Experimental harvesting of wild peas in Israel: implications for the origins of Near East farming

Previous evaluations of the potential advantage of farming over foraging have been based on experimental harvesting of wild cereals in Near Eastern ecosystems. The highly successful combination of cereals and legumes, “the Near Eastern crops assemblage”, is considered a natural choice due to the complementary nutritional value of the two crop types. Yet, legumes were rarely referred to in models describing early farming and the transition from foraging to farming. Wild legumes differ from wild cereals in several attributes pertaining to domestication, including population density (patchy vs. dense), growth habit (indeterminate vs. determinate), dispersal units (camouflaged seeds vs. awned spikelets), seed dormancy (90–80% vs. 50%), and year to year establishment (erratic vs. regular). We have analyzed the yield potential of three wild pea species (Pisum elatius, P. humile, and P. fulvum) in several ecosystems in Israel. All three pea species have patchy distribution patterns and the yield potential of the wild populations is highly variable. A harvest rate of 0.6–610 grams (calculated per collector) of clean dry grain per hour of harvest time was estimated. Among the three species studied the one adopted for domestication appears to be the least productive. This suggests that the potential productivity of wild peas was not the only or even the major consideration for domestication.     

Charred remains of grains and seeds from the medieval high-status farm site of Reykholt in western Iceland

Abstract

Iceland was settled, primarily by peoples from Norway and the northern British Isles, in the 9th and 10th centuries. The first settlers brought with them from their homelands an agricultural system based on animal husbandry, of which cereal cultivation was an element and also with inputs from fishing, hunting and gathering of wild plants. There are strong indications that barley was cultivated during the first centuries in some parts of the country and that cultivation was at least attempted in other areas. However, Iceland is near the climatic limits of the barley-growing zone, and it is open to discussion how reliable a food source locally grown barley would have been. This paper discusses a seed assemblage of cultivated barley and archetypical weeds of cereal crops dated to between the 10th and 12th centuries AD obtained during archaeological excavations at the high status farm of Reykholt in western Iceland.     

Manuring and stable nitrogen isotope ratios in cereals and pulses: towards a new archaeobotanical approach to the inference of land use and dietary practices

This paper explores the impact of animal manure application on the δ¹⁵N values of a broad range of crops (cereals and pulses), under a range of manuring levels/regimes and at a series of locations extending from northwest Europe to the eastern Mediterranean. We included both agricultural field experiments and areas where ‘traditional’ farming is practised. Our aim is to ground-truth interpretation of δ¹⁵N values in archaeobotanical crop remains as evidence of past growing conditions and husbandry practices. The results confirm the potentially radical impact of manuring on δ¹⁵N values in cereals, depending on manuring level, but indicate only a slight effect on pulses, which can fix atmospheric nitrogen. The expected geographical trend towards greater δ¹⁵N with increasing climatic aridity is not apparent, probably because the growing conditions for crops are ‘buffered’ through crop management. Each of these observations has fundamental implications for archaeobotanical interpretation of δ¹⁵N values as evidence of land use practices and (together with analysis of bone collagen/tooth enamel in potential consumers) palaeodiet.     

Agricultural change in Copper Age Croatia (ca. 4500–2500 cal B.C)?

The Copper Age in the Carpathian Basin is marked by a distinct change in settlement patterns, material culture, social traditions and subsistence practices; however, few studies address the nature of crop cultivation in the region. This paper examines new archaeobotanical data from 13 Copper Age (ca. 4500–2500 cal BC) sites located in continental Croatia, in order to assess the extent to which crop agriculture may have changed and contributed to overall subsistence economies in the Copper Age. From the archaeobotanical results, a dominance in einkorn and emmer is seen followed by barley. Less frequently millet, naked wheat and spelt/new glume wheat are also recovered, but due to their limited numbers, it is less clear whether they were grown as crops or represent weeds. Pulses (e.g. lentil, pea and grass pea), fruit remains (e.g. cornelian cherry and chinese lantern) and wild plant and weed species are also recovered, although more commonly from the late Copper Age sites. The archaeobotanical results show a clear reduction in the quantity and range of plant species recovered during the early/middle Copper Age; however, this is likely the result of taphonomic bias rather than a reduction in crop cultivation. The results therefore highlight problems of recovery bias in the region, which makes comparisons between sites as well as the reconstruction of crop husbandry regimes difficult. Overall, the results from continental Croatia suggest that the type of crops cultivated continued relatively unchanged from the late Neolithic, although it is clear that more research is desperately needed to explore the relationship between crop agriculture and the changing socio-economic environment of the Copper Age in the region.     

