The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914)—implications for the identification of Pseudisograptus manubriatus subspecies

VandenBerg, A.H.M. & Maletz, J., April 2016. The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914)—implications for the identification of Pseudisograptus manubriatus subspecies. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914) has been rediscovered in the collections of Museum Victoria and is recognized to belong to Pseudisograptus manubriatus harrisi as described by Cooper & Ni (1986). This taxon becomes a junior synonym of P. m. manubriatus. The selected neotype of Pseudisograptus manubriatus manubriatus (Hall, 1914) can be referred to Pseudisograptus manubriatus texanus Cooper & Ni, 1986. The proximal development in all known relief specimens of Pseudisograptus and their descendants indicates that a symmetrical development of the manubrium must be regarded as the rule, and the interpretation of a strong asymmetry in Pseudisograptus manubriatus texanus is rejected.

Alfons H.M. VandenBerg [avandenberg@museum.vic.gov.au, lanceolatus@hotmail.com], Geology, Museum Victoria, GPO Box 666, Vic 3001, Australia; Jörg Maletz [yorge@zedat.fu-berlin.de], Department of Earth Sciences, FU Berlin, Malteserstrasse 74-100, D-12249 Berlin, Germany.     

Conodonts and tabulate corals from the Upper Ordovician Angullong Formation of central New South Wales,Australia

Zhen, Y.Y., Wang, G.X. &; Percival, I.G., August 2016. Conodonts and tabulate corals from the Upper Ordovician Angullong Formation of central New South Wales, Australia. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

The Angullong Formation is the youngest Ordovician unit exposed in the Cliefden Caves area of central New South Wales. Its maximum age is constrained by a Styracograptus uncinatus graptolite Biozone fauna at the very top of the underlying Malongulli Formation, but the few fossils previously reported from higher in the Angullong Formation are either long-ranging or poorly known. From allochthonous limestone clasts in the middle part of the formation, we document a conodont fauna comprising Aphelognathus grandis, A. solidum, Aphelognathus sp., Aphelognathus? sp., Belodina confluens, Drepanoistodus suberectus, Panderodus gracilis, Panderodus sp., Phragmodus undatus, Pseudobelodina inclinata and Pseudobelodina? sp. aff. P. obtusa, which supports correlation with the Aphelognathus grandis Biozone (late Katian) of the North American Midcontinent succession. The species concepts of Aphelognathus and Pseudobelodina are reviewed in detail. Associated corals are exclusively tabulates, dominated by agetolitids, including Agetolites angullongensis sp. nov., Heliolites orientalis, Hemiagetolites breviseptatus, Hemiagetolites sp. cf. H. spinimarginatus, Navoites sp. cf. N. circumflexa, Plasmoporella bacilliforma, P. marginata, Quepora sp. cf. Q. calamus and Sarcinula sp. Affinities of the coral fauna from the Angullong Formation are closer to faunas from northern NSW and northern Queensland than to the locally recognized Fauna III of late Eastonian age in central NSW. We propose a subdivision of Fauna III to account for this difference, with the late Katian Fauna IIIB characterized by the incoming of agetolitid corals. The currently known distribution of representatives of this group with adequate age constraints suggests that agetolitids possibly originated in North China, subsequently migrating to Tarim, South China and adjacent peri-Gondwanan terranes while also spreading eastward to northern Gondwana, where they progressively moved through eastern Australia to reach the central NSW region by the early Bolindian.

Yong Yi Zhen* (yong-yi.zhen@industry.nsw.gov.au) and Ian G. Percival (ian.percival@industry.nsw.gov.au), Geological Survey of New South Wales, W.B. Clarke Geoscience Centre, 947–953 Londonderry Road, Londonderry, NSW 2753, Australia; Guangxu Wang (gxwang@nigpas.ac.cn), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road Nanjing 210008 PR China.     

