Technology based evolution? A biometric test of the effects of handsize versus tool form on efficiency in an experimental cutting task

The use of stone cutting tools opened a novel adaptive niche for hominins. Hence, it has been hypothesised that biomechanical adaptations evolved to maximise efficiency when using such tools. Here, we test experimentally whether biometric variation influences the efficiency of simple cutting tools (n = 60 participants). Grip strength and handsize were measured in each participant. 30 participants used flint flakes, while the other 30 used small (unhafted) steel blades. Variations in basic parameters of tool form (length, width, thickness, cutting edge length) were recorded for the 30 flint flakes. It was ensured that mean handsize and strength in each participant group were not significantly different to investigate the effect of tool variation. The experimental task required cutting through a 10 mm-diameter hessian rope. Cutting efficiency was measured using both ‘Number of cutting strokes required’ and ‘Total time taken’. Results show that both efficiency measures were significantly correlated with handsize using all 60 participants. However, no significant differences were found between the flake and blade groups in terms of mean efficiency. Nor was any significant relationship found between tool form parameters and efficiency in the flint flake group. We stress that our results do not imply that tool form has no impact on tool efficiency, but rather that – all other things being equal – biometric variation has a statistically significant influence on efficiency variation when using simple cutting tools. These results demonstrate that biomechanical parameters related directly to efficiency of use, may plausibly have been subject to selection in the earliest stone tool-using hominins.     

A Statistical Examination of Flake Edge Angles Produced During Experimental Lineal Levallois Reductions and Consideration of Their Functional Implications

Recent studies have indicated that Levallois-style core reduction offered potential practical benefits to hominin populations. However, none of these studies have yet considered one of the most important functional attributes of flake tools, which is edge angle. To address this shortcoming, we statistically examined flakes produced experimentally during “classic” or “lineal” Levallois core production and reduction. The primary aim of our analyses was to statistically test the null hypothesis of “no difference” between the edge angles of “Levallois” products and the flakes involved in their production. We employ existing edge angle analytical techniques and develop new comparative methodologies to assess flake blank standardization through the case of Levallois core reduction. Having determined the statistical properties of our experimental Levallois reductions, we thereafter evaluated to what extent edge angles produced may, or may not, have been useful to prehistoric hominins. Our analyses demonstrated that the experimentally produced Levallois edge angles were indeed statistically different from the flakes involved in their production. These differences were evident both in terms of relatively higher (i.e., more obtuse) edge angles than debitage flakes and in being significantly less variable around their higher mean edge angles compared to debitage flakes. However, based on current knowledge of how flake edge angle properties relate to issues of functionality, such differences would not have been detrimental to their functionality. Indeed, the edge angle properties they possess would have provided distinct benefits to hominins engaged in their manufacture. Most notably, Levallois-style core organization and reduction would have provided hominins with a reliable means of consistently producing flakes (i.e., “Levallois flakes”) possessing average flake angles that are beneficial in terms of providing a viable cutting edge yet not being so acute as to be friable upon application. Hence, edge angle properties join an array of other features that provide logical motive for why hominins may have organized core production and reduction around Levallois-style patterns at various times and places during the Mid-Late Pleistocene.     

Investigating interrelationships between Lower Palaeolithic stone tool effectiveness and tool user biometric variation: implications for technological and evolutionary changes

Lower Palaeolithic hominins are thought to have been dependent upon stone tools during the acquisition and processing of food resources. Hence, it is hypothesized that the evolutionary advantages provided by efficient stone tool use may have selected for anatomical changes observed in the hand during this period. Similarly, hominin manipulative capabilities are suggested to have been of consequence to Lower Palaeolithic technological choices and tool use capabilities. The extent and character of these relationships are not, however, fully understood and it is not known whether these hypothesized co-evolutionary and co-dependent relationships are consistent across varying technological and task-type conditions. Here, six key biometric parameters of the hand are investigated in terms of their statistical relationship with cutting efficiency using both flakes and handaxes over extended periods of use and in multiple types of cutting task. Results indicate that (1) both handaxe and flake cutting efficiencies are significantly related with biometric variation of individual tool users, (2) relationships between biometric parameters and efficiency are consistent across extended durations but vary dependent upon task-type conditions, (3) manipulative strength is the most significant biometric trait in terms of predicting flake efficiency, while (4) hand size is the strongest predictor of handaxe cutting efficiency. These results demonstrate the long-term impact that stone tool use likely had on the evolution of hominin biometric variation during the Lower Palaeolithic, while also highlighting the variable influence of different tool use contexts. Most notably, results indicate that the onset of the Acheulean may have been dependent, a priori, upon hand dimensions that are close to the modern human range, and that prior to the appearance of this anatomy, handaxe use would have been an impractical (i.e. inefficient) tool use behaviour compared to the use of flakes.     

