Fossil wood of the Mimosoideae from the early Paleocene of Patagonia,Argentina

Brea, M., Zamuner, A.B., Matheos, S.D., Iglesias, A. & Zucol, A.F., December, 2008. Fossil wood of the Mimosoideae from the early Paleocene of Patagonia, Argentina. Alcheringa 32, 427–441. ISSN 0311-5518.

An anatomically preserved mature stem from the Salamanca Formation (early Paleocene) at Palacio de Los Loros, central Patagonia, Argentina, is described and assigned to Paracacioxylon frenguellii sp. nov. The material was preserved by siliceous permineralization and shows features of the secondary xylem typical of subfamily Mimosoideae. This species represents the oldest record of the genus and of the Leguminosae along the western border of Gondwana, and is the world's second oldest record of Leguminosae wood. The species is characterized by ring-porous to semi-ring-porous vessels that are solitary, in multiples of 2–4 and clustered, simple perforation plates, alternate and vestured inter-vessel pitting, homocellular 1–6 seriate rays, tyloses, crystals and diffuse apotracheal, vasicentric paratracheal and confluent axial parenchyma. Paracacioxylon frenguellii has anatomical similarities to Acacia Miller. The presence of Paracacioxylon frenguellii associated with pulvinate leaves suggests that the legumes might have been a component of mesothermal forests developed along the western margin of the Golfo San Jorge Basin during the early Paleocene.     

The United Nations as the ‘Parliament of Man’

Abstract

Paul Kennedy. The Parliament of Man: The Past, Present, and Future of the United Nations. Toronto, ON: HarperCollins, 2006. Pp. xvii, 361. $36.95 (CDN); Ronald St John Macdonald and Douglas M. Johnston, eds. Towards World Constitutionalism: Issues in the Legal Ordering of the World Community. Leiden: Brill, 2005. Pp. xviii, 968. €235.00; $317.00 (US); S. Neil MacFarlane and Yuen Foong Khong. Human Security and the UN: A Critical History. Bloomington and Indianapolis, IN: Indiana University Press, 2006. Pp. xix, 346. $35.00 (US), paper; David M. Malone. The International Struggle over Iraq: Politics in the UN Security Council, 1980–2005. New York, NY: Oxford University Press, 2006. Pp. xiv, 398. $59.95 (CDN); Michael J. Matheson. Council Unbound: The Growth of UN Decision Making on Conflict and Postconflict Issues after the Gold War. Washington, DC: United States Institute of Peace Press, 2006. Pp. xvi, 422. $19.95 (US), paper; Ramesh Thakur. The United Nations, Peace, and Security: From Collective Security to the Responsibility to Protect. New York, NY: Cambridge University Press, 2006. Pp. xvi, 388. $32.99 (US), paper.     

Fixation of the type species of the genus Protoretepora de Koninck, 1878 (Bryozoa,Fenestrata)

Wyse Jackson, P.N., Reid, C.M. & McKinney, F.K., iFirst article, 2011. Fixation of the type species of the genus Protoretepora de Koninck, 1878 (Bryozoa, Fenestrata). Alcheringa, 1–2. ISSN 0311-5518.

The type species of the Palaeozoic bryozoan genus Protoretepora de Koninck, 1878 was originally fixed as Fenestella ampla Lonsdale in Darwin, 1844, but this taxon has been shown to belong to the bryozoan genus Parapolypora Morozova & Lisitsyn, 1996 Morozova, I. P. and Lisitsyn, D. V. 1996. Revision of the genus Polypora. Paleontologicheskii Zhurnal, 1996(4): 38–47. [English translation: Paleontological Journal30(5), 530–541] [Google Scholar]. The original type species designation for Protoretepora de Koninck, 1878 is set aside, and in accordance with Article 70.3 of the International Code of Zoological Nomenclature (4th edition, 1999) the nominal species Protoretepora crockfordae Wyse Jackson, Reid & McKinney, 2011 from the Permian of Tasmania, Australia is herein fixed as the type species.     