New,early ostracods from the Ordovician (Tremadocian) of Iran: systematic,biogeographical and palaeoecological significance

New Tremadocian ostracod material from the Alborz Mountains of Iran confirms the early and widespread occurrence of the Ordovician genus Nanopsis, and the apparently simultaneous first appearance of ostracods in the fossil record at the level of the P. deltifer conodont biozone (485.5 Ma) from China to Argentina. Nanopsis pairidaeza sp. nov. adds to the pool of species diversity for the Early Ordovician, though documented Tremadocian ostracod generic diversity remains low, with only four genera. The presence of Early Ordovician ostracods in Alborz, their occurrence elsewhere in palaeocontinental Gondwana, Baltica and China coupled to their marked absence from the Tremadocian of Laurentia and Siberia, supports the notion of the earliest occurrence of ostracods centred on Gondwana/Baltica.     

Upper Ordovician Phacopida (Trilobita) from Tasmania

Eastonian trilobite faunas of the Gordon Group in Tasmania include the new species Ceraurinella oepiki, Erratencrinurus trippi, and Pliomerina trisulcata, as well as a reedocalymeninid probably allied to Sarrabesia Hammann & Leone, 1997. Ceraurinella and Erratencrinurus have not previously been reported from Australia, the former being predominantly Laurentian but also known from NE China, the Himalaya, and Vietnam, and the latter mostly Baltic/Laurentian. Peri-Gondwanan species of Ceraurinella appear to form a clade, within which Tasmanian and Indian (central Himalayan) taxa are closest relatives.     

Revision of two phragmodontid species (Conodonta) from the Darriwilian (Ordovician) of the Canning Basin in Western Australia and phylogeny of the Cyrtoniodontidae

Zhen, Y.Y. 9 July 2019. Revision of two phragmodontid species (Conodonta) from the Darriwilian (Ordovician) of the Canning Basin in Western Australia and phylogeny of the Cyrtoniodontidae. Alcheringa XX, xxx–xxx. ISSN 0311-5518

Based on re-examination of the material used in the original study from the subsurface Goldwyer and Nita Formations (middle Darriwilian, Middle Ordovician) of the Canning Basin, Western Australia, two phragmodontid species (Phragmodus polystrophos Watson and Ph. spicatus Watson) are revised as having a septimembrate apparatus including geniculate (Ph. polystrophos) or nongeniculate (Ph. spicatus) M, triform alate Sa, modified tertiopedate (Ph. polystrophos) or tripennate (Ph. spicatus) Sb, modified bipennate Sc, modified quadriramate Sd, carminate Pa and pastinate Pb elements. Characterized by a carminate Pa element in their respective species apparatuses, these two species demonstrate a close phylogenetic relationship with Phragmodus cognitus Stauffer from the Late Ordovician (Sandbian) of North America. These distinctive shared characters have allowed their accommodation within a new genus, Protophragmodus gen. nov., which represents an evolutionary lineage separated from species of Phragmodus Branson & Mehl (sensu stricto). In addition, it is postulated that the Cyrtoniodontidae might have originated in the early–middle Darriwilian from ‘Plectodina’ in shallow-water settings, with Phragmodus (sensu stricto), the most derived part of the family, perhaps directly evolving from Protophragmodus gen. nov. in the late Darriwilian and then becoming cosmopolitan, deeper-water dwellers in the Late Ordovician.

Y.Y. Zhen [yong-yi.zhen@planning.nsw.gov.au], Geological Survey of New South Wales, W.B. Clarke Geoscience Centre, 947–953 Londonderry Road, Londonderry, NSW 2753, Australia.     

Genyornis eggshell (Dromornithidae; Aves) from the Late Pleistocene of South Australia

Fossil eggshell fragments from a sand dune near Port Augusta are attributed to the extinct dromornithid, Genyornis newtoni Stirling & Zietz. Shell curvature measurements show that the eggs were larger than those of the Emu, Dromaius novaehollandiae. Radiocarbon dates indicate an age in excess of 40,680 BP. Holes pierced through some fragments are attributed to the action of predators.     

