Permian (Kungurian) Foraminifera from Western Australia described by Walter Parr in 1942: reassessment and additions

Haig, D.W., October 2017. Permian (Kungurian) Foraminifera from Western Australia described by Walter Parr in 1942: reassessment and additions. Alcheringa 42, 37–66. ISSN 0311-5518.

Exceptionally well-preserved siliceous agglutinated Foraminifera originally recorded by Walter Parr in 1942 are redescribed and illustrated by rendered multifocal reflected-light images. Significant new observations are made on wall texture and apertural morphology. The specimens are from the Quinnanie Shale and lower Wandagee Formation in the Merlinleigh Sub-basin of the Southern Carnarvon Basin, a marginal rift that splayed from the East Gondwana interior rift. During the Early Permian, a restricted shallow sea inundated the rift. The formations are part of sequence III of the Byro Group and belong within the Kungurian Stage (Cisuralian, Lower Permian). Of the 14 agglutinated species described by Parr, six are retained under their original names, viz., Hyperammina coleyi Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], H. rudis Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], Ammodiscus nitidus Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], A. wandageeensis Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar], Tolypammina undulata Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar] and Reophax tricameratus Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]; one is transferred to a different species, viz., Thurammina texana Cushman &; Waters, 1928a Cushman, J.A. &; Waters, J.A., 1928a. Some Foraminifera from the Pennsylvanian and Permian of Texas. Contributions from the Cushman Laboratory for Foraminiferal Research 4, 31–55. [Google Scholar]; six are placed with other genera, viz., Thuramminoides pusilla (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Teichertina teicherti (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Sansabaina acicula (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Tolypammina? adhaerens (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Kunklerina subasper (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]), Trochamminopsis subobtusa (Parr, 1942 Parr, W.J., 1942. Foraminifera and a tubicolous worm from the Permian of the North-West Division of Western Australia. Journal of the Royal Society of Western Australia 27, 97–115. [Google Scholar]); and a species of Ammobaculites Cushman, 1910 Cushman, J.A., 1910. A monograph of the Foraminifera of the North Pacific Ocean. Part 1. Astrorhizidae and Lituolidae. United States National Museum, Bulletin 71(1), 134 pp. [Google Scholar] identified by Parr is now left in open nomenclature. From Parr's material, eight additional species are described: two new species, viz., Hyperammina parri sp. nov. and Gaudryinopsis raggatti sp. nov.; rare representatives of Aaptotoichus quinnaniensis Haig, 2003 Haig, D.W., 2003. Palaeobathymetric zonation of foraminifera from lower Permian shale deposits of a high-latitude southern interior sea. Marine Micropaleontology 49, 317–334. 10.1016/S0377-8398(03)00051-3[Crossref], [Web of Science ®] , [Google Scholar]; and very rare species of Lagenammina Rhumbler, 1911 Rhumbler, L., 1911. Die Foraminiferen (Thalamophoren) der Plankton-Expedition, Erster Teil, Die allgemeinen Organizationsverhaltnisse der Foraminiferen. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, Kiel u. Leipzig, 3L.c. (1909), 1–331. [Google Scholar], Giraliarella Crespin, 1958 Crespin, I., 1958. Permian foraminifera of Australia. Bureau Mineral Resources, Geology and Geophysics, Bulletin 48, 1–207. [Google Scholar], Glomospira Rzehak, 1885 Rzehak, A., 1885. Bemerkungen über einige Foraminiferen der Oligocän Formation. Verhandlungen des Naturforschenden Vereins in Brünn 1884(23), 123–129. [Google Scholar], Hormosinella Shchedrina, 1969 Shchedrina, Z.G., 1969. O nekotorykh izmeneniyakh v sisteme semeystv Astrorhizidae i Reophacidae (Foraminifera). Voprosy Mikropaleontologii 11, 157–170. [Google Scholar], and Reophax Denys de Montfort, 1808 Denys de Montfort, P., 1808. Conchyliologie Systématique et Classification Méthodique des Coquilles, Volume 1. F. Schoell, Paris, 409. 10.5962/bhl.title.10571[Crossref] , [Google Scholar], all of which are left in open nomenclature. Hyperammina rudis is the type species of Hyperamminita Crespin, 1958 Crespin, I., 1958. Permian foraminifera of Australia. Bureau Mineral Resources, Geology and Geophysics, Bulletin 48, 1–207. [Google Scholar], a genus now considered a junior subjective synonym of Hyperammina Brady, 1878 Brady, H.B., 1878. On the reticularian and radiolarian Rhizopoda (Foraminifera and Polycystina) of the North Polar Expedition of 1875–76. Annals and Magazine of Natural History, ser. 1(6), 425–440. 10.1080/00222937808682361[Taylor &; Francis Online] , [Google Scholar]. 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David W. Haig [david.haig@uwa.edu.au] Centre for Energy Geoscience, School of Earth Sciences, University of Western Australia, 35 Stirling Highway, Crawley WA 6009, Australia.     

