The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri

Rich, T.H., Hopson, J.A., Gill, P.G., Trusler, P., Rogers-Davidson, S., Morton, S., Cifelli, R.L., Pickering, D., Kool, L., Siu, K., Burgmann, F.A., Senden, T., Evans, A.R., Wagstaff, B.E., Seegets-Villiers, D., Corfe, I.J., Flannery, T.F., Walker, K., Musser, A.M., Archer, M., Pian, R. & Vickers-Rich, P., June 2016. The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri. Alcheringa 40, xx–xx. ISSN 0311-5518.

The monotreme Teinolophos trusleri Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren, 1999 Rich, T.H., Vickers-Rich, P., Constantine, A., Flannery, T.F., Kool, L. & van Klaveren, N., 1999. Early Cretaceous mammals from Flat Rocks, Victoria, Australia. Records of the Queen Victoria Museum and Art Gallery 106, 1–34. [Google Scholar] from the Early Cretaceous of Australia is redescribed and reinterpreted here in light of additional specimens of that species and compared with the exquisitely preserved Early Cretaceous mammals from Liaoning Province, China. Together, this material indicates that although T. trusleri lacked a rod of postdentary bones contacting the dentary, as occurs in non-mammalian cynodonts and basal mammaliaforms, it did not share the condition present in all living mammals, including monotremes, of having the three auditory ossicles, which directly connect the tympanic membrane to the fenestra ovalis, being freely suspended within the middle ear cavity. Rather, T. trusleri appears to have had an intermediate condition, present in some Early Cretaceous mammals from Liaoning, in which the postdentary bones cum ear ossicles retained a connection to a persisting Meckel’s cartilage although not to the dentary. Teinolophos thus indicates that the condition of freely suspended auditory ossicles was acquired independently in monotremes and therian mammals. Much of the anterior region of the lower jaw of Teinolophos is now known, along with an isolated upper ultimate premolar. The previously unknown anterior region of the jaw is elongated and delicate as in extant monotremes, but differs in having at least seven antemolar teeth, which are separated by distinct diastemata. The dental formula of the lower jaw of Teinolophos trusleri as now known is i2 c1 p4 m5. Both the deep lower jaw and the long-rooted upper premolar indicate that Teinolophos, unlike undoubted ornithorhynchids (including the extinct Obdurodon), lacked a bill.

Thomas H. Rich [trich@museum.vic.gov.au], Sally Rogers-Davidson [srogers@museum.vic.gov.au], David Pickering [dpick@museum.vic.gov.au], Timothy F. Flannery [tim.flannery@textpublishing.com.au], Ken Walker [kwalker@museum.vic.gov.au], Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; James A. Hopson [jhopson@uchicago.edu], Department of Organismal Biology & Anatomy, University of Chicago,1025 East 57th Street, Chicago, IL 60637, USA; Pamela G. Gill [pam.gill@bristol.ac.uk], School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, U.K. and Earth Science Department, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Peter Trusler [peter@petertrusler.com.au], Lesley Kool [koollesley@gmail.com], Doris Seegets-Villiers [doris.seegets-villiers@monash.edu], Patricia Vickers-Rich [pat.rich@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Victoria 3800, Australia; Steve Morton [steve.morton@monash.edu], Karen Siu [karen.siu@monash.edu], School of Physics and Astronomy, Monash University, Victoria 3800, Australia; Richard L. Cifelli [rlc@ou.edu] Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, OK 73072, USA; Flame A. Burgmann [flame.burgmann@monash.edu], Monash Centre for Electron Microscopy, 10 Innovation Walk, Monash University, Clayton, Victoria 3800, Australia; Tim Senden [Tim.Senden@anu.edu.au], Department of Applied Mathematics, Research School of Physical Sciences and Engineering, The Australian National University, Canberra, Australian Capital Territory 0200, Australia; Alistair R. Evans [alistair.evans@monash.edu], School of Biological Sciences, Monash University, Victoria 3800, Australia; Barbara E. Wagstaff [wagstaff@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Ian J. Corfe [ian.corfe@helsinki.fi], Institute of Biotechnology, Viikinkaari 9, 00014, University of Helsinki, Finland; Anne M. Musser [anne.musser@austmus.gov.au], Australian Museum, 1 College Street, Sydney NSW 2010 Australia; Michael Archer [m.archer@unsw.edu.au], School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia; Rebecca Pian [rpian@amnh.org], Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA. Received 7.4.2016; accepted 14.4.2016.     

A new Miocene carnivorous marsupial,Barinya kutjamarpensis (Dasyuromorphia), from central Australia

Binfield, P., Archer, M., Hand, S.J., Black, K.H., Myers, T.J., Gillespie, A.K. & Arena, D.A., June 2016. A new Miocene carnivorous marsupial, Barinya kutjamarpensis (Dasyuromorphia), from central Australia. Alcheringa 41, xx–xx. ISSN 0311-5518.

