Who brought in the rabbits? Taphonomical analysis of Mousterian and Solutrean leporid accumulations from Gruta do Caldeirão (Tomar,Portugal)

In the Iberian Peninsula, leporids, and specifically rabbits, play a key role in the understanding of hunter-gatherer economies. They appear to have been especially important in the Tardiglacial, when large numbers of small prey animals and of the European rabbit (Oryctolagus cuniculus) in particular, are a ubiquitous feature of faunal assemblages from archaeological sites. Since a large number of non-human predators can also contribute to the formation of such assemblages, the ability to discriminate between bones accumulated by humans and by other kinds of predators is a key prerequisite to their interpretation. On the basis of systematic actualistic studies carried out on modern leporid remains produced by mammalian terrestrial carnivores, nocturnal and diurnal raptors, and humans, we identified diagnostic taphonomic indicators of the different predators. In this paper, the patterns observed on the modern material are applied to the taphonomical analysis of two archaeological samples of rabbit and hare remains from Mousterian and Solutrean layers of Gruta do Caldeirão, a cave site located in Central Portugal. Our results suggest that Eagle Owl (Bubo bubo) were mainly responsible for the Mousterian accumulations, whilst the Solutrean ones were most likely the result of human activity. These data support the notion that, in Iberia, significant reliance on rabbits does not become a feature of subsistence strategies until later Upper Palaeolithic times.     

Big Game and Small Prey: Paleolithic and Epipaleolithic Economy from Valencia (Spain)

This study analyzes the faunal assemblages corresponding to the Middle and Upper Paleolithic and the Early Epipaleolithic for the central Spanish Mediterranean Region dated from between 135,000 and 9,000 B.P. To interpret these data, we employ a zooarchaeological and paleoeconomic perspective. Human large and small game hunting systems are analyzed through the identification of the origins of faunal assemblages, bone breackage patterns, and prey selection patterns. Our study shows that hunting systems changed significantly through time, which may be related to the characteristics of the Mediterranean ecosystem and, above all, to cultural changes.     

When wildcats feed on rabbits: an experimental study to understand the taphonomic signature of European wildcats (<Emphasis Type="Italic">Felis silvestris silvestris</Emphasis>)

Studies of the feeding ecology of the European wildcat (Felis silvestris silvestris) demonstrate that leporids, mostly European rabbit (Oryctolagus cuniculus), dominate their diet in regions where they are present. The remains of wildcats have been found at Pleistocene and Holocene archaeological sites, raising the possibility that they actively accumulated leporid bones in caves and shelters shared with other terrestrial carnivores, raptors and humans. We present the first taphonomic study of rabbit remains consumed by this terrestrial carnivore, with the ultimate aim of understanding their role in bone accumulations at archaeological sites. An experimental study was carried out with a wildcat female, who was fed with nine complete rabbit carcasses. Non-ingested remains and scats were recovered for the analysis of anatomical representation, breakage and bone surface modification. This revealed that non-ingested remains and scats of the European wildcat can be discriminated from most other agents of accumulation. The referential framework provided will permit the discrimination of hominids and wildcats as agents of fossil accumulations of rabbits.     

On the Causes of the Extinction of Some Animals at the end of the Pleistocene

A mathematical theory is developed to demonstrate the disputed hypothesis that hunting tribes of the Upper Paleolithic in Europe did play a decisive role in bringing about the extinction of the mammoth and other large herbivorous animals at the end of the Pleistocene. Formulas for the periods of time during which hunting did not greatly affect animal numbers and during which the numbers declined to zero are used to show that mammoth numbers remained relatively stable during most of the Upper Paleolithic (a period of 10,000 to 25,000 years) and then dropped rapidly within a few centuries to complete extinction. The theory is used to explain why only the largest animals became extinct and why elephants continued to survive in tropical areas while the mammoth vanished in the north.     

Exploitation of Lithic Raw Material in The Northwestern Caucasus Upper Paleolithic

Sources of lithic raw material discovered by the authors are described with special regard to methods of transporting stone to Upper Paleolithic sites in the northwestern Caucasus. The analysis of obsidian artifacts suggests that materials were imported from remote regions such as the central and South Caucasus, evidencing mobility patterns in the Upper Paleolithic.     