Ordovician stromatoporoids from Tasmania

The Hunter Siltstone near Grenfell, New South Wales, contains a rich Upper Devonian fish fauna including the sinolepid Grenfellaspis and the new antiarchs Bothriolepis grenfellensis sp. nov. and Remigolepis redcliffensis sp. nov. Bothriolepis grenfellensis sp. nov. is the first bothriolepid species described from N.S.W., and R. redcliffensis sp. nov. is the first species of Remigolepis described from Australia. Traditionally, the Hunter Siltstone was considered to be uppermost Famennian or earliest Carboniferous in age based on the presence of Grenfellaspis, and the related taxon Sinolepis, which is known from the Wutung and Sanmentan formations of southeastern China. However, available data indicates the Hunter Siltstone may be early Famennian in age. Ongoing work suggests that all Famennian Bothriolepis from N.S.W., including B. grenfellensis, possess a trifid preorbital recess, but differ in other aspects of headshield morphology. In North China, the Famennian Zhongning Formation contains six species of Remigolepis and a species of Sinolepis. However, R. redcliffensis does not show any similarity to these species beyond those of Remigolepis as a whole.     

Phylogenetic relationships of leptellinid brachiopods

The relationships among the diverse genera comprising the family Leptellinidae (Brachiopoda) are reviewed in the light of the revised edition of the Treatise on Invertebrate Palaeontology. Taxonomic work reassessed all the genera identified as Leptellinidae in the most current classification. Four genera were discarded, namely Bekkerella, Benignites, Leptastichidia and Nikitinamena. Cladistic analysis reveals the paraphyly of these genera; their abandonment leading to more morphologically coherent subdivisions of the family. Two subfamilies, Leptellininae and Palaeostrophomeninae, are emended and taxonomically restructured. The palaeogeographical history of the Leptellinidae is complicated. The Leptellinidae are first recorded in Baltica in the late Floian (Early Ordovician) and rapidly dispersed to circum-Iapetus palaeocontinents by the Dapingian, thence to most terranes composing Gondwana by the Darriwilian (Middle Ordovician).     

Tremadocian (Ordovician) graptolite diversification events in China

Study of the taxonomy, evolution and distribution of Tremadoc graptolites and their sequences in China and other representative sections around the world indicates that five graptolite diversity events can be recognised during the Tremadoc interval. The taxonomic validity of the suborder Graptodendroidina is advocated.     

Ichnofacies of the Stairway Sandstone fish-fossil beds (Middle Ordovician,Northern Territory,Australia)

The Stairway Sandstone is a 30–560 m thick succession of Middle Ordovician siliciclastic sedimentary rocks within the Amadeus Basin of central Australia, deposited in the epeiric Larapintine Sea of northern peri-Gondwana. The Stairway Sandstone is significant as one of only two known Gondwanan successions to yield articulated arandaspid (pteraspidomorph agnathan) fish. Herein we use the ichnology of the Stairway Sandstone to reveal insights into the shallow marine habitat of these early vertebrates, and compare it with that of other known pteraspidomorph-bearing localities from across Gondwana. The Stairway Sandstone contains a diverse Ordovician ichnofauna including 22 ichnotaxa of Arenicolites, Arthrophycus, Asterosoma, Cruziana, Didymaulichnus, Diplichnites, Diplocraterion, ?Gordia, Lockeia, Monocraterion, Monomorphichnus, Phycodes, Planolites, Rusophycus, Skolithos and Uchirites. These ichnofauna provide a well-preserved example of a typical Ordovician epeiric sea assemblage, recording the diverse ethologies of tracemakers in a very shallow marine environment of flashy sediment accumulation and regularly shifting sandy substrates. New conodont data refine the age of the Stairway Sandstone to the early Darriwilian, with ichnostratigraphic implications for the Cruziana rugosa group and Arthrophycus alleghaniensis.