Raw stone material supply for Upper Pleistocene settlements in Sierra de Atapuerca (Burgos,Spain): flint characterization using petrographic and geochemical techniques

Microscopy, mass spectrometry and X-ray diffraction methods were used to analyse 415 samples of natural and archaeological flint from Sierra de Atapuerca (Burgos, Spain) in order to define the different types from Neogene and Cretaceous formations in the study area, infer their genetic context and ascertain the supply sources used by hunter-gatherers who exploited this area in the Upper Pleistocene. A statistical classification model was also designed using linear discriminatory analysis and support vector machines which permitted the differentiation of the flint on an age basis.     

Significance of diadematid echinoid ossicles in micropalaeontological samples,Miocene–Pliocene of Indonesia

Fragments of diadematoid echinoids from the early and middle Miocene, and late Miocene–Pliocene, respectively, of Java, Kalimantan and Sulawesi, Indonesia, are identified as diadematid spp. indet. (radioles from all sites) and Centrostephanus sp. (an interambulacral plate; early Miocene, Java). The radioles are probably a mixture of Diadema ± Centrostephanus ± Echinothrix. This is the first report of identifiable fossil diadematoid remains from Indonesia and demonstrates that these echinoids, so common in modern reef environments, were present in the Neogene of the region. Even though classified in open nomenclature, Centrostephanus sp. nevertheless provides further evidence for the Cenozoic record of a genus in which the only nominal species are of Late Cretaceous and Holocene age.     

A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile

Hampe, O., Witzmann, F. & Asbach, P., 2014. A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile. Alcheringa 38, xxx–xxx. ISSN 0311–5518.

A pathology of the fossil baleen whale ‘Megaptera’ hubachi from the early Pliocene of Chile is described. It is a bony outgrowth on the left side of the supraoccipital, which is interpreted as a benign bone-forming tumour (osteoma). This diagnosis is based on X-ray imaging and CT scans of the abnormal bone, revealing a homogeneously dense internal structure with no evidence for lytic areas. The osteoma described here in ‘Megaptera’ hubachi is the first unequivocal evidence of a bone tumour in a cetacean, fossil or extant.

Oliver Hampe [oliver.hampe@mfn-berlin.de] and Florian Witzmann [florian.witzmann@mfn-berlin.de], Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, D-10115 Berlin, Germany; Patrick Asbach [patrick.asbach@charite.de], Institut für Radiologie, Charité—Universitätsmedizin Berlin, Charitéplatz 1, D-10117 Berlin, Germany. Received 28.8.2013, revised 7.11.2013, accepted 12.11.2013.     

Traditional and computed tomographic (CT) techniques link modern and Cenozoic fruits of Pleiogynium (Anacardiaceae) from Australia

Computed tomography (CT) and traditional techniques were used to study the internal morphology of modern fruits of Pleiogynium timoriense (DC.) Leenh., and phosphatized and silicified fruits that were considered to belong to the genus. The results demonstrate that phosphatized fruits from the Oligocene–Miocene Dunsinane site at Riversleigh and silicified fruits from the Oligocene Glencoe Site, which are both in Queensland, are referable to Pleiogynium. The Riversleigh material (Pleiogynium wannanii Rozefelds, Dettmann & Clifford sp. nov.) is very similar to extant P. timoriense (DC.) Leenh. Both possess locules that, in the vertical axial plane, are asymmetrically reniform and enclosed by a two-layered endocarp, the inner layer woody and resistant to decay, the outer of fibres and parenchyma, surrounded by a mesocarp of fibres and sclereids and a parenchymatous exocarp. The outer mesocarp is sculpted by meridionally arranged, elongate depressions (germination apertures) situated at the dorsal surface of each locule; erosion of the mesocarp and outer endocarp provides passage for emergence of the embryo’s radicle on germination. The silicified material from Glencoe (P. parvum Rozefelds, Dettmann & Clifford sp. nov.) is smaller in all its parts and has locules that are ovate in the vertical axial plane. Discrete, cap-like, opercula are not evident in the fruit wall of the fossil and extant fruits studied. The presence of Pleiogynium, and associated taxa, indicates the existence of rainforest communities at Glencoe, and mixed open forest/rainforest at Riversleigh during Oligocene–Miocene times. The multilocular, fossil fruit described as Pleiogynium mitchellii Collinson, Manchester and Wilde from the Eocene deposits at Messel Quarry, Germany, is not considered to belong in the genus, as evidence of elongate depressions on the dorsal surface of the fruit stones has not been demonstrated conclusively, and its internal morphology/anatomy differs significantly from that of Pleiogynium.     