Reappraisal of Austrosaurus mckillopi Longman, 1933 from the Allaru Mudstone of Queensland,Australia’s first named Cretaceous sauropod dinosaur

Poropat, S.F., Nair, J.P., Syme, C.E., Mannion, P.D., Upchurch, P., Hocknull, S.A., Cook, A.G., Tischler, T.R. &; Holland, T. XX.XXXX. 2017. Reappraisal of Austrosaurus mckillopi Longman, 1933 Longman, H.A., 1933. A new dinosaur from the Queensland Cretaceous. Memoirs of the Queensland Museum 10, 131–144. [Google Scholar] from the Allaru Mudstone of Queensland, Australia’s first named Cretaceous sauropod dinosaur. Alcheringa 41, 543–580. ISSN 0311-5518

Austrosaurus mckillopi was the first Cretaceous sauropod reported from Australia, and the first Cretaceous dinosaur reported from Queensland (northeast Australia). This sauropod taxon was established on the basis of several fragmentary presacral vertebrae (QM F2316) derived from the uppermost Lower Cretaceous (upper Albian) Allaru Mudstone, at a locality situated 77 km west-northwest of Richmond, Queensland. Prior to its rediscovery in 2014, the type site was considered lost after failed attempts to relocate it in the 1970s. Excavations at the site in 2014 and 2015 led to the recovery of several partial dorsal ribs and fragments of presacral vertebrae, all of which clearly pertained to a single sauropod dinosaur. The discovery of new material of the type individual of Austrosaurus mckillopi, in tandem with a reassessment of the material collected in the 1930s, has facilitated the rearticulation of the specimen. The resultant vertebral series comprises six presacral vertebrae—the posteriormost cervical and five anteriormost dorsals—in association with five left dorsal ribs and one right one. The fragmentary nature of the type specimen has historically hindered assessments of the phylogenetic affinities of Austrosaurus, as has the fact that these evaluations were often based on a subset of the type material. The reappraisal of the type series of Austrosaurus presented herein, on the basis of both external morphology and internal morphology visualized through CT data, validates it as a diagnostic titanosauriform taxon, tentatively placed in Somphospondyli, and characterized by the possession of an accessory lateral pneumatic foramen on dorsal vertebra I (a feature that appears to be autapomorphic) and by the presence of a robust ventral mid-line ridge on the centra of dorsal vertebrae I and II. The interpretation of the anteriormost preserved vertebra in Austrosaurus as a posterior cervical has also prompted the re-evaluation of an isolated, partial, posterior cervical vertebra (QM F6142, the ‘Hughenden sauropod’) from the upper Albian Toolebuc Formation (which underlies the Allaru Mudstone). Although this vertebra preserves an apparent unique character of its own (a spinopostzygapophyseal lamina fossa), it is not able to be referred unequivocally to Austrosaurus and is retained as Titanosauriformes indet. Austrosaurus mckillopi is one of the oldest known sauropods from the Australian Cretaceous based on skeletal remains and potentially provides phylogenetic and/or palaeobiogeographic context for later taxa such as Wintonotitan wattsi, Diamantinasaurus matildae and Savannasaurus elliottorum.