The relative effects of core surface morphology on flake shape and other attributes

It is long been thought that many flake attributes, including both size and shape, are largely due to the morphology of a core’s flaking surface, yet this has never been tested under strictly controlled conditions. Using molded glass cores with surface morphologies that highly resemble prehistoric ones, this experiment demonstrates that while core surface morphology does exhibit some influence on flake size and shape, a high degree of variation in flakes produced with the same core surface morphology shows that the effects of other independent variables, such as exterior platform angle and platform depth, have an even stronger effect. A major implication of these results is that current approaches to reconstruct prehistoric knapping strategies are overlooking significant sources of variation.     

Experimental assessment of plan-view and profile-view gross-edge curvature on stone flake slicing efficiency

Separating two or more aspects of an object via cutting was likely an important factor in the origin and evolution of flaked stone technology. In recent years experiments have demonstrated that several stone tool attributes can influence different kinds of cutting behaviour: slicing, cleaving, scraping, sawing, drilling, piercing and abrading. Here we experimentally assessed the role of stone flake plan- and profile-view gross-edge curvature in a controlled slicing task. We also assessed the role of edge length. A total of 21 participants, using 252 stone flakes with distinct gross-edge curvatures and edge lengths, were asked to cut through a standardized substrate, and their efficiency in the task was measured over time. Flakes with longer edge lengths increased the efficiency of the cutting task, but increasing either plan- or profile-view edge curvature decreased the efficiency of the cutting task. These results have implications for the emergence of particular tool forms or reduction sequences throughout the Pleistocene, and may in part explain why certain forms were favoured by Paleolithic people, leading to their convergent evolution or widespread transmission.     

Factors affecting Early Stone Age cut mark cross-sectional size: implications from actualistic butchery trials

Early Stone Age cut marks are byproducts of hominins' tool-assisted animal carcass consumption and provide a potential avenue of inference into the paleoecology of hominin carnivory. If diagnostic cut mark characteristics can be linked to flake and core tool use or the completion of distinct butchery actions, it may be possible to infer ancient tool preferences, reconstruct the consumption of specific muscular tissues, and illuminate landscape-scale stone resource use. Recently, diagnostic morphological criteria including cut mark width and depth have been used to identify marks made by different classes of experimental and archaeological stone tools (Bello, S.M., Parfitt, S.A., Stringer, C., 2009. Quantitative micromorphological analyses of cut marks produced by ancient and modern handaxes. Journal of Archaeological Science 36: 1869–1880; de Juana, S., Galan, A.B., Dominguez-Rodrigo, M., 2010. Taphonomic identification of cut marks made with lithic handaxes: an experimental study. Journal of Archaeological Science 37: 1841–1850; Dominguez-Rodrigo, M., de Juana, S., Galan, A. B., Rodriguez, M., 2009. A new protocol to differentiate trampling marks from butchery cut marks. Journal of Archaeological Science 36: 2643–2654). The work presented here adds to this experimental butchery database by using measurements of cut mark cross-section taken from bone surface molds to investigate how stone tool characteristics including flake versus core tool type, edge angle, and tool weight, influence cut mark width and depth, ultimately testing whether cut mark size is a useful indicator of tool identity. Additionally, these experiments investigate the influence of contextual factors, including butchery action, carcass size, and bone density on cut mark size. An experienced butcher used replicated Oldowan flakes and bifacial core tools in experimental trials that isolated skinning, bulk and scrap muscle defleshing, and element disarticulation cut marks on goat and cow skeletons. This sample explores cut mark traces generated under realistic butchery scenarios and suggests the following results: 1) Core and flake tools were equally efficient at completing all butchery tasks in size 1 and 3 bovid carcasses. 2) Samples of cut mark width and depth produced by core and flake tools were similar and cut marks could not be accurately classified to a known tool type. 3) Skinning and disarticulation activities produced significantly wider and deeper marks than defleshing activities. 4) Cut marks on cows tended to be wider and deeper than those on goats. 5) Cut mark width is negatively correlated with bone density when carcass size and bone portion are taken into consideration. These results suggest that a general quantitative model for inferring tool type or edge characteristics from archaeological cut mark size is not warranted.     

