Australian Middle Cambrian molluscs and their bearing on early molluscan evolution

Thirty ammonite taxa are recognised in the Wangarlu Mudstone of the Bathurst Island Group from the Cox Peninsula and Shoal Bay, Northern Territory, Australia. Included are a new heteromorph genus Notostreptites, type species N. exilis, and four additional new species and subspecies: Pseudhelicoceras gracilis, Labeceras (L.) tumidum, Labeceras (Appurdiceras) decorum, and Idiohamites dorsetensis laticostatus. The assemblage is best collectively correlated with the Mortoniceras (M.) inflatum Zone of the standard European Albian zonation but some of its members may represent the lower part of the Stoliczkaia dispar Zone. It is broadly correlative with faunas from the Eromanga Basin, which relate to an extensive eastern Australian Late Albian epicontinental sea, but is strikingly different in aspect. The Wangarlu Mudstone assemblage has relatively high diversity and an abundance of cosmopolitan heteromorph taxa well known from Europe and elsewhere, whereas Eromanga Basin assemblages are of relatively low diversity and dominated by Austral heteromorph genera known only from Australasia, southern Africa and Malagasy. The Austral character of Eromanga Basin assemblages is attributed to evolution in a restricted epicontinental sea environment and modest dispersal whereas the continental margin position of the Wangarlu Mudstone ensured an influx of pandemic elements drawn from the mid-Cretaceous world ocean.     

Thambetolepis delicata gen. et sp. nov., an enigmatic fossil from the Early Cambrian of South Australia

The acid insoluble sclerites of Thambetolepis delicata gen. et sp. nov. of late Tommotian or immediately post-Tommotian age from near Ardrossan, South Australia, are described. These sclerites, as well as those of Sachites Meshkova 1969 and Halkieria Poulsen 1967, are from multi-sclerite animals similar to Wiwaxia Walcott 1911 together with which they constitute the new class Thambetolepidea. Biological affinities of these fossils are discussed in the light of the complex internal structure of the sclerites. Possible assignments to the opisthobranch gastropods, the polyplacophorans, or the annelids are discussed but the taxonomic position of Thambetolepis and the Thambetolepidea is unresolved.     

Occurrence of the ammonite Goodhallites goodhalli (J. Sowerby) in the Eromanga Basin,Queensland: an index species for the late Albian (Cretaceous)

Mortoniceratid ammonites of the Eromanga Basin of Queensland, although uncommon, are reviewed and reassessed utilising all known collections. Representatives of this group are restricted to the Allaru Formation and almost all specimens are Goodhallites goodhalli, a well-known species from the English Gault. The Allaru Formation overlies the Toolebuc Formation, widely considered to be an essentially isochronous unit because of its unusual sedimentary and geochemical character. Using G. goodhalli, the middle and upper Allaru Formation can be directly correlated with the early late Albian orbignyi and auritus Subzones of the Mortoniceras inflatum Zone as recognized in the reference ammonite zonation embedded in the standard Cretaceous time scale. Overlapping ranges of G. goodhalli and Labeceras and Myloceras allow these common Austral heteromorph genera to be also confidently assigned a late Albian age in Australia, matching their biostratigraphic occurrence in South Africa.