A new Late Silurian plant with complex branching from Xinjiang,China

Wang Yi, Fu Qiang, Xu Honghe, & Hao Shougang, June, 2007. A new Late Silurian plant with complex branching from Xinjiang, China. Alcheringa 31, 111-120. ISSN 0311-5518.

A new fossil plant is described from the middle part of the Wutubulake Formation (late Pridoli) of Xinjiang, China. This plant demonstrates at least two orders of branching. The first-order axis has pseudomonopodial branching with alternately attached second-order axes. Fertile units are alternately inserted along the second-order axis, and consist of a branching system and two sporangia at each tip. Sporangia are narrowly obovate with rounded apex and tapering base. This plant is characterized by more complex branching than other Silurian and Early Devonian plants, and is named Wutubulaka multidichotoma gen. et sp. nov., and placed under open higher-order nomenclature.     

A new Late Permian brachiopod fauna from Tanggula,Qinghai-Tibet Plateau and its palaeogeographical implications

Weihong He, Jianjun Bu, Zhijun Niu & Yang Zhang, June, 2009. A new Late Permian brachiopod fauna from Tanggula, Qinghai-Tibet Plateau and its palaeogeographical implications. Alcheringa 33, 113–132. ISSN 0311-5518.

A brachiopod fauna described from the Late Permian of the Gongri and Lizai villages, northwest of the Qoima Co Lake, Tanggula area, southern Qinghai, northwest China, includes ten species in nine genera. This fauna from the (Qiangtang Block) has a strong affinity to coeval faunas of South China, sharing 74% of its species. In addition, the Late Permian Tanggula brachiopod assemblages also demonstrate a clear link with the Middle Permian brachiopod faunas of neighbouring microcontinents including the Indochina block (Malaysia and Cambodia) and the Shan-Thai block (Thailand), as suggested by the presence of Caricula sp. cf. C. salebrosa, Transennatia termierorum and Strophalosiina. This phenomenon indicates that there were relatively narrow seaways between these microcontinents to enable ready interchange of brachiopods during the Permian, and that brachiopods tended to retreat towards the palaeoequatorial region throughout the period.     

A new dragonfly,Austroprotolindenia jurassica (Odonata: Anisoptera), from the Upper Jurassic of Australia

Beattie, R.G. & Nel, A., June 2012. A new dragonfly, Austroprotolindenia jurassica (Odonata: Anisoptera), from the Upper Jurassic of Australia. Alcheringa, 189–193. ISSN 0311-5518.

Austroprotolindenia jurassica gen. et sp. nov., a new Mesozoic Australian dragonfly, is described from the Talbragar Fossil Fish Bed (Upper Jurassic) of eastern Australia. It shows some similarities with the Eurasian Mesozoic petalurid family Protolindeniidae, but its incomplete state of preservation prevents us assigning it to a particular anisopteran clade.

Robert G. Beattie [rgbeattie@bigpond.com] PO Box 320, Berry 2535, NSW, Australia. André Nel [anel@mnhn.fr] CNRS UMR 7205, CP 50, Entomologie, Muséum National d'Histoire Naturelle, 45 rue Buffon, F-75005 Paris, France. Received 6.4.2011; revised 8.6.2011; accepted 15.6.2011.     

Editorial

Kear, Benjamin P., 11 November 2019. Editorial. Alcheringa 43, XX–XX. ISSN 0311-5518.

Benjamin P. Kear [benjamin.kear@em.uu.se], Museum of Evolution, Uppsala University, 752 36 Uppsala, Sweden.     

New Late Permian rugose corals from Pahang,peninsular Malaysia

Aye Ko Aung, Ng Tham Fatt, Kyaw Kyaw Nyein & Myo Htut Zin, 2013. New Late Permian rugose corals from Pahang, peninsular Malaysia. Alcheringa 37, 422–434. ISSN 0311-5518.

Late Permian rugose corals are described from a limestone unit of the Gua Musang Formation at Selborne Estate, Padang Tengku area, Pahang, peninsular Malaysia. These include one genus, Iranophyllum, which is reported for the first time from Malaysia, with two new species Iranophyllum aequabilis and I. pahangense belonging to Waagenophyllidae. A Late Permian age is confirmed by a Paleofusulina–Colaniella–Reichelina foraminiferal fauna co-preserved with the corals.

