolarian Rhizopoda (Foraminifera and Polycystina) of the North Polar Expedition of 1875–76. Annals and Magazine of Natural History, ser. 1(6), 425–440. 10.1080/00222937808682361[Taylor &; Francis Online] , [Google Scholar]. Thuramminoides pusilla is considered a senior subjective synonym of T. sphaeroidalis Plummer, 1945 Plummer, H.J., 1945. Smaller Foraminifera in the Marble Falls, Smithwick, and Lower Strawn strata around the Llano Uplift in Texas. The University of Texas, Publication 4401, 209–271. [Google Scholar], the type species of Thuramminoides Plummer, 1945 Plummer, H.J., 1945. Smaller Foraminifera in the Marble Falls, Smithwick, and Lower Strawn strata around the Llano Uplift in Texas. The University of Texas, Publication 4401, 209–271. [Google Scholar]. Imagery is presented confirming that the simple cylindrical canals through the wall of Teichertia teicherti differ from the branching canals in Crithionina rotundata Cushman, 1910 Cushman, J.A., 1910. A monograph of the Foraminifera of the North Pacific Ocean. Part 1. Astrorhizidae and Lituolidae. United States National Museum, Bulletin 71(1), 134 pp. [Google Scholar], type species of Oryctoderma Loeblich &; Tappan, 1961 Loeblich, A.R. &; Tappan, H., 1961. Remarks on the systematics of the Sarkodina (Protozoa), renamed homonyms and new and validated genera. Proceedings of the Biological Society of Washington 74, 213–234. [Google Scholar]. The collection contains some of the earliest representatives of the revised family Verneuilinoididae Suleymanov, 1973 Suleymanov, I.S., 1973. Nekotorye voprosy sistematiki semeystva Verneuilinidae Cushman 1927 v svyazi s usloviyami obitaniya. Dokladari Uzbekiston SSR. Fanlar Akademiyasining, Tashkent 1973, 35–36. [Google Scholar], herein elevated from subfamily rank, and considered to include Pennsylvanian–Cisuralian representatives of Mooreinella Cushman &; Waters, 1928a Cushman, J.A. &; Waters, J.A., 1928a. Some Foraminifera from the Pennsylvanian and Permian of Texas. Contributions from the Cushman Laboratory for Foraminiferal Research 4, 31–55. [Google Scholar], Aaptotoichus Loeblich &; Tappan, 1982 Loeblich, A.R. &; Tappan, H., 1982. A revision of mid-Cretaceous textularian foraminifers from Texas. Journal of Micropalaeontology 1, 55–69. 10.1144/jm.1.1.55[Crossref] , [Google Scholar], Digitina Crespin &; Parr, 1941 Crespin, I. &; Parr, W.J., 1941. Arenaceous Foraminifera from the Permian rocks of New South Wales. Journal and Proceedings of the Royal Society of New South Wales 74, 300–311. [Google Scholar], Gaudryinopsis Podobina, 1975 Podobina, V.M., 1975. Foraminifery Verkhnego Mela i Paleogena zapadno-Sibirskoy nizmennosti, ikh znachenie dlya stratigrafii. Tomsk University Press, Tomsk, 264. [Google Scholar], Caronia Brönnimann, Whittaker &; Zaninetti, 1992 Brönnimann, P., Whittaker, J.E. &; Zaninetti, L., 1992. Brackish water foraminifera from mangrove sediments of southwestern Viti Levu, Fiji Island, Southwest Pacific. Revue de Paléobiologie 11, 13–65. [Google Scholar] (=Palustrella Brönnimann, Whittaker &; Zaninetti, 1992 Brönnimann, P., Whittaker, J.E. &; Zaninetti, L., 1992. Brackish water foraminifera from mangrove sediments of southwestern Viti Levu, Fiji Island, Southwest Pacific. Revue de Paléobiologie 11, 13–65. [Google Scholar]) and Verneuilinoides Loeblich &; Tappan, 1949 Loeblich, A.R. &; Tappan, H., 1949. New Kansas Lower Cretaceous Foraminifera. Journal of the Washington Academy of Sciences 39, 90–92. [Google Scholar].

David W. Haig [david.haig@uwa.edu.au] Centre for Energy Geoscience, School of Earth Sciences, University of Western Australia, 35 Stirling Highway, Crawley WA 6009, Australia.