The Upper Paleolithic of the Russian steppe zone

The vast steppe area North of the Black Sea has been populated since the early Middle Paleolithic. The number of known sites dating to the Upper Paleolithic is increasing as research progresses, and today about 150 Upper Paleolithic sites have received some (although varied) degree of study. For a long time, the steppe zone was considered to have had a separate cultural-economic adaptation that differed drastically from that found in the periglacial province farther north. The difference was expressed in the specialization in bison hunting and in the very high mobility of the population. More recent archaeological research, described briefly in this article, presents a different picture: Subsistence was more varied, hunting was not limited to bison, and the settlement system indicates long periods of occupation of the sites. There were many local archaeological cultures, and study of the material remains indicates numerous contacts between the people of the steppe zone and those in Central Europe and the Caucasus and, by way of the Caucasus, with Western Asia.     

Neandertals,early modern humans,and rodeo riders

In 1995 Berger and Trinkaus (J. Archaeol. Sci. 22, 841–852) proposed that the anatomical distribution of Neandertal trauma, with a predominance of upper body lesions, reflected close-quarter ambush hunting as dictated by the available Middle Paleolithic weaponry (the “Rodeo rider” hypothesis). The necessity for mobility among these Late Pleistocene foragers, as a factor possibly reducing the number of preserved lower limb injuries, was considered as an alternative explanation. The accumulating data on Upper Paleolithic injuries and Middle Paleolithic weaponry, considerations of differential skeletal susceptibility of minor trauma, and evidence of interhuman violence, plus the importance of mobility for Late Pleistocene human existence, suggest that hunting injuries may explain only part of the pattern. The purpose of this note is not to resolve to ultimate factors behind the anatomical distribution of traumatic lesions among the Neandertals (or early modern humans). It is 1) to emphasize that there are multiple probable contributing factors other than close-quarter ambush hunting due to the limitations of Middle Paleolithic weaponry, and 2) to open the discussion to alternative interpretations.     

Upper Paleolithic Net-Hunting, Small Prey Exploitation, and Women's Work Effort: A View from the Ethnographic and Ethnoarchaeological Record of the Congo Basin

Recent interest in Upper Paleolithic small prey acquisition focuses on the significance of fiber-based hunting technologies. Some researchers believe the advent of these technologies and presence of small faunas reflect efficient communal net-hunts driven by women's labor. We evaluate different small prey hunting techniques, using ethnographic data from foragers in the Congo Basin. These and other ethnographic data suggest that net-hunting is a high risk endeavor that often has high opportunity costs. We argue that the high costs associated with net-hunting have profound implications for human technological choice, and we evaluate the circumstances that would favor the use of different small prey hunting technologies in the Upper Paleolithic.     

Streams as Entanglement of Nature and Culture: European Upper Paleolithic River Systems and Their Role as Features of Spatial Organization

Large river valleys have long been seen as important factors to shape the mobility, communication, and exchange of Pleistocene hunter-gatherers. However, rivers have been debated as either natural entities people adapt and react to or as cultural and meaningful entities people experience and interpret in different ways. Here, we attempt to integrate both perspectives. Building on theoretical work from various disciplines, we discuss the relationship between biophysical river properties and sociocultural river semantics and suggest that understanding a river’s persona is central to evaluating its role in spatial organization. By reviewing the literature and analyzing European Upper Paleolithic site distribution and raw material transfer patterns in relation to river catchments, we show that the role of prominent rivers varies considerably over time. Both ecological and cultural factors are crucial to explaining these patterns. Whereas the Earlier Upper Paleolithic record displays a general tendency toward conceiving rivers as mobility guidelines, the spatial consolidation process after the colonization of the European mainland is paralleled by a trend of conceptualizing river regimes as frontiers, separating archaeological entities, regional groups, or local networks. The Late Upper Paleolithic Magdalenian, however, is characterized again by a role of rivers as mobility and communication vectors. Tracing changing patterns in the role of certain river regimes through time thus contributes to our growing knowledge of human spatial behavior and helps to improve our understanding of dynamic and mutually informed human-environment interactions in the Paleolithic.     

Mobility patterns and core technologies in the Middle Paleolithic of the Levant

Mobility is thought to be a significant source of Middle Paleolithic archaeological variability in the East Mediterranean Levant. However, models of Levantine Middle Paleolithic land-use have historically been based on rare and taphonomically sensitive evidence from a limited number of sites. Because lithic artifacts are the most ubiquitous archaeological remains available to the prehistorian, relationships between stone tool technology and mobility patterns can improve tests of hypotheses about prehistoric land-use strategies. This paper examines variation in Middle Paleolithic mobility strategies in the Levant from the perspective of core technology. A model linking expedient core reduction techniques and decreased mobility is adapted from one developed for late prehistoric contexts in the New World. Incorporating core data from numerous Levantine Middle Paleolithic assemblages, this study tests hypotheses about diachronic change, synchronic geographic variation, and possible hominin behavioral differences in mobility strategies.     