Aye Ko Aung [akaung.mm@gmail.com], Ng Tham Fatt [thamfatt@gmail.com], Kyaw Kyaw Nyein [konyein@gmail.com], Department of Geology, University of Malaya, 50603, Kuala Lumpur, Malaysia and Myo Htut Zin [myohtutgreat@googlemail.com], Lab. Services, Pte. Co. Ltd., Singapore. Received 16.10.2012; revised 5.1.2013; accepted 17.1.2012.     

Emwadea microcarpa gen. et sp. nov.—anatomically preserved araucarian seed cones from the Winton Formation (late Albian), western Queensland,Australia

Dettmann, M.E., Clifford, H.T., Peters, M., June 2012. Emwadea microcarpa gen. et sp. nov.—anatomically preserved araucarian seed cones from the Winton Formation (late Albian), western Queensland, Australia. Alcheringa, 217–237. ISSN 0311-5518.

A new genus and species, Emwadea microcarpa Dettmann, Clifford & Peters, is established for ovulate/seed cones with helically arranged cone scales bearing a centrally positioned, inverted ovule from the basal Winton Formation (late Albian), Eromanga Basin, Queensland. The cones are small, prolate ellipsoidal (9.5–14 mm vertical axis, 6.3–8.7 mm transverse axis) with wedge-shaped cone scales bearing winged seeds attached adaxially to the scale only by tissues surrounding the vasculature entering the ovule. Ovuliferous tissue that is free from the cone scale extends distally from the chalaza; the seeds' lateral wings are derived from the integument. Foliage attached to the cones is spirally arranged, imbricate and with spreading and incurved bifacial blades with acute tips; stomata are arranged in longitudinal files and are confined to the adaxial surface. The cone organization testifies to placement within the Araucariaceae, and is morphologically more similar to Wollemia and Agathis than to Araucaria.

Mary Dettmann [mary.dettmann@qm.qld.gov.au] and Trevor Clifford, Queensland Museum, PO Box 3300, South Brisbane, Q 4101, Australia; Mark Peters, PO Box 366 Gumeracha, SA 5233, Australia. Received 31.3.2011; revised 23.8.2011; accepted 5.9.2011.

     

A Multi-Method Approach to Reconstructing Occupational History and Activity Areas: A Case Study at the Formative Site of Tlalancaleca,Central Mexico

While surface collection constitutes an important component of field research at the site and regional levels, the association between surface and sub-surface materials needs to be examined through independent lines of evidence. This study employed manual bucket auger probing and soil geochemistry at the Formative site of Tlalancaleca, Central Mexico (800 b.c.–a.d. 250), to evaluate the results of surface collection and to understand the formation process of the anthropogenic landscape. The combined results at one of the largest architectural complexes at Tlalancaleca indicate that it was built during the Late Formative period (500–100 b.c.) and rebuilt during the subsequent Terminal Formative period (100 b.c.–a.d. 250), which is corroborated by radiocarbon dates. Moreover, the results suggest segregated uses of space such as cooking/storage, waste disposal, and craft production areas within the complex. Overall results demonstrate the effectiveness of this multi-method approach for reconstructing occupational history and activity areas.     

The Regional History of the Guan River Valley,China: Cultural Periphery,Ecological Diversity,and the Development of Social Complexity before A.D. 220

Regional settlement pattern studies in China have overlooked interregional interaction in the development of social complexity. This study examines settlement patterns, land use practices, sociopolitical structure, and interregional interaction from 4000 b.c. to the early Imperial period (before a.d. 221) in the Guan River valley, China. The Guan valley is located at the inter-montane corridor to three cultural core zones in China. An international collaborative team used regional full-coverage survey to locate 96 sites in 135?km² in the Middle Guan valley, the earliest of which dated to the middle Yangshao period (4000–3500 b.c.). In the late Yangshao period (3500–2900 b.c.), occupation increased rapidly and expanded to the upper reach and tributaries of the river. After an occupational collapse during the Early States period (1900–771 b.c.), the population recovered rapidly and reached a new level of organizational complexity in the Eastern Zhou period (770–221 b.c.). This pattern continued into the Qin-Han period (220 b.c.–a.d. 220), when hamlets increased in number and expanded into hilly areas.     