Practical quantitative lithic use-wear analysis using multiple classifiers

Although use-wear analysis of prehistoric stone tools using conventional microscopy has proven useful to archaeologists interested in tool function, critics have questioned the reliability and repeatability of the method. The research presented here shows it is possible to quantitatively discriminate between various contact materials (e.g., wood, antler) using laser scanning confocal microscopy in conjunction with conventional edge damage data. Experiments with replica and prehistoric tools suggest the quantitative method presented here provides valid functional inferences and is flexible enough to accommodate other relevant sources of data on tool function.     

The technology of Stone Age colonization: an empirical,regional-scale examination of Clovis unifacial stone tool reduction,allometry, and edge angle from the North American Lower Great Lakes region

There is now broad consensus that the appearance of Clovis in Northeastern North America (Great Lakes, New England) represents a colonization pulse into recently deglaciated landscapes. Due to the increased resource uncertainty that comes with colonizing unfamiliar landscapes, it was hypothesized that the majority tool component of Clovis assemblages, unifacial stone tools, should have been knapped on tool blanks possessing the design properties of longevity and functional flexibility to facilitate exploration mobility and guard against the absence of toolstone sources in the new landscape. These properties are optimized by large, flat flakes, possessing large surface area relative to flake thickness. Since discarded and, at times, exhausted unifacial stone tools do not preserve the original dimensions of the blank upon which they were created – necessary items for a true test of blank morphology selection – this study presents a set of predictions for inferring whether Clovis unifacial stone tool blanks were selected for the properties of longevity and functional flexibility based on evidence that Clovis people actually capitalized on those properties. Due to the nature of Clovis unifacial stone tools, tool size was of necessity used as a proxy for tool reduction, on the grounds that smaller tools are more likely to have been resharpened than larger tools, at least in the case of unifacial flake tools. The results showed that less resharpened tools possessed flatter, less spherical shapes than the more resharpened tools, which possessed more globular, spherical shapes, suggesting Clovis foragers exploited the retouch potential afforded by the larger, flatter blanks. Edge angles showed no relationship with tool reduction, suggesting that Clovis foragers exploited the functional flexibility afforded by flatter blanks by adjusting the edge angle to be either higher or lower as needed. These results are consistent with the notion that human colonizers, who did not know the abundance or location of stone outcrops prior to settling an unfamiliar territory, not only “geared up” before leaving a stone source, but geared up as efficiently as possible by carefully selecting the blanks they chose to carry. Broader implications for such careful unifacial stone tool blank selection are discussed.     

Experimental Flake Scatter-Patterns: a New Interpretative Technique

Abstract

Scatters of flint waste flakes are among the commonest finds on prehistoric sites. To understand how such patterns are formed, we made a series of Neolithic axe roughouts and other tools and recorded the distribution of waste flakes. The most important variable affecting the size and shape of flake scatter patterns seems to be the knapper's position. The applications of these experiments to prehistoric data are considered.     

How flakes shatter: a critical evaluation of quartz fracture analysis

Despite its worldwide use as a stone tool raw material, quartz is known to be a difficult material for archaeologists. The main reason for this is the tendency of quartz flakes to fragment during detachment, which complicates the use of traditional lithic analyses. In this article we present an experimental study of quartz flake fragmentation. We evaluate the method called fracture analysis that has been developed and used explicitly for the study of quartz assemblages. The method assumes high predictability of quartz flake fragmentation, but our experiments show that there is significant variation in fragmentation that fracture analysis does not take into account. Our results indicate that this variation is partly explained by indenter hardness, the relative thickness of the detached flake, as well as individual knapper-related factors. These results undermine the applicability of quartz fracture analysis in its current form. In addition, we discuss the effects of flake fragmentation on the technological organisation of prehistoric quartz users and suggest that it has affected reduction strategies as well as blank and tool dimensions. We also suggest that there should be mobility-related differences in archaeological assemblages in terms of the quality of the quartz raw material and that the curation of quartz should be low in relation to better quality raw materials used parallel with it.     