Stephen F. Poropat* [sporopat@swin.edu.au; stephenfporopat@gmail.com] Department of Chemistry and Biotechnology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Jay P. Nair [j.nair@uq.edu.au; jayraptor@gmail.com] School of Biological Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Caitlin E. Syme [caitlin.syme@uqconnect.edu.au] School of Biological Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Philip D. Mannion [philipdmannion@gmail.com] Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, UK; Paul Upchurch [p.upchurch@ucl.ac.uk] Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK; Scott A. Hocknull [scott.hocknull@qm.qld.gov.au] Geosciences, Queensland Museum, 122 Gerler Rd, Hendra, Queensland 4011, Australia; Alex G. Cook [alex.cook@y7mail.com] School of Earth Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Travis R. Tischler [travisr.tischler@outlook.com] Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia; Timothy Holland [drtimothyholland@gmail.com] Kronosaurus Korner, 91 Goldring St, Richmond, Queensland 4822, Australia. *Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia.     

An Early Permian brachiopod–gastropod fauna from the Calytrix Formation,Barbwire Terrace,Canning Basin,Western Australia

Taboada, A.C., Mory, A.J., Shi, G.R., Haig, D.W. & Pinilla, M.K., 12.11.2014. An Early Permian brachiopod–gastropod fauna from the Calytrix Formation, Barbwire Terrace, Canning Basin, Western Australia. Alcheringa 39, xxx–xxx. ISSN 0311-5518

A small brachiopod–gastropod fauna from a core close to the base of the Calytrix Formation within the Grant Group includes the brachiopods Altiplecus decipiens (Hosking), Myodelthyrium dickinsi (Thomas), Brachythyrinella narsarhensis (Reed), Neochonetes (Sommeriella) obrieni Archbold, Tivertonia barbwirensis sp. nov. and the gastropod Peruvispira canningensis sp. nov. The fauna has affinities with that of the late Sakmarian?early Artinskian Nura Nura Member directly overlying the Grant Group in other parts of the basin but, as with all lower Cisuralian (and Pennsylvanian) glacial strata in Western Australia, its precise age remains poorly constrained, especially in terms of correlation to international stages. Although the Calytrix fauna lies within the Pseudoreticulatispora confluens Palynozone, the only real constraint on its age (and that of the associated glacially influenced strata) is from Sakmarian (Sterlitamakian) and stratigraphically younger faunas. A brief review of radiometric ages from correlative strata elsewhere in Gondwana shows that those ages need to be updated. The presence of Asselian strata and the position of the Carboniferous?Permian boundary remain unclear in Western Australia.

Arturo César Taboada [ataboada@unpata.edu.ar], CONICET-Laboratorio de Investigaciones en Evolución y Biodiversidad (LIEB), Facultad de Ciencias Naturales, Sede Esquel, Universidad Nacional de la Patagonia ‘San Juan Bosco’, Edificio de Aulas, Ruta Nacional 259, km. 16,5, Esquel U9200, Chubut, Argentina; Arthur Mory [arthur.mory@dmp.wa.gov.au], Geological Survey of Western Australia, 100 Plain Street, East Perth, WA 6004, School of Earth and Environment, The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia; Guang R. Shi [grshi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; David W. Haig [david.haig@uwa.edu.au], School of Earth and Environment (M004), The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia; María Karina Pinilla [mkpinilla@fcnym.unlp.edu.ar], División Paleozoología Invertebrados, Museo de Ciencias Naturales de La Plata, Paseo del Bosque s/n, 1900 La Plata, Buenos Aires, Argentina.     

Early Aptian (Early Cretaceous) freshwater bivalves from the Australian–Antarctic rift,southeast Victoria

The brissid echinoid Schizobrissus insignis (Duncan &; Sladen, 1883) is described from the Fulra Limestone (late middle Eocene), in Kachchh, India. Formerly assigned to Peripneustes or to Meoma it is shown that the species possesses a complete subanal fasciole throughout ontogeny, a characteristic feature of Schizobrissus. Eocene species of Schizobrissus and Meoma show several morphological similarities, but diverged during the late Paleogene and early Neogene.     