Pliocene Mollusca (Bivalvia,Gastropoda) from the Sørsdal Formation,Marine Plain,Vestfold Hills,East Antarctica: taxonomy and implications for Antarctic Pliocene palaeoenvironments

Quilty, P.G., Darragh, T.A., Gallagher, S.J. & Harding, L.A. July 2016. Pliocene Mollusca (Bivalvia, Gastropoda) from the Sørsdal Formation, Marine Plain, Vestfold Hills, East Antarctica: taxonomy and implications for Antarctic Pliocene palaeoenvironments. Alcheringa 40, XXX–XXX. ISSN 0311-5518.

Pliocene shallow-water marine sediments at Marine Plain (centred on 68°37.7?S; 78°07.8?E) and covering approximately 10 km² in the Vestfold Hills, East Antarctica, have yielded six species of gastropods, and 11 species of bivalves from two beds within the Sørsdal Formation. Most of the material is close to in situ but some specimens have been disturbed from their life position; there is no evidence of significant transport. The gastropods include Nacella concinna (Strebel, 1908), Falsimargarita parvispira Quilty, Darragh, Gallagher & Harding sp. nov., indeterminate species of trochids and naticids, Chlanidota (Chlanidota) sp. cf. C. signeyana Powell, 1951, and two species of Trophon/Trophonella. Bivalves include Ennucula sp. aff. E. grayi (d’Orbigny, 1846), Aequiyoldia defossata Quilty, Darragh, Gallagher & Harding, sp. nov., ‘Pectunculina’ sp., Lissarca sp., Austrochlamys anderssoni (Hennig, 1911), Ruthipecten campestris Quilty, Darragh, Gallagher & Harding sp. nov., Adamussium necopinatum Quilty, Darragh, Gallagher & Harding sp. nov., Limatula (Antarctolima) sp. cf. L. hodgsoni (Smith, 1907), Cyclocardia magna Quilty, Darragh, Gallagher & Harding sp. nov., ?Hiatella sp. cf. H. arctica (Linnaeus, 1767) and Laternula elliptica (King, 1832). Preservation varies considerably owing to recrystallization, dissolution or distortion through compaction, so several species are left in open nomenclature. Oxygen isotope data indicate that water temperature was 4–7.5°C at the time of shell growth. Many species or species groups are now extinct or have migrated away from the Antarctic to the sub-Antarctic region. An Antarctic mollusc fauna has been characteristic of the region for much of the Cenozoic.

Patrick G. Quilty [p.quilty@utas.edu.au], Discipline of Earth Sciences, University of Tasmania, Private Bag 79, Hobart, Tasmania 7001, Australia; Thomas A. Darragh [tdarragh@museum.vic.gov.au], Museum Victoria, GPO Box 666 Melbourne, Victoria 3000, Australia; Stephen J. Gallagher [sjgall@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Lucy A. Harding [harding.lucy.a@edumail.vic.gov.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia.     

A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales,Australia)

?erňanský, A. & Hutchinson, M.N., 2012. A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales, Australia). Alcheringa, 1–6. ISSN 0311-5518.

We describe an isolated frontal bone referable to a new species, Tiliqua laticephala (Scincidae), from the Pliocene Big Sink doline of the Wellington Caves, central eastern New South Wales, Australia. The bone is very robust, is unusually broad and thick, especially around the bases of the subolfactory processes and represents a large and heavily built lizard. The fossil has multiple fragmentary osteoderms in the frontal region, showing asymmetry in shape and thickness that do not correspond to the more regularly arranged anterior head shields of other scincoids. The specimen shares two unusual character states with extant Tiliqua, especially the large armoured species, T. rugosa. Other large skinks related to T. rugosa (other Tiliqua spp., Corucia zebrata, Egernia cunninghamii, Bellatorias major, Liopholis kintorei) are less similar in terms of frontal shape, thickness, sculpture, osteoderm ornamentation, and positioning of adjacent bones.     

Systematic status of the Miocene darter ‘Liptornis’ hesternus Ameghino, 1895 (Aves,Suliformes, Anhingidae) from Patagonia,Argentina

Diederle, J.M., 1.8.2015. Systematic status of the Miocene darter ‘Liptornis’ hesternus Ameghino, 1895 (Aves, Suliformes, Anhingidae) from Patagonia, Argentina. Alcheringa 39, 589–594. ISSN 0311-5518.