Biometric variables predict stone tool functional performance more effectively than tool‐form attributes: a case study in handaxe loading capabilities

Both the form of a stone tool and the anatomy of the individual using it have potential to influence its cutting performance. To date, however, the selective pressures acting on stone‐tool form and hominin biometric/biomechanical attributes have been investigated in isolation and their relative influence on performance have never been compared directly. This paper examines the influence of both tool‐form attributes and biometric variation on the functional performance of Acheulean handaxes. Specifically, it investigates the impact of 13 tool attributes and eight biometric traits on the working forces applied through the edge of 457 replica tools. The relative contribution of tool‐form and biometric attributes to handaxe loading levels were examined statistically. Results identify that both tool‐form attributes and biometric traits are significantly related to loading; however, tool–user biometric variation has a substantially greater impact relative to tool‐form attributes. This difference was demonstrated by up to a factor of 10. These results bear directly on the co‐evolutionary relationships of stone tools and hominin anatomy, and the comparative strength of selective pressure acting on each. They also underline why handaxe forms may have been free to vary in form across time and space without necessarily incurring critical impacts on their functional capabilities.     

Could woodworking have influenced variation in the form of Acheulean handaxes?

The relationship between tool form and function is fundamental to hominin behaviour and evolution. Acheulean handaxes are known for their general consistency across more than a million years and three continents, albeit with some variation in size and shape. However, the influence of this variation on cutting has only rarely been studied, mostly in either butchery or generalized cutting tasks. Yet evidence indicates that handaxes were used for woodworking by at least 1.5 mya. Here, we experimentally tested whether woodworking could have exerted selective pressures on handaxe form. Additionally, these data were compared with a previous experiment that tested flakes during woodworking. For handaxes, no significant relationships were detected in woodworking efficiency. Accordingly, woodworking likely did not exert strong selective pressures on handaxe variability. However, the effectiveness shown by handaxes further demonstrates the functional flexibility of this technology, which is likely a factor contributing to their use over broad temporal, geographical and ecological spans. When comparing handaxes with flakes, only the smallest flakes were found to be significantly less efficient than handaxes. Therefore, the functional demands of woodworking were likely influencing hominin technological decisions about flake morphology, or the selection of flakes for woodworking, more than they were handaxe morphology.     

Measuring skill in the production of bifacial pressure flaked points: a multivariate approach using the flip-test

Skill is an aspect of prehistoric technology that can inform us on many areas of investigation. This article discusses the notion of skill in prehistoric contexts and how skill is to be formally defined in relation to lithic bifacial tools. The nature of bifacial manufacture entails simultaneous attention to the facial, profile- and cross- section morphology of the core, since each flake removal affects all features. It is argued that bifacial skill can be measured using a multivariate approach, which takes all these features into account. An index measure, the “Bifacial Skill Score”, is proposed and evaluated using both experimental and archaeological data. This measure is argued to constitute a good proxy for skill in bifacial technology and a useful tool for comparative research.     

Milky Quartz Bipolar Reduction and Lithic Miniaturization: Experimental Results and Archaeological Implications

We present experimental data examining the energetics and identification of axial bipolar reduction in contexts of lithic miniaturization on milky quartz. These experiments answer two specific questions. First, does bipolar reduction provide any benefits over freehand reduction? Second, can axial bipolar reduction be distinguished from freehand reduction? Our data show that bipolar reduction requires significantly less time to reduce a percentage unit of core mass and to produce a millimeter of cutting edge on milky quartz than freehand reduction. Milky quartz bipolar reduction surpasses even the cutting edge production efficiency of obsidian pressure blades. We outline a series of quantitative criteria for identifying bipolar cores and flakes. Our results show that bipolar cores and flakes can be distinguished from those produced using freehand reduction by quantifying platform crushing, distal flake rebound scars, bulb shearing, as well as axial, bipolar, and splintered flake terminations. Our results challenge the widely held perceptions about the wastefulness of bipolar reduction and provide clear guidelines for identifying this reduction strategy in archaeological milky quartz assemblages.     