Changes in crop cultivation during the last five centuries before the Roman conquest: archaeobotanical investigation in the Clermont-Ferrand basin,Massif Central,France

For several decades, a growing number of protohistoric sites in Auvergne (Massif Central, France)—mostly excavated through preventive archaeology—have been subject to the collection of archaeobotanical analysis. This study presents the archaeobotanical results from ten La Tène sites dating from the last five centuries before the Roman conquest. In addition to providing an inventory of taxa, this paper concerns crops and crop cultivation methods. These results are interpreted in a wider context, dealing with crop husbandry, arable farming, storage and consumption practices. The sites are located in the Limagne plain within a 50-km radius around the city of Clermont-Ferrand. A total of 23,579 carpological remains were identified from 163 sediment samples totalizing a volume of 1096 l. Comparable to other sites in Gaul, the spectrum of cereals and pulses is restricted. Spelt (Triticum spelta), einkorn (Triticum monococcum) and foxtail millet (Setaria italica) are disappearing or becoming rare. A better control of agricultural techniques and increased yields is supposed during the ca. five centuries of the La Tène period. Mixed cultivation is hereby replaced by the sowing of monocrops. At the beginning of this period, farming systems are dominated by a polyculture of cereals and pulses. They become more specialized during the last two centuries before the Roman conquest; diversity of cereals and pulses is decreased.     

Bread and surpluses: the Anglo-Saxon ‘bread wheat thesis’ reconsidered

This paper presents a fundamental new assessment of crop husbandry in the Mid Saxon period in England (c. AD 650–850), using data from charred plant remains. While recent archaeological studies have begun to emphasise the importance of agricultural development in this period – focusing especially upon field systems and livestock – crops have received comparatively little attention. This study challenges one popular model of Anglo-Saxon arable farming, here dubbed the ‘bread wheat thesis’, which posits a Mid Saxon shift whereby bread wheat supplanted hulled barley as the most important cereal crop in this period. The empirical basis for this model is here re-examined in the light of an updated archaeobotanical dataset from selected regions in southern Britain. No evidence for bread wheat supplanting hulled barley is discovered. It is argued instead that rye and oats became substantially more important in the 7th–9th centuries, regional patterns in cereal cultivation in this period correlate with differences in the natural environment and Anglo-Saxon farmers were able to produce greater arable surpluses from the 7th century onwards.     

东南亚地区农业起源研究综论

自20世纪50年代始,东南亚地区的农业起源探索历程至今已半个多世纪。大量考古发现表明,东南亚早期农业的性质是依赖于根系作物的驯化和块茎作物的栽培,该地区早期农业的特点与树木栽培的实践紧密相连,但农业起源的时间尺度仍未很好地建立。基于东南亚地区植物遗存和农业性质的特殊性,多种研究方法得以运用,也相继提出了许多不同的理念。东南亚地区农业起源的研究对中国华南地区早期农业的探索具有很好的启示作用。     

Chickpea domestication in the Neolithic Levant through the nutritional perspective

An alternative approach to the process of selection and domestication of grain crops in early history based on nutritional value is proposed. Selection by a long trial and error process among a number of wild large seeded legumes gave rise to a nutritionally superior domesticated chickpea among the selected “founder crops” of the Neolithic Near Eastern agriculture. We found considerably higher free tryptophan levels in cultivated stocks (44 desi and 29 kabuli types from 25 countries; 1.10 mg/g seed dry weight), compared with the wild progenitor Cicer reticulatum (15 accessions; 0.33 mg/g seed dry weight). Dietary tryptophan determines brain serotonin synthesis, which in turn affects certain brain functions and human behaviour. We suggest that these nutritive facts may explain the decision of prehistoric farmers to choose this rare species and struggle to keep such an agronomically complicated crop under domestication.     