Liptornis hesternus was established by Ameghino in the late 1800s on the basis of a cervical vertebra (NHMUK-A599) from the Santa Cruz Formation (Santacrucian, South American Land Mammal Age, early Miocene, Burdigalian Stage) of Patagonia, Argentina. Although taxonomic attributions were controversial, the specimen is now confidently assigned to Anhingidae. Recently, however, L. hesternus was designated a nomen dubium because of its uninformative diagnostic characters and apparent loss of the holotype. Nevertheless, NHMUK-A599 has been relocated and is redescribed here prompting referral to Anhinga. A combination of traits are shared with the extant Anhinga anhinga and A. melanogaster, and the material is dimensionally compatible with A. anhinga. The estimated body size of NHMUK-A599 would have been larger than A. minuta but less than A. grandis, A. subvolans, A. fraileyi and A. walterbolesi. Finally, Anhinga hesterna is considered valid and represents the stratigraphically oldest occurrence of the genus in South America and the southernmost yet recorded.

Juan M. Diederle [juandiederle@cicyttp.org.ar], Laboratorio de Paleontología de Vertebrados, Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción (CICYTTP-CONICET), Materi y España, E3105BWA Diamante, Entre Ríos, Argentina.     

The first fossil sea turtles (Testudines: Cheloniidae) from the Cenozoic of Australia

An isolated dentary and costal identified as cf. Pacifichelys and Cheloniidae indet., respectively, are described from the upper Miocene–lower Pliocene Black Rock Sandstone of Beaumaris, Victoria, Australia. These remains represent the first fossil evidence of sea turtles from the Cenozoic of Australia. Neither of the fossils can be referred to living genera, indicating that extinct cheloniids occurred in southeast Australian coastal waters for at least part of the late Neogene. Thus, the taxonomic composition of the current sea turtle fauna of Australia was apparently established within the last five to six million years.     

Occurrence of Euowenia grata (De Vis, 1887) (Diprotodontidae,Marsupialia) from the Pliocene Spring Park Local Fauna,northeastern Queensland

Mackness, B.S., Black, K.H. & Price, G.J., 1.10.2014. Occurrence of Euowenia grata (De Vis, 1887 De Vis, C.W. [In Anon.] 1887. Untitled. The Brisbane Courier 9224 (44) (8 August), 6. [Google Scholar]) (Diprotodontidae, Marsupialia) from the Pliocene Spring Park Local Fauna, northeastern Queensland. Alcheringa 39, 000?000. ISSN 0311-5518

Ten specimens including several dentaries and maxillae, recovered from the Pliocene Spring Park Local Fauna, northern Australia, are referred to the diprotodontine Euowenia grata (De Vis). The fossils exhibit minimal dental wear and reveal new characters that are unrecognizable in the holotype. The remains represent at least three animals, effectively doubling the previous number of individuals known for this rare megaherbivore. The new records also provide a significant northern geographic range extension for the species and allow an assessment of intraspecific variation, sexual dimorphism and phylogenetic relationships for the species. Euowenia grata is most similar in morphology to the monotypic Pliocene diprotodontid Meniscolophus mawsoni.

Brian S. Mackness [deceased] and Karen H. Black [k.black@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, NSW, 2052, Australia; Gilbert J. Price [g.price1@uq.edu.au], Department of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia.     

Development of successive karstic systems within the Baix Penedès Fault zone (onshore of the Valencia Trough,NW Mediterranean)

The Baix Penedès Fault zone records successive karstic systems. The outcrops studied correspond to different segments of the fault, which were temporarily connected and disconnected, allowing for different diagenetic processes to occur. The first karstic system affected the Mesozoic rocks due to subaerial exposure after Paleogene compression, an event characterized by widespread dissolution and the generation of vug and cavern porosity. The δ¹⁸O values of the dolomitic sediment filling the initial vuggy porosity are similar to those of the host dolomite, indicating that the sediment comes from the erosion and reworking of the host rock. The second karstic system is related to the upward propagation of the Baix Penedès Fault. This deformation was characterized by random‐fracture fabrics with dolomite cement and sediment. The stable isotopes values and Sr/Ca ratios of both the dolomite cement and sediments are similar to those of the host rock. In contrast, the more depleted δ¹³C values indicate the influence of soil‐derived CO₂ and the opening of the system to meteoric waters. During the third karstic event, the δ¹⁸O, δ¹³C, Sr/Ca ratios, and ⁸⁷Sr/⁸⁶Sr values of diagenetic cements suggest a marine signature, indicating that the karstic sediments were dolomitized under the influence of late Burdigalian‐Langhian marine waters. These marine waters were probably expelled from poorly buried sediments and circulated through faults producing dolomitization of the karstic sediments. A final karst system developed during a period of uplift and subaerial exposure. The δ¹⁸O values, the Mg/Ca and Sr/Ca ratios, and the high radiogenic values of the calcite cements formed during this period indicate precipitation from meteoric waters. The results of this study have implications for carbonate hydrocarbon reservoir analogs subject to karstic influence in the Valencia Trough and elsewhere.