Introducing a new experimental design for controlled studies of flake formation: results for exterior platform angle,platform depth,angle of blow,velocity, and force

Potential variables that underlie variation in flake size, and in some instances shape, are investigated in a newly designed experiment. This new design, which utilizes glass cores molded to a specific shape, results in flakes that are identical to archaeological ones. Variation in exterior platform angle, platform depth and angle of blow all directly affect flake size, and in the case of exterior platform angle, flake shape as well (in spite of constant core surface morphology). In treating velocity and force independently, neither is found to affect flake size or shape. These results have implications for understanding different strategies that flintknappers may employ to control the size of their flake products.     

Levallois economics: an examination of ‘waste’ production in experimentally produced Levallois reduction sequences

Mathematical modelling has suggested that Levallois core morphology represents a reduction strategy driven by economic considerations; particularly the minimization of ‘waste’ while aiming to maximize cutting edge length of flakes obtained from cores of a given size. Such models are elegant in that they facilitate formal modelling of economic considerations that potentially motivate patterns seen in prehistoric data. However, the abstract nature of such models means that they do not take full account of all the practical difficulties and material challenges involved in reproducing Levallois-style reductions in stone. In particular, such models have only examined nodule morphology in two-dimensions, and did not take account of the fact that in the case of classic (lineal) Levallois reduction, core surfaces must be re-prepared between successive stages of flake removal. Hence, the potential economic implications of these factors are currently unknown, potentially undermining the significance of models that assume specific economic conditions. Here, we undertook to examine these factors using a series of experimentally produced Levallois reduction sequences. A total of 3957 flaking events were considered in our analyses, and we used six specific measures of economy to examine Levallois reduction across successive phases. Our analyses found that key assumptions of mathematical models suggesting that Levallois core morphology was driven by economic considerations (i.e. conservation of raw material when attempting to remove flakes with long cutting edges) can be upheld under the practical challenges of replicating Levallois-style reduction in stone. In supporting the notion that Levallois reduction has advantageous economic properties, our results emphasize the importance of considering why Levallois reduction did not emerge earlier in the archaeological record, and indeed, why even during the later Pleistocene the temporal and geographic distribution of Levallois technology varies. Our results also re-emphasize the value of formally modelling lithic reduction strategies in specific economic terms.     

Laboratory evidence of the use of metal tools at Machu Picchu (Peru) and environs

An interpretation of use-wear marks on metal artifacts is developed from the principles of metal cutting and brittle fracture and applied to surficial markings and microstructural damage on bronze tools from Machu Picchu and environs. Most of the tools have blunt edges, relatively low tin contents, and were not work hardened before use; they appear to have been designed for work that involved breaking chips from hard, brittle material. Use-wear marks on these tools are interpreted as due to sliding contacts and impacts with rock. One tool with a relatively sharp edge has a higher alloy content than those with blunt edges and has been work hardened; it appears to have been designed for cutting wood and use-wear markings suggest it was so used. A long bronze bar carries markings that suggest use by stonemasons. Many of the tools are broken and study of their microstructures shows that the bronze used has poor mechanical properties because of porosity and bands of sulphide inclusions.     

The Analysis of Stone Tool Procurement, Production, and Maintenance

Researchers who analyze stone tools and their production debris have made significant progress in understanding the relationship between stone tools and human organizational strategies. Stone tools are understood to be morphologically dynamic throughout their use-lives; the ever-changing morphology of stone tools is intimately associated with the needs of tool users. It also has become apparent to researchers that interpretations of lithic analysis are more productive when the unique contexts and situations for which lithic artifacts were made, used, modified, and ultimately discarded are considered. This article reviews the recent literature on stone tool production with an emphasis on raw material procurement, manufacturing techniques, and tool maintenance processes as they relate to adaptive strategies of toolmakers and users.