Latitudinal variation in a photoperiod response gene in European barley: insight into the dynamics of agricultural spread from ‘historic’ specimens

Between ca. 6000 BC and ca. 500 BC, barley cultivation spread across the continent of Europe from the extreme south to the extreme north. Carbon-dating would suggest that this spread, and indeed the spread of crop cultivation generally, varied in its pace, with ‘delays’ at certain points along its route. Such delays in the spread of agriculture have been explained as resulting from the slow assimilation of agricultural practices by existing indigenous human populations or as the time taken for the crops to adapt to novel climatic conditions, such as altered temperature regimes and day-lengths. A mutant form of the photoperiod response gene, Ppd-H1, causes barley to be non-responsive to long days, while the wild-type responsive form allows plants to flower in response to long days. We sequenced this gene in 65 ‘historic’ barley accessions, from the late 19th and early 20th centuries, in order to explore the potential role of environmental adaptation in the spread of agriculture. We chose to use ‘historic’ material, to complement the richer patterns in extant genetic lines, by spreading the data range in both time and space. Our ‘historic’ barley data shows a latitudinal divide in the Ppd-H1 gene similar to that found in extant lines, but with clearer geographical resolution, and extending northwards into the Arctic Circle. We discuss the implications of our results in relation to the dynamics of agricultural spread across Europe.     

关于中国稻作起源证据的讨论与商榷

本文是对傅稻镰等人最近的文章批评中国栽培水稻起源研究缺乏证据、提出长江下游水稻栽培开始于公元前4000年这一观点一文的回应。我们首先考察中国从野生稻采集到水稻栽培过程研究的相关文献．进而从水稻形态学和考古学的角度对早期水稻遗存加以研究。讨论集中在以下三个方面：长江流域最早水稻栽培的时间：长江下游和淮河流域初期栽培水稻的遗存;考古遗存所见水稻粒形态大小变化的意义。我们也考察了傅文用来区分野生水稻和栽培水稻的几个标准：考古纪录中所见的未成熟稻遗存、水稻粒的增大和水稻粒的形态。根据已经发表的资料和我们对水稻的研究,我们认为到全新世早期（距今9000年前后）,南北中国的新石器时代居民就可能已经收获野生稻并开始水稻栽培了。     

Early Processed Triticeae Food Remains in The Yanghai Tombs,Xinjiang, China

Visible food remains can provide evidence regarding ancient food processing, the spread of cereals and cultural communication. Some desiccated food remains were discovered in the Yanghai Tombs, Turpan district, in Xinjiang, China (2600–2900 bp ). These food remains were analysed by Fourier transform infrared (FT–IR) spectroscopy combined with plant microfossils, including starch grains and cross cells of pericarp from the cereal bran fragments. The results showed that these food remains were cooked dough food made from wheat (Triticum aestivum) and barley (Hordeum spp.). The cross‐sections of these remains look very dense, not porous under a microscope, which suggests that no fermentation had happened, so these foodstuffs may be some kind of flatbread. Although wheat and barley had been introduced into China by at least the third millennium bc , these remains are still the earliest known direct evidence that wheat and barley were ground into flour and then processed as foodstuffs in north‐western China.     

On the Origins and Dissemination of Domesticated Sorghum and Pearl Millet across Africa and into India: a View from the Butana Group of the Far Eastern Sahel

Four decades have passed since Harlan and Stemler (1976) proposed the eastern Sahelian zone as the most likely center of Sorghum bicolor domestication. Recently, new data on seed impressions on Butana Group pottery, from the fourth millennium BC in the southern Atbai region of the far eastern Sahelian Belt in Africa, show evidence for cultivation activities of sorghum displaying some domestication traits. Pennisetum glaucum may have been undergoing domestication shortly thereafter in the western Sahel, as finds of fully domesticated pearl millet are present in southeastern Mali by the second half of the third millennium BC, and present in eastern Sudan by the early second millennium BC. The dispersal of the latter to India took less than 1000 years according to present data. Here, we review the middle Holocene Sudanese archaeological data for the first time, to situate the origins and spread of these two native summer rainfall cereals in what is proposed to be their eastern Sahelian Sudan gateway to the Red Sea and the Indian Ocean trade.     

Agricultural and household activities in Vestfold,Southeast Norway,as illustrated by pollen data and the charred remains of crops and wild plants

