Thanerocleridae is a small family of Cleroidea with no fossil representatives to date. Here we describe and figure the first fossil representative of Thanerocleridae, Cretozenodosus fossilis gen. et sp. nov., from the mid-Cretaceous amber of northern Myanmar. Cretozenodosus is referred to the extant subfamily Zenodosinae as evidenced by its open procoxal cavities and transverse procoxae. Cretozenodosus has close affinities with the North American Zenodosus Wolcott, suggesting that modern Zenodosinae is probably a relict group. Our discovery of a new thaneroclerid genus from Burmese amber suggests that Thanerocleridae originated no later than the mid-Cretaceous.
Chenyang Cai [cycai@nigpas.
A new fossil species, Luanpingia daohugouensis sp. nov., belonging to the family Sinoalidae is described from the Middle to Upper Jurassic Daohugou beds of Inner Mongolia, China, on the basis of two well-preserved complete specimens. The described species of Sinoalidae are reviewed and Jiania gracila is considered a junior synonym of Jiania crebra. The new discovery increases the palaeodiversity of sinoalids from the Daohugou beds. It also indicates stratigraphic correlation between the Daohugou beds, the Haifanggou Formation at Haifeng, Beipiao City, West Liaoning Province, and the Jiulongshan Formation at Zhouyingzi, Luanping County, Hebei Province. All of these units host the ‘early assemblage’ of the Yanliao biota.
Yanzhe Fu [yzfu41@163.
Six conodont and one fusuline zones are recognized on basis of a total of 25 conodont and 13 fusuline species (including seven unidentified species or species given with cf. or aff. in total) from the Bamchi Formation, Yeongwol, Korea. The conodont zones include the Streptognathodus bellus, S. isolatus, S. cristellaris, S. sigmoidalis, S. fusus and S. barskovi zones in ascending order, which can be correlated with the conodont zones spanning the uppermost Gzhelian to Asselian Age of the Permian globally. The fusuline zone is named the Rugosofusulina complicata–Pseudoschwagerina paraborealis zone. The co-occurrence of the conodont Streptognathodus isolatus (the Global Boundary Stratotype Section and Point index for the base of Permian) and Pseudoschwagerina (a Permian inflated fusuline) indicates that the Carboniferous–Permian boundary can be placed in the lower part of the Bamchi Formation in South Korea.
Qiulai Wang* [qlwang@nigpas.
Omma Newman is an extant ommatid genus currently endemic to Australia. A new Omma species, O. daxishanense sp. nov. is described and illustrated based on a compression fossil from the Upper Jurassic Tiaojishan Formation at Daxishan, a fossil locality well known for yielding mammals, feathered dinosaurs and diverse pterosaurs. Omma daxishanense is very similar morphologically to the extant O. sagitta, but differs from the latter by its broader body and prominent temples. The new discovery documents the first valid Omma species from the Mesozoic of China and highlights the antiquity and palaeodiversity of the extant Australian endemic genus.
Chenyang Cai [cycai@nigpas.
The smooth atrypoid brachiopod Thulatrypa gen. nov. incorporates two species, a younger (T. gregaria) from Norway, and an older (T. orientalis) from South China, which collectively span the middle Rhuddanian through Aeronian. In Baltica, the genus thrived just below the storm wave base in a tropical BA4 setting extending slightly into BA3 and BA5 respectively, whereas in South China, its representative occurs in a much shallower assemblage (BA2–3). Their palaeobiogeographical implications are carefully investigated. This study supports the arguments that Thulatrypa may have originated in South China in the middle Rhuddanian and extended its range to eastern Baltica in the late Rhuddanian. Larvae may have drifted along a channel from the east to the southwest of Baltica, which supports the reconstructions of palaeocurrents in the early Silurian in previous palaeogeographical studies.
Bing Huang [bhuang@nigpas.
A nearly complete specimen of Juraheterophlebia cancellosa sp. nov., the third species of the family Juraheterophlebiidae, is described from the Middle–Late Jurassic of China and shows the exact structure of its forewing discoidal space. As a consequence, this family is restored, separated from Erichschmidtiidae, and its diagnosis amended. It is transferred from Heterophlebioptera to Stenophlebioptera, the first clade being now only known from the Early Jurassic. Erichschmidtiidae includes the sole species Erichschmidtia nigrimontana, and this family is now considered of uncertain systematic position.