Late Prehistoric High-Altitude Hunter-Gatherer Residential Occupations in the Argentine Southern Andes

Test excavations were conducted at Risco de los Indios (RDLI), a site at 2480 masl with 29 residential features and a well-developed midden containing abundant floral, faunal, lithic, and ceramic materials. Analyses indicate the site was intensively used ca. 500 cal b.p. as a residential base for groups focused on hunting guanaco, supplemented by locally-available wild flora and fauna as well as domestic beans transported from the lowlands. Ceramic and obsidian artifacts indicate these groups were highly mobile and in contact with groups on the eastern and western margins of the Andes. These patterns compare favorably to those seen in the region’s other high altitude villages. It appears that the development of these patterns began with population increase and economic intensification in the lowlands ca. 2000 cal b.p. and that the move to slightly lower elevation settings like RDLI may have been conditioned by the onset of the Little Ice Age.     

A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales,Australia)

?erňanský, A. & Hutchinson, M.N., 2012. A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales, Australia). Alcheringa, 1–6. ISSN 0311-5518.

We describe an isolated frontal bone referable to a new species, Tiliqua laticephala (Scincidae), from the Pliocene Big Sink doline of the Wellington Caves, central eastern New South Wales, Australia. The bone is very robust, is unusually broad and thick, especially around the bases of the subolfactory processes and represents a large and heavily built lizard. The fossil has multiple fragmentary osteoderms in the frontal region, showing asymmetry in shape and thickness that do not correspond to the more regularly arranged anterior head shields of other scincoids. The specimen shares two unusual character states with extant Tiliqua, especially the large armoured species, T. rugosa. Other large skinks related to T. rugosa (other Tiliqua spp., Corucia zebrata, Egernia cunninghamii, Bellatorias major, Liopholis kintorei) are less similar in terms of frontal shape, thickness, sculpture, osteoderm ornamentation, and positioning of adjacent bones.     

On the integration of Ordovician conodont and graptolite biostratigraphy: new examples from Gansu and Inner Mongolia in China

Wang, Z.H., Bergström, S.M., Zhen, Y.Y., Chen, X. & Zhang, Y.D., 2013. On the integration of Ordovician conodont and graptolite biostratigraphy: New examples from Gansu and Inner Mongolia in China. Alcheringa 37, 510–528. ISSN 0311-5518.

Few Ordovician successions in the world contain both biostratigraphically highly diagnostic conodonts and graptolites permitting an integration between standard biozones based on these fossil groups. The Sandbian Guanzhuang section in the vicinity of Pingliang in the Gansu Province has an outstanding graptolite record through most of the Nemagraptus gracilis and Climacograptus bicornis graptolite biozones. Calcareous interbeds in the succession yield biostratigraphically important conodonts, including some species used for biozonations in Baltoscandia and the North American Midcontinent. Likewise, the middle–upper Darriwilian Dashimen section in the Wuhai region of Inner Mongolia hosts both diverse graptolites of the Pterograptus elegans, Didymograptus murchisoni and lowermost Nemagraptus gracilis biozones, and conodonts of Midcontinent and Baltoscandic types. The distribution patterns of these index fossil groups provide an unusual opportunity to closely correlate conodont and graptolite biozones in the middle to upper Darriwilian to Sandbian interval. For instance, the base of the C. bicornis Biozone is approximately coeval with the base of the Baltoscandic B. gerdae Subbiozone and a level near the middle of the North American P. aculeata Biozone.