Remains of a Late Cretaceous pterosaur from the Molecap Greensand of Western Australia

Pterosaur remains are very rare in Australasia and especially in Upper Cretaceous strata. Thus, the discovery of a jaw fragment from the Cenomanian–Coniacian Molecap Greensand near Gingin in Western Australia represents an important new stratigraphical occurrence for the region. Although the teeth are not preserved, the presence of labio-lingually compressed alveoli that are anterolaterally oriented, variable in shape/size (inferring heterodonty) and very widely spaced is reminiscent of ornithocheirids—a geographically cosmopolitan clade of predominantly Early Cretaceous pterodactyloids. If correct, this identification could extend the known range of Ornithocheiridae through to the Late Cretaceous in the Southern Hemisphere.     

Leaf galls on Dicroidium hughesii (Feistmantel) Lele from the Triassic of India—a new record

Disc-like galls are reported on the Triassic corystosperm leaf Dicroidium hughesii (Feistmantel) Lele from the Parsora Formation of South Rewa Gondwana Basin, central India. Although there have been numerous reports of arthropod–plant interactions from Permian and Lower Cretaceous successions, this is the first unequivocal report of arthropod–plant interactions from the Triassic succession of Peninsular India. The new record adds to global evidence that arthropod herbivory and gall formation, in particular, had rediversified by the Late Triassic in the wake of the end-Permian mass extinction.     

An Early Permian brachiopod fauna of Gondwanan affinity from the Baoshan block,western Yunnan,China

The present paper describes and illustrates an Early Permian brachiopod fauna collected from two localities from the upper part of the type Dingjiazhai Formation near Youwang, 30 km south of Baoshan in the Baoshan block, western Yunnan, China. The brachiopod fauna is dominated by Stenoscisma sp. and Elivina yunnanensis sp. nov. and exhibits strong generic and some specific links with faunas from the Bisnain assemblage of Timor and the Callytharra Formation of Western Australia and, to a lesser extent, faunas from the Jilong Formation of southern Tibet, the Tashkazyk Formation of southeastern Pamir, the lower Toinlungkongba Formation of northwestern Tibet, the upper Pondo Group of central Tibet, and the Jimba Jimba Calcarenite of the Carnarvon Basin, Western Australia. Based on these correlations, a Late Sakmarian (Sterlitamakian) age is preferred for the Dingjiazhai brachiopod fauna. Two new species are proposed: Globiella youwangensis sp. nov. and Elivina yunnanensis sp. nov.     

Umkomasia (megasporophyll): part 1 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed

The genus Umkomasia, a megasporophyll, belonging to the pteridosperms (seed ferns) in the family Umkomasiaceae (Corystospermaceae), is reassessed comprehensively worldwide. All previous records are analysed. Certain fertile structures previously attributed are reclassified. Umkomasia is shown to be restricted to the Triassic of Gondwana where it is associated with the genus Pteruchus, a microsporophyll, and the genus Dicroidium, a vegetative leaf. It is well represented from Argentina, Australia and southern Africa where the Molteno Formation is by far the most comprehensively sampled with eight species described. Two specimens from the upper Permian of India attributed to Umkomasia are reclassified as cf. Arberiopsis sp. A whorled fertile structure from Antarctica, previously assigned to Umkomasia, is reclassified in a new genus as Axsmithia uniramia. Another compression fossil and the permineralized Umkomasia resinosa remain as valid records from Antarctica. The material described as Umkomasia from the Triassic of China is reclassified as Stenorachis asiatica. The Lower Jurassic record from Germany is placed in a new genus as Kirchmuellia franconica. The records of Umkomasia sp. from the Rhaetic of Germany are reclassified as cf. Kirchmuellia sp. and the single specimen from the Jurassic of Libya as genus et sp. indet. The Lower Cretaceous record from Mongolia has been reclassified by other researchers as Doylea mongolica. A pictorial key to Umkomasia species is provided, geographic and stratigraphic distributions are tabulated.     