Diying Huang [dyhuang@nigpas.
Extant tettigarctids are also known as hairy cicadas because they are covered by long and abundant hairs. This character had not been reported in fossil species of Tettigarctidae because previous examples were poorly preserved or lacked long hairs. Hirtaprosbole erromera gen. et sp. nov. (Tettigarctidae) with a hairy body, from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China, is described here. This new species provides evidence that tettigarctids with long dense hairs had appeared by the latest Middle Jurassic and lived at high altitudes.
Xiao-hui Liu [liuxh8917@163.
A new form of dendroid multicellular thallophyte is documented in the Ediacaran Doushantuo phosphorite at Weng’an, Guizhou Province, southern China. The dendroid thallophytes have variable forms, possibly owing to heteromorphic variation. Many lateral branches extend from the upper portions of the main axes; the lateral branches bear terminal vegetative vesicles, reproductive vesicles, monosporangium-like discoidal vesicles and urn-shaped pseudoparenchymatous structures. The vegetative vesicles give rise to clavate pseudoparenchymatous structures, characterized by differentiation of the thallus medulla/cortex, which might represent an early stage of thallus development. An oogamous conceptacle arising from one carpogonial vesicle forms a highly specialized goblet-shaped conceptacle. The discovery of the new dendroid multicellular thallophytes provides not only the first fossil-based evidence of the morphological complexity and tissue differentiation in the Precambrian organisms but also insights into the life cycle of the Precambrian red algae.
Wei Du* [duwei@ea.
Palynological analysis was carried out on Middle to Upper Triassic strata from Tulong, Nyalam County, southern Xizang (Tibet), China. Well-preserved miospore (pollen and spore) assemblages and sparse acritarch occurrences were identified. We recognized four formal and one informal biozones based on stratigraphically important taxa and compositional changes through the succession, in ascending order: the Triplexisporites Interval Zone (Anisian), the Staurosaccites quadrifidus Taxon-range Zone (upper Anisian to lower Norian), the Striatella Interval Zone (lower Norian), the Craterisporites rotundus Taxon-range Zone (middle to upper Norian) and the informal ‘Dictyophyllidites harrisii zone’ (Rhaetian). The zonation was supported by marine fossils (e.g., ammonoids and conodonts), and compositional similarity between the zones was examined using non-metric multidimensional scaling (NMDS). Correlation with other representative palynological sequences across Gondwana was also conducted. The presence of miospore taxa not previously recovered from the Late Triassic North and South China palynofloral provinces (e.g., Ashmoripollis reducta, Craterisporites rotundus, Enzonalasporites vigens, Minutosaccus crenulatus, Samaropollenites speciosus and Staurosaccites quadrifidus) calls for a new province in southwestern China, i.e., the Southern Xizang Province. It is proposed here that the modern expression of the northern boundary runs along the Yarlung Zangbo Suture, the remnant of the Tethys that separated the Indian Plate (southern Xizang) and the Lhasa Block during the Late Triassic. This new palynofloral province comprises typical elements of the Onslow Microflora, indicating the need for an extension of this microflora in southern Xizang, China.
Jungang Peng [jgpeng@nigpas.
New gazelle fossils are described from the Siwalik Group of Pakistan. The material includes horncores, maxilla and mandible fragments, and isolated teeth. The available samples are assigned to three Gazella species: Gazella sp. in the Lower Siwalik Subgroup (ca 14.2–11.2 Ma), and G. lydekkeri and G. superba in the Middle Siwalik Subgroup (ca 10.2–3.4 Ma). Based on a review of the Siwalik Group gazelles, G. padriensis is synonymized with G. lydekkeri. Gazella superba Pilgrim, 1939 sensu stricto is a large form and is a valid species of the genus in the Siwalik Group.
Muhammad Akbar Khan [akbaar111@yahoo.