Zhi-hao Wang [zhwang@nigpas.ac.cn] Xu Chen [xu1936@gmail.com], and Yuan-dong Zhang [ydzhang@nigpas.ac.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; Stig M. Bergström [stig@geology.ohio-state.edu], School of Earth Sciences, Division of Earth History, The Ohio State University, Columbus, OH 43210, USA; Yong Yi Zhen [yongyi.zhen@austmus.gov.au], Australian Museum, 6 College Street, Sydney NSW 2010, Australia.     

Pteruchus (microsporophyll): part 2 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed

Anderson, H.M., Barbacka, M., Bamford, M.K., Holmes, W.B.K. & Anderson, J.M., 3 July 2019. Pteruchus (microsporophyll): part 2 of a reassessment of Gondwana Triassic plant genera and a reclassification of some attributed previously. Alcheringa XXX, X–X. ISSN 0311-5518

The microsporophyll genus Pteruchus, belonging to the pteridosperms (seed ferns) in the family Umkomasiaceae (Corystospermaceae), is reassessed comprehensively worldwide and emended. All records are analysed, and some fertile structures previously attributed are reclassified. The Lower Jurassic record of Pteruchus from Germany is ascribed to a new genus as Muelkirchium septentrionalis. Pteruchus is shown to be restricted to the Triassic of Gondwana and is clearly affiliated with the megasporophyll genus Umkomasia and the vegetative leaf genus Dicroidium. It is well represented from Argentina, Antarctica, Australia and southern Africa; the Molteno Formation of southern Africa is by far the most comprehensively sampled, yielding three species and 425 specimens from 22 localities. Nomenclatural problems with the species of Pteruchus are addressed. A key to Pteruchus species is provided; geographic and stratigraphic distributions are tabulated.

Heidi M. Anderson [hmsholmes@googlemail.com], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa; Maria Barbacka [mariabarbacka@gmail.com], W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, 31-512 Kraków, Poland; Hungarian Natural History Museum, Botanical Department, H-1431 Budapest, Pf. 137, Hungary; Marion K. Bamford [marionbamford@witsacza], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa; W.B. Keith Holmes* [wbkholmes@hotmail.com], 46 Kurrajong Street, Dorrigo, NSW 2453, Australia; John M. Anderson [jmandersongondwana@googlemail.com], Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg 20150, South Africa. *Also affiliated with: University of New England, Armidale, NSW 2351, Australia.     

Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia

Benson, R.B.J., Fitzgerald, E.M.G., Rich, T.H. & Vickers-Rich, P., 2013. Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia. Alcheringa 37, 1–6. ISSN 0311-5518

We report a large plesiosaurian tooth from the freshwater early–middle Aptian (Early Cretaceous) Eumeralla Formation of Victoria, Australia. This, combined with records of smaller plesiosaurian teeth with an alternative morphology, provides evidence for a multitaxic freshwater plesiosaurian assemblage. Dental and body size differences suggest ecological partitioning of sympatric freshwater plesiosaurians analogous to that in modern freshwater odontocete cetaceans. The evolutionarily plastic body plan of Plesiosauria may have facilitated niche differentiation and helped them to exclude ichthyosaurs from freshwater environments during the Mesozoic. However, confirmation of this hypothesis requires the discovery of more complete remains.

Roger B.J. Benson [roger.benson@earth.ox.ac.uk], Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK; Erich M.G. Fitzgerald [efitzgerald@museum.vic.gov.au], Thomas H. Rich [trich@museum.vic.gov.au], Museum Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia; Thomas H. Rich and Patricia Vickers-Rich [pat.rich@monash.edu], School of Geosciences, Monash University, Clayton, Victoria 3800, Australia. Received 30.10.2012; revised 27.1.2013; accepted 31.1.2013.     

A white‐hot liquid earth and geological time

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The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri

Rich, T.H., Hopson, J.A., Gill, P.G., Trusler, P., Rogers-Davidson, S., Morton, S., Cifelli, R.L., Pickering, D., Kool, L., Siu, K., Burgmann, F.A., Senden, T., Evans, A.R., Wagstaff, B.E., Seegets-Villiers, D., Corfe, I.J., Flannery, T.F., Walker, K., Musser, A.M., Archer, M., Pian, R. & Vickers-Rich, P., June 2016. The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri. Alcheringa 40, xx–xx. ISSN 0311-5518.