Shifts in Neandertal mobility,technology and subsistence strategies in western France

We propose a reassessment of Neandertal mobility strategies by crossing technological and zooarchaeological data. A broad comparative approach to the Middle Paleolithic series from western France shows that the Levallois and laminar flaking systems, the Mousterian of Acheulian Tradition (MTA) shaping system and the Quina and discoidal-denticulate flaking systems, vary significantly in terms of duration of reduction sequences, blank versatility and tool maintenance. These technological systems, which prevail in this context over different time periods, reflect distinct mobility strategies as a response to differing hunting practices. This new approach to Middle Paleolithic technologies and related mobility patterns gives new insights into Mousterian diversity. It also highlights the determinant role played by large game hunting strategies in the organization of late Neandertal societies.     

The Upper Paleolithic in Mediterranean Spain: A Review of Current Evidence

In spite of an active and sophisticated archaeological research program, the Paleolithic of the Iberian peninsula remains comparatively little known to English-speaking prehistorians, with the exception of Cantabrian Spain. The rich data set compiled by Spanish prehistorians and their colleagues over the past several decades stands to make a valuable and unique contribution to our understanding of the Pleistocene prehistory of Europe. We present a detailed overview of Upper Paleolithic chronology, sites, and assemblages for Mediterranean Spain, an area of over 1,600,000 km ² that extends from the French border to the Straits of Gibraltar. To interpret these data, we employ a regional perspective that emphasizes studies of paleoeconomy (especially zooarchaeology) and settlement. The Middle–Upper Paleolithic transition and Upper Paleolithic art also receive detailed treatment, and the Upper Paleolithic of Mediterranean Spain is discussed in the broader context of the late Upper Pleistocene of western Europe and the Mediterranean Basin.     

Taphonomic analysis of leporid remains obtained from modern Iberian lynx (Lynx pardinus) scats

Distinguishing leporid bones accumulated by diurnal raptors, owls, mammals and humans is essential to understand not only past human subsistence activities but also past ecology. This is particularly relevant in Iberian Palaeolithic sites where leporid remains usually constitute the most abundant taxon. As far as terrestrial mammal carnivores are concerned, the Iberian lynx (Lynx pardinus) would be the main leporid predator in the Iberian Peninsula. Up until now its contribution as a rabbit bone accumulator in archaeological sites has never been properly assessed. In order to investigate the taphonomic signature of this predator the analysis of 33 samples of modern Iberian lynx scats was carried out. Results show that leporid remains consumed by this carnivore appear very fragmented and exhibit a strong degree of corrosion digestion. Cranial elements are most abundant followed by long bones. Whereas there seem to be clear differences in relation to breakage and digestion alteration caused by owls, some of the patterns obtained are closest to those established with diurnal raptors.     

The Middle Paleolithic from Mujina Pećina,Dalmatia, Croatia

Abstract

Mujina Pe?ina in Dalmatia, Croatia is the only systematically excavated and well-dated Middle Paleolithic cave site in the eastern Adriatic. Its stratigraphic sequence covers the period between 45 and 39 uncalibrated years ka. Results of sedimentological, lithic, faunal, and paleobotanical analyses of the upper levels (Levels B, C, D1, and D2) show that this was a hunting site only sporadically visited by humans. During these occupation episodes humans processed animal remains (mostly large bovids, cervids, and caprids) and produced tools from locally available raw material. Carnivore use of the cave is also indicated. This work contributes to our understanding of lifestyles and mobility patterns of the Middle Paleolithic inhabitants of this region.     

Neanderthal settlement patterns in Crimea: A landscape approach

Traditional settlement pattern analysis involves classifying sites within a region using previously established functional categories. Faunal and lithic data are frequently combined as archaeological indicators of site function. The results are interpreted in terms of a settlement system from which inferences about mobility patterns and social organization are drawn. The previously published site of Karabi Tamchin is used here as a spring-point for a discussion of the problems inherent with established site typologies and some of the settlement models that have been proposed in the past. The rich Middle Paleolithic archaeological record of Crimea is used to demonstrate the usefulness of the landscape approach as a means of supplementing existing information about Middle Paleolithic settlement patterns.     