The discovery of a well-preserved dragonfly forewing in the Upper Jurassic Talbragar Fish Bed near Gulgong and attributed to Austroprotolindenia jurassica Beattie & Nel allows this taxon to be placed in Protolindeniidae. It extends the palaeogeographical distribution of this family, previously known only from the Jurassic of Europe, to Australia.
André Nel [anel@mnhn.
Campterophlebiidae is the most diverse family of fossil odonatans in China with ten genera recovered mostly from Middle Jurassic strata of Inner Mongolia. We describe a well-preserved campterophlebiid damsel-dragonfly from the Middle Jurassic Yanan Formation in Shanxi Province, northwestern China. This discovery adds to the diversity of Campterophlebiidae and identifies a new Middle Jurassic insect fossil locality in China. Within Campterophlebiidae, the new taxon most closely resembles Ctenogampsophlebia from the Middle Jurassic of Inner Mongolia but differs from other genera in having vein AA with four parallel posterior branches uncrossed in the anal triangle.
Daran Zheng [dranzheng@gmail.
A new insect species, Cixius discretus (Hemiptera, Fulgoromorpha), from the Lower Miocene Garang Formation of Zeku County, Qinghai Province (northeastern Tibetan Plateau) is described. This species can be assigned to Cixiidae and represents the first fossil representative of this family from Qinghai Province. Based on the recent single-origin hypothesis and the distribution of tectonic plates in the Cretaceous, we consider that ancient Cixius had dispersed globally prior to the Cretaceous. Through analysis of the habitats of extant Cixius, the palaeoclimate and fossil flora of the Zeku area during the Miocene, we interpret the climate of Zeku in the early Miocene to have been warm-temperate and mildly arid. The new species constitutes evidence of wooded and shrubby habitats in Zeku during the Miocene.
Yi Li [liy0124@sina.
We describe an isolated osteoderm from the Albian Griman Creek Formation where it is exposed near the town of Lightning Ridge in central-northern New South Wales, Australia. Several lines of evidence allow referral of this element to the Ankylosauria—a group that epitomises body armour and ubiquitous osteodermal coverage among dinosaurs. Despite the abundant record of fossil vertebrates from this interval, ankylosaurians have not been previously reported, although, they have been described from penecontemporaneous deposits in western Queensland and Victoria. This discovery, therefore, provides an important link between the northerly faunas (including the Griman Creek Formation) that flourished at the edge of the epeiric Eromanga Sea, with those from the sub-polar rift-valley system of Victoria during the mid-Cretaceous.
Phil R. Bell [pbell23@une.
Two new Psocodea, Sinopsyllipsocus fushunensis gen. et sp. nov. and Eotriplocania sinica gen. et sp. nov., are described from Eocene amber of Fushun City, China. They are distinctly different from all known Psocodea from Fushun amber in their three-segmented tarsi. Sinopsyllipsocus fushunensis is the second unequivocal fossil of Psyllipsocidae. Eotriplocania sinica is the first Asiatic and oldest representative of the Neotropical family Ptiloneuridae, and reveals a formerly global distribution of the family. The discovery of these two families in Eocene Fushun amber suggests a rather warm palaeoclimate for the Fushun amber locality.
Qingqing Zhang [qqzhang@nigpas.
Historical fossil insect collections from England were re-examined and the taxa revised. Lateophlebia gen. nov. is erected for Liassophlebia anglicanopsis (Zeuner) in Campterophlebiidae. Petrophlebia anglicana Tillyard is confirmed in this family and Archithemis liassina (Strickland) is transferred to this family. Lastly, Archithemis brodiei (Geinitz), Archithemis Handlirsch, and Architemistidae Tillyard (reduced to this sole species) are transferred to the Heterophlebioidea.
Richard Kelly [richard.
A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.
Philippa M. Chamberlain [philippa.
An isolated turtle xiphiplastron similar to that of Puppigerus sp. is described from the upper Oligocene (27.3–25.2 Ma) Pomahaka Formation near Tapanui, Otago, New Zealand. The bone is unlike any previously described turtle from the Cenozoic of New Zealand and is from a newly recognized estuarine vertebrate locality. It represents the first Oligocene cheloniid turtle bone described from the southwest Pacific.
Henry J. L. Gard [henrygard@hotmail.