The monotreme Teinolophos trusleri Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren, 1999 Rich, T.H., Vickers-Rich, P., Constantine, A., Flannery, T.F., Kool, L. & van Klaveren, N., 1999. Early Cretaceous mammals from Flat Rocks, Victoria, Australia. Records of the Queen Victoria Museum and Art Gallery 106, 1–34. [Google Scholar] from the Early Cretaceous of Australia is redescribed and reinterpreted here in light of additional specimens of that species and compared with the exquisitely preserved Early Cretaceous mammals from Liaoning Province, China. Together, this material indicates that although T. trusleri lacked a rod of postdentary bones contacting the dentary, as occurs in non-mammalian cynodonts and basal mammaliaforms, it did not share the condition present in all living mammals, including monotremes, of having the three auditory ossicles, which directly connect the tympanic membrane to the fenestra ovalis, being freely suspended within the middle ear cavity. Rather, T. trusleri appears to have had an intermediate condition, present in some Early Cretaceous mammals from Liaoning, in which the postdentary bones cum ear ossicles retained a connection to a persisting Meckel’s cartilage although not to the dentary. Teinolophos thus indicates that the condition of freely suspended auditory ossicles was acquired independently in monotremes and therian mammals. Much of the anterior region of the lower jaw of Teinolophos is now known, along with an isolated upper ultimate premolar. The previously unknown anterior region of the jaw is elongated and delicate as in extant monotremes, but differs in having at least seven antemolar teeth, which are separated by distinct diastemata. The dental formula of the lower jaw of Teinolophos trusleri as now known is i2 c1 p4 m5. Both the deep lower jaw and the long-rooted upper premolar indicate that Teinolophos, unlike undoubted ornithorhynchids (including the extinct Obdurodon), lacked a bill.

Thomas H. Rich [trich@museum.vic.gov.au], Sally Rogers-Davidson [srogers@museum.vic.gov.au], David Pickering [dpick@museum.vic.gov.au], Timothy F. Flannery [tim.flannery@textpublishing.com.au], Ken Walker [kwalker@museum.vic.gov.au], Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; James A. Hopson [jhopson@uchicago.edu], Department of Organismal Biology & Anatomy, University of Chicago,1025 East 57th Street, Chicago, IL 60637, USA; Pamela G. Gill [pam.gill@bristol.ac.uk], School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, U.K. and Earth Science Department, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Peter Trusler [peter@petertrusler.com.au], Lesley Kool [koollesley@gmail.com], Doris Seegets-Villiers [doris.seegets-villiers@monash.edu], Patricia Vickers-Rich [pat.rich@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Victoria 3800, Australia; Steve Morton [steve.morton@monash.edu], Karen Siu [karen.siu@monash.edu], School of Physics and Astronomy, Monash University, Victoria 3800, Australia; Richard L. Cifelli [rlc@ou.edu] Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, OK 73072, USA; Flame A. Burgmann [flame.burgmann@monash.edu], Monash Centre for Electron Microscopy, 10 Innovation Walk, Monash University, Clayton, Victoria 3800, Australia; Tim Senden [Tim.Senden@anu.edu.au], Department of Applied Mathematics, Research School of Physical Sciences and Engineering, The Australian National University, Canberra, Australian Capital Territory 0200, Australia; Alistair R. Evans [alistair.evans@monash.edu], School of Biological Sciences, Monash University, Victoria 3800, Australia; Barbara E. Wagstaff [wagstaff@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Ian J. Corfe [ian.corfe@helsinki.fi], Institute of Biotechnology, Viikinkaari 9, 00014, University of Helsinki, Finland; Anne M. Musser [anne.musser@austmus.gov.au], Australian Museum, 1 College Street, Sydney NSW 2010 Australia; Michael Archer [m.archer@unsw.edu.au], School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia; Rebecca Pian [rpian@amnh.org], Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA. Received 7.4.2016; accepted 14.4.2016.