This paper presents and interprets two data sets from Vestfold, Southeast Norway: the pollen record is from a small lake basin, isolated from the sea in Mid Mesolithic (8100–6400 cal BC), and with a record of sediment deposition up to recent time. Charred plant remains from six settlement sites ranging in date from the Late Neolithic (2400–1800 cal BC) to the Merovingian Period (cal AD 570–800). Soil from archaeological contexts that was recovered from several prehistoric settlement features (two- and three-aisled houses, a rock shelter and a pit) has also been investigated. The number and concentrations of identifiable charred macro remains are low from all features except one, but the records contribute to the interpretation of agriculture and wild plant use. Carbonised cereals dated to the Late Neolithic/Early Bronze Age are reported from a two-aisled house. Naked barley was the main cereal identified and a few weed seeds were found with the cereal grains. In a rock shelter nearby, cereals and seeds of flax were found, demonstrating cultivation in the Late Bronze Age. Pollen of ribwort plantain recorded in lake deposits in Nordbytjern, 0·5?km to the southwest, also indicates agricultural activity in the southern part of Vestfold during the Late Bronze Age. Archaeobotanical samples from Early Iron Age houses contained low concentrations of carbonised cereal remains, mainly hulled barley, but also wheat and oat. Seeds/fruits of weeds, plants of moist/wet habitats and grasses increase in abundance from the end of Roman Period. The high concentration of hulled barley found in a pit at the site of Ringdal 13 confirms that hulled barley was a cereal used in the Iron Age. Throughout the Iron Age, cereal pollen has a continuous curve in the Nordbytjern pollen diagram, demonstrating the significance of cereal cultivation in Vestfold. Flax was also cultivated in the vicinity of and probably processed in Nordbytjern. Large numbers of rush seeds and sedge nutlets indicate a possible involvement in basketry and cordage making and/or as animal fodder.     

Plant remains from Roman period town of Obulco (today Porcuna) in Andalusia (Spain)—distribution and domestication of olive in the west Mediterranean region

The aim of this work is (1) to identify the key characteristics of agriculture and the role of olive in Porcuna (Andalusia) of Roman period and (2) to derive a conclusion about the origin of cultivated olive. The study of the literature showed that the olive is one of the most esteemed fruit species in the Classical Mediterranean world and is considered to have been domesticated first in Levant region. The signs of cultivation appear since the Neolithic. The authors analysed 18 samples from archaeological sites. Sorting and identification were carried out using a binocular microscope with the help of comparative collection of seeds/fruits and identification atlases (Digital Seed Atlas of the Netherlands: Cappers et al. 2006). A total number of 19,616 remains was studied. The remains belong to groups of fruit trees, cereals, pulses and synanthropic plants. The cultivated species were used for domestic purposes. The cereals, pulses and mesocarps of the fruits served as food. Important characteristic trait was the overrepresentation of olives. The authors give a summary of the development that led to the cultivation and domestication of the olive tree and to reconstruct its early diffusion using different sources of information, such as archaeobotanical analysis, archaeological data and genetical studies. As a result, a better understanding of the domestication and use of olive on a Mediterranean scale is given. This study also discusses cultivated and synanthropic species in Roman settlements on the Iberian Peninsula.     

Variety in cereal cultivation in the Late Bronze and Early Iron Ages in relation to environmental conditions

The objective of this paper is to assess the relationship between the cereals cultivated in the Late Bronze and Early Iron Ages (ca. 1250–400 BC) within the area of the present-day Czech Republic, and their environmental settings. The various charred caryopses of cereal species represented in the archaeobotanical assemblages from 35 archaeological sites differ, especially in the proportion of wheat and barley. The cereal assemblages were compared regarding site altitude, weather conditions, soils and soil productivity. The most important environmental variable influencing the choice of a particular crop seemed to be altitude which is correlated with other variables such as the length of growing season, mean annual temperature, soil quality etc. Although the ecological requirements of cereals cultivated in the Late Bronze and Early Iron Ages are not known, they presumably thrived under similar conditions to present-day species/varieties, and the strategy of past crop husbandry was based on similar principles as today, e.g. flexible adaptation to local environmental conditions, in an effort to achieve optimal yields and reduce the danger of crop failure.     

Hesitant hunters: a review of the introduction of agriculture in western Norway

Abstract

Evidence for the introduction of agriculture in western Norway is presented, using three categories of data: (1) palaeobotanical data, including pollen diagrams from lakes, bogs and archaeological sites, focusing on the presence of cereals, Plantago lanceolata L. and anthropogenic pollen indicators, and charred macro remains of cereals from archaeological sites; (2) osteological data, focusing on the occurrences of bones of cattle, sheep and goats in three rock-shelters, and the bone material from one open-air Neolithic site; (3) archaeological data, including artefacts indicating agricultural practices, distribution of residential settlement sites, and stray finds. The evidence for agricultural activity at the beginning of the fourth millennium BC (Early Neolithic, EN) is low, whereas the presence of both cereals and animal husbandry is indicated in the palaeobotanical material from the Middle Neolithic A (MNA, 3400–2600 cal. BC). The earliest record of domesticated animal bones is dated to the Middle Neolithic B (MNB, 2600–2200 cal. BC), while palynological and archaeological data also indicate an expansion in the area cultivated by early farmers. All data confirm the establishment of an agrarian society and animal husbandry in the Late Neolithic (LN, 2200–1700 cal. BC). It is concluded that agriculture was introduced into western Norway by the indigenous hunter-fisher populations. During this process, social and ideological factors played principal roles.     