Paleocene flora from Seymour Island,Antarctica: revision of Dusén’s (1908) angiosperm taxa

The Paleocene flora from Seymour Island, Antarctica, is one of the most diverse floras of this age in the Southern Hemisphere. First collected on the Swedish South Polar Expedition (1901–1903), it was described by Dusén in 1908 as having 87 leaf taxa. Forty-seven angiosperm taxa were described and/or illustrated. Many species are based on single specimens, and the flora has not been re-examined in its entirety since it was first described. This study is the first reassessment of the flora updating the original research using current methodologies, and permitting evaluation of the flora in the context of modern ideas on plant evolution and palaeogeography. This paper continues the revision of the material first studied by Dusén; a previous paper described the ferns and gymnosperms; here we describe the angiosperms. The revision is based on the original collections held at the Swedish Museum of Natural History, together with the first major new collections held at the British Antarctic Survey. Among the taxa recognized by Dusén, we recognize only three entire-margined and 11 tooth-margined angiosperms. This revision to 14 species notably lacks the two tropical elements originally described from the flora, Mollinedia seymourensis and Miconiiphyllum austral. Hence, its status as a ‘Mixed Flora’ comes into question and influences climatic interpretations based on Gondwanan floras. Anne-Marie Tosolini^? [a.tosolini@unimelb.edu.au], Department of Earth Sciences, University of Leeds, Leeds LS2 9JT, UK, and Environmental Geoscience, Latrobe University, Victoria, 3086, Australia; David Cantrill [david.cantrill@rbg.vic.gov.au], National Herbarium of Victoria, Royal Botanic Gardens Melbourne, Private Bag 2000, Birdwood Ave, South Yarra, Victoria, 3141, Australia; Jane Francis [j.francis@earth.leeds.ac.uk], Department of Earth Sciences, University of Leeds, Leeds LS2 9JT, UK. ?Present address: School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia. Received 5.5.2012; revised 13.12.2012; accepted 7.1.2012.     

Pteruchus (microsporophyll): part 2 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed

Anderson, H.M., Barbacka, M., Bamford, M.K., Holmes, W.B.K. & Anderson, J.M., 3 July 2019. Pteruchus (microsporophyll): part 2 of a reassessment of Gondwana Triassic plant genera and a reclassification of some attributed previously. Alcheringa XXX, X–X. ISSN 0311-5518

The microsporophyll genus Pteruchus, belonging to the pteridosperms (seed ferns) in the family Umkomasiaceae (Corystospermaceae), is reassessed comprehensively worldwide and emended. All records are analysed, and some fertile structures previously attributed are reclassified. The Lower Jurassic record of Pteruchus from Germany is ascribed to a new genus as Muelkirchium septentrionalis. Pteruchus is shown to be restricted to the Triassic of Gondwana and is clearly affiliated with the megasporophyll genus Umkomasia and the vegetative leaf genus Dicroidium. It is well represented from Argentina, Antarctica, Australia and southern Africa; the Molteno Formation of southern Africa is by far the most comprehensively sampled, yielding three species and 425 specimens from 22 localities. Nomenclatural problems with the species of Pteruchus are addressed. A key to Pteruchus species is provided; geographic and stratigraphic distributions are tabulated.

Heidi M. Anderson [hmsholmes@googlemail.com], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa; Maria Barbacka [mariabarbacka@gmail.com], W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, 31-512 Kraków, Poland; Hungarian Natural History Museum, Botanical Department, H-1431 Budapest, Pf. 137, Hungary; Marion K. Bamford [marionbamford@witsacza], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa; W.B. Keith Holmes* [wbkholmes@hotmail.com], 46 Kurrajong Street, Dorrigo, NSW 2453, Australia; John M. Anderson [jmandersongondwana@googlemail.com], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa. *Also affiliated with: University of New England, Armidale, NSW 2351, Australia.