Bennett’s wallaby (Macropus rufogriseus) marrow quality vs quantity: evaluating human decision-making and seasonal occupation in late Pleistocene Tasmania

A pilot study on the fatty acid composition of the bone marrow, brain tissue and muscle (flesh) of the medium-sized marsupial the Bennett’s wallaby (Macropus rufogriseus), aimed to explore the mechanisms behind the dominance of this human prey species in late Pleistocene southwest Tasmania. In particular it was designed to investigate why humans frequently selected and transported the wallaby hindquarters, breaking open their longbones and metatarsals to expose the marrow cavity, when these animals could easily have been transported whole. Initial results indicated that the percentage of unsaturated fatty acids particularly palmitoleic acid (C_16:1) and oleic acid (C_18:1) in the bone marrow increased distally away from the body core temperature or the heart, towards the hands and feet. There is a corresponding decrease in the percentage of the saturated fatty acids, particularly palmitic acid (C_16:0) and stearic acid (C_18:0). This is similar to the pattern identified in placental artiodactyls (even-toed ungulates) and rodents. The preliminary study was then expanded to a year-long seasonal experiment involving wallabies from different Tasmanian environmental and climatic zones to see if this can better help explain the apparent seasonal occupation of caves indicated from recent Bennett’s wallaby teeth annuli (Skeletochronological studies). It was found that the Bennett’s wallaby remained a relatively stable and dependable resource throughout the year, with environmental, altitudinal and seasonal variability having only a minor effect on the fatty acid composition or nutritional quality of their bone marrow, muscle and brain tissue. Interestingly, the animals had increased desirable unsaturated fatty acids in their marrow, muscle and brain during the winter and spring with decreased amounts during the summer and autumn. This lends support to an earlier Kidney Fat Index (KFI) analysis for the Bennett’s wallaby which recorded significant seasonal differences within and between regions, indicating an environmental effect on overall body condition. Fatty acid analysis of the Common wombat Vombatus ursinus, the Tasmanian pademelon Thylogale billardierii, and the Forester or Eastern Grey kangaroo Macropus giganteus were also included to ascertain why these animals remained constant, although minor, prey species throughout the late Pleistocene. While the results from these animals were similar to the Bennett’s wallaby, there were some unexpected and surprising patterns that require further investigation. All of these fatty acid analyses have important implications for understanding human decision-making and occupation of southwest Tasmania during the Last Glacial Maximum, for modern dietary and palaeodietary benefits, and for understanding the physiology of these unique Australian marsupials.     

The image of the Frankish kings in the poetry of Venantius Fortunatus

The poetry of Venantius Fortunatus is a sadly neglected historical source for sixth-century Gaul. Amongst the literary material that has survived from that age, the works of Gregory of Tours loom large. Since Gregory provides us with the sole narrative history of Gaul for much of this century, we are forced to see Merovingian society through his eyes. Venantius wrote panegyric, and an age such as ours, which values sincerity of expression, finds little that is attractive in that genre. Despite this, Venantius' poetry affords us a vantage point from which to view the Frankish kings. It also provides important evidence for the nature of the cultural fusion of Germanic, Roman and Christian elements that was taking place in the Gaul of Gregory of Tours and King Chilperic. The poems written for the Merovingian monarchs suggest that Venantius sensed a Frankish hankering after the trappings of Roman imperial authority. He wrote, perhaps with didactic intent, to give full exposition to the traditional Roman conception of the just ruler, coupled with the more recent ideal of the orthodox Christian monarch that was still current in the Byzantine Empire. When Venantius Fortunatus journeyed to the courts of the barbarian kings, he brought with him his cultural baggage from Byzantine Ravenna.