Cereals,calories and change: exploring approaches to quantification in Indus archaeobotany

Several major cereal groups have been identified as staples used by the pre-urban, urban and post-urban phase populations of the Indus Civilisation (3200–1500 BCE): wheat, barley, a range of small hulled millets and also rice, though their proportional exploitation is variable across space and over time. Traditional quantification methods examine the frequency, intensity and proportionality of the use of these crops and help ascertain the ‘relative importance’ of these cereals for Indus populations. However, this notion of ‘importance’ is abstracted from the daily lives of the people using these crops and may be biased by the differential production (as well as archaeological survival) of individual cereals. This paper outlines an alternative approach to quantifying Indus cereals by investigating proportions of calories. Cereals are predominantly composed of carbohydrates and therefore provided much of the daily caloric intake among many late Holocene farming populations. The four major cereal groups cultivated by Indus farmers, however, vary greatly in terms of calories per grain, and this has an impact on their proportional input to past diets. This paper demonstrates that, when converted to proportions of calories, the perceived ‘importance’ of cereals from five Indus sites changes dramatically, reducing the role of the previously dominant small hulled millet species and elevating the role of Triticoid grains. Although other factors will also have affected how a farmer perceived the role and importance of a crop, including its ecological tolerances, investments required to grow it, and the crop’s role in the economy, this papers suggests that some consideration of what cereals meant in terms of daily lives is needed alongside the more abstracted quantification methods that have traditionally been applied.     

Phylogeographic analysis of barley DNA as evidence for the spread of Neolithic agriculture through Europe

Results of analyses of the photoperiod response gene (PPD-H1) and simple sequence repeats (SSRs) in modern landraces of cultivated barley were used as evidence for the mechanism of agricultural spread in Neolithic Europe. In particular, we explored the usefulness of considering adaptive genes as indicators of past selective pressures acting on crops, during their spread through Europe. In some areas, such as the Alpine region, Britain and Scandinavia, we have evidence to suggest that the adaptation of crops to certain climatic conditions may have contributed to the timing of agricultural spread. At the northern fringes of Europe, and in higher altitude locations in central Europe, the introduction of more suitably adapted cereals may have facilitated successful agriculture to trigger agricultural expansion. This research opens up the possibility of investigating other genetic adaptations to climate, which would permit a fuller evaluation of the relative contributions of climate/crop and forager/farmer interactions in the process of agricultural spread.     

Wild lentil and chickpea harvest in Israel: bearing on the origins of Near Eastern farming

Several small seeded plants were domesticated as grain crops in the Near East and other domestication centers. In this study we investigate the potential of small seeded wild lentils, and chickpea as a food source for hunters-gatherers. The yield potential and the return in terms of grams of seeds per hour of collection time were evaluated in several wild populations in Israel. The yield figures never exceeded 50 g/h, and in most cases were below 20 g/h. These data reaffirm Ladizinsky's claim that wild lentils are unlikely to have been a staple resource for hunter-gatherers prior to plant domestication. The result presented herein may be significant vis-a-vis the role attributed to small seeded (‘inefficient’) plants in the Broad Spectrum hypothesis concerning late Paleolithic, pre-agricultural societies. It may also contribute to a more careful interpretation of plant remains recovered from pre-agricultural sites.     

Cultivation of Naked Barley by Early Iron Age Agro-pastoralists in Xinjiang,China

ABSTRACT

We report an archaeobotanical analysis of flotation samples taken from Shirenzigou, an Early Iron Age agro-pastoralist site dated between the fourth and first century BC, located on the northern slope of the Tianshan Mountains in Xinjiang, China. The charred macro-botanical assemblage is dominated by naked barley grains with a few broomcorn and foxtail millet grains. In the context of Trans-Eurasian exchange of cereal crops, southwest Asian crops (wheat and barley) and two Asian millets (broomcorn and foxtail) were introduced to Xinjiang a few centuries to a millennium before Shirenzigou was occupied. The choice of barley cultivation in preference to wheat and millet may have been possibly driven by the relatively extreme local environment and the scheduling requirements of mobile pastoralism. Barley is well suited to this environment, and the choice of naked barley in preference to hulled barley may have been driven by the whole grain tradition prevailing in East Asia.