Changes in the size and shape of fallow deer—evidence for the movement and management of a species

The European fallow deer (Dama dama dama) is native to the eastern Mediterranean and whilst it is clear that its dispersion from this region was the result of human transportation, the timing and circumstances of its post‐glacial diffusion are still uncertain. Archaeological fallow deer remains offer perhaps the best opportunity to understand the deep history of the species, with measurements of ancient bones providing important information about an individual's sex and size. Unfortunately, the fragmentary nature of archaeological remains means that metrical samples are usually too small to draw any meaningful conclusions. Biometrical scaling techniques can increase the size of the sample available for comparison and in this paper log ratios are used in combination with traditional metrical analysis to provide new information about archaeological fallow deer populations, in particular highlighting the size variations across time and space and shifts in hunting and management practices. Copyright © 2011 John Wiley & Sons, Ltd.     

Standing at the gates of Europe: Human behavior and biogeography in the Southern Carpathians during the Late Pleistocene

This study presents a behavioral analysis of Middle and Upper Paleolithic lithic assemblages from 14 sites located in the southern Carpathian Mountains. Using a whole assemblage behavioral indicator, we show that the hominins that manufactured those stone tools do not appear to have differed in terms of the flexibility of the mobility strategies they employed to exploit their landscapes. Rather than biological change, we argue that large-scale climate changes are likely more important drivers of behavioral changes during the Late Pleistocene of the region, including during the Middle–Upper Paleolithic transition. These results agree well with the results of studies having employed this methodology in other regions, suggesting that this is a generalized feature of the transition across Eurasia. Recasting the transition as a mainly ecological rather than purely biocultural process allows us to generate new perspectives from which to approach the question of behavioral change during the Late Pleistocene, and ultimately suggests that the process referred to as the ‘Middle–Upper Paleolithic transition’ is essentially a brief segment of a much more extensive process driven by prehistoric human–environment interactions that would culminate in the highly logistical mobility strategies documented throughout the continent at the Last Glacial Maximum.     

Hunting lesions caused by osseous projectile points: experimental results and archaeological implications

The identification of projectile impact traces on archaeological faunal remains is an important issue for understanding prehistoric hunting behavior, especially in the Paleolithic and Mesolithic. From the beginning of the Upper Paleolithic in Europe, and earlier in Africa, projectiles tipped with osseous points were of great importance for subsistence; but thus far, no specific experimental reference has been developed to help identify the traces left by these points. In 2003 and 2004 two series of projectile experiments with antler points of Magdalenian design were organized, involving two ox calves and two female fallow deer as targets for bow and spearthrower shooting. The subsequent study demonstrates that positive identifications of impact traces left by osseous points can be made. The observation of 127 impact traces allowed us to distinguish three main types of traces: notches, punctures and perforations. The relationships between the nature of the impact traces and the (i) target species, (ii) characteristics of the impacted bones, and (iii) type of weapon are presented. Synthesized results are then discussed within the context of the European Upper Paleolithic and Mesolithic.     

Animal Bones from the Late Neolithic Site of Ponte da Azambuja 2 (Évora,Portugal)

Neolithic faunal assemblages are scarce in Portugal, and although some trends related to the beginning of domestication are now becoming understood, more data and further zooarchaeological analyses are necessary to fully understand them. Ponte da Azambuja 2 is a set of three Late Neolithic ditch enclosures located in the Alentejo region, Southern Portugal. It was discovered in 2008 by the archaeological unit Crivarque during the construction of a new irrigation system. Ditch enclosure 1 was excavated in two trenches, Locus 1 and Locus 2. A small faunal assemblage was recovered, showing high fragmentation mainly due to post‐depositional processes, which made the interpretation rather problematic. Although animal herding strategies are difficult to assess, it was possible to confirm the domesticate status of some of the taxa. Domestication appears to have been complemented by hunting – both red deer and leporids being present. The identification of burning, cut marks and worked bone confirms an anthropogenic faunal accumulation. Worked bone fragments suggest needle production and the production of potential ornaments. Copyright © 2016 John Wiley & Sons, Ltd.     

The Enisei River of Central Siberia in the Late Pleistocene

Recent Paleolithic work along the middle Enisei River of central Siberia has revealed a long history of occupation that almost certainly begins in the Middle Pleistocene. Although the evidence for the Lower Paleolithic is somewhat tentative, there is good reason to believe that hunter-gatherers had periodically occupied the middle Enisei before the last interglacial. The steppe environment of the region during the Upper Pleistocene was relatively bountiful; more than 200 Upper Paleolithic sites, both before and after the Last Glacial Maximum, have been located. The region appears to have been abandoned during the Last Glacial Maximum. Most of Soviet and Russian archaeological work has been guided by a cultural–historical orientation, but recently there has been increased interest in developing adaptationist and ecological research strategies. The middle Enisei and the wider central Siberian region are key to understanding early adaptations to the north and the dimensions of Paleolithic population movements.