A new dasyuromorphian, Barinya kutjamarpensis sp. nov., is described on the basis of a partial dentary recovered from the Miocene Wipajiri Formation of northern South Australia. Although about the same size as the only other species of this genus, B. wangala from the Miocene faunal assemblages of the Riversleigh World Heritage Area, northwestern Queensland, it has significant differences in morphology including a very reduced talonid on M₄ and proportionately wider molars. Based on the structural differences and the more extensive wear on its teeth, the central Australian species might have consumed harder or more abrasive prey in a more silt-rich environment than its congener, which hunted in the wet early to middle Miocene forests of Riversleigh.

Pippa Binfield [pippa.binfield@outlook.com], Michael Archer [m.archer@unsw.edu.au], Suzanne J. Hand [s.hand@unsw.edu.au], Karen H. Black [k.black@unsw.edu.au], Troy J. Myers [t.myers@unsw.edu.au] Anna K. Gillespie [a.gillespie@unsw.edu.au] and Derrick A. Arena [rick@sitc.net.au], PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales 2052, Sydney, Australia.     

Early Cretaceous polar biotas of Victoria,southeastern Australia—an overview of research to date

Poropat, S.F., Martin, S.K., Tosolini, A.-M.P., Wagstaff, B.E, Bean, L.B., Kear, B.P., Vickers-Rich, P. &; Rich, T.H., May 2018. Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date. Alcheringa 42, 158–230. ISSN 0311-5518.

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in Victoria since the 1960s, including the Koonwarra Fossil Bed (Strzelecki Group) near Leongatha, the Dinosaur Cove and Eric the Red West sites (Otway Group) at Cape Otway, and the Flat Rocks site (Strzelecki Group) near Cape Paterson. Systematic exploration over the past five decades has resulted in the collection of thousands of fossils representing various plants, invertebrates and vertebrates. Some of the best-preserved and most diverse Hauterivian–Barremian floral assemblages in Australia derive from outcrops of the lower Strzelecki Group in the Gippsland Basin. The slightly younger Koonwarra Fossil Bed (Aptian) is a Konservat-Lagerstätte that also preserves abundant plants, including one of the oldest known flowers. In addition, insects, crustaceans (including the only syncaridans known from Australia between the Triassic and the present), arachnids (including Australia’s only known opilione), the stratigraphically youngest xiphosurans from Australia, bryozoans, unionoid molluscs and a rich assemblage of actinopterygian fish are known from the Koonwarra Fossil Bed. The oldest known—and only Mesozoic—fossil feathers from the Australian continent constitute the only evidence for tetrapods at Koonwarra. By contrast, the Barremian–Aptian-aged deposits at the Flat Rocks site, and the Aptian–Albian-aged strata at the Dinosaur Cove and Eric the Red West sites, are all dominated by tetrapod fossils, with actinopterygians and dipnoans relatively rare. Small ornithopod (=basal neornithischian) dinosaurs are numerically common, known from four partial skeletons and a multitude of isolated bones. Aquatic meiolaniform turtles constitute another prominent faunal element, represented by numerous isolated bones and articulated carapaces and plastrons. More than 50 specimens—mostly lower jaws—evince a high diversity of mammals, including monotremes, a multituberculate and several enigmatic ausktribosphenids. Relatively minor components of these fossil assemblages are diverse theropods (including birds), rare ankylosaurs and ceratopsians, pterosaurs, non-marine plesiosaurs and a lepidosaur. In the older strata of the upper Strzelecki Group, temnospondyl amphibians—the youngest known worldwide—are a conspicuous component of the fauna, whereas crocodylomorphs appear to be present only in up-sequence deposits of the Otway Group. Invertebrates are uncommon, although decapod crustaceans and unionoid bivalves have been described. Collectively, the Early Cretaceous biota of Victoria provides insights into a unique Mesozoic high-latitude palaeoenvironment and elucidates both palaeoclimatic and palaeobiogeographic changes throughout more than 25 million years of geological time.

Stephen F. Poropat*? [sporopat@swin.edu.au; stephenfporopat@gmail.com], Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Sarah K. Martin*? [sarah.martin@dmirs.wa.gov.au; martin.sarahk@gmail.com] Geological Survey of Western Australia, 100 Plain St, East Perth, Western Australia 6004, Australia; Anne-Marie P. Tosolini [a.tosolini@unimelb.edu.au] and Barbara E. Wagstaff [wagstaff@unimelb.edu.au] School of Earth Sciences, The University of Melbourne, Melbourne, Victoria 3010, Australia; Lynne B. Bean [lynne.bean@anu.edu.au] Research School of Earth Sciences, Australian National University, Acton, Canberra, Australian Capital Territory 2001, Australia; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden; Patricia Vickers-Rich§ [prich@swin.edu.au; pat.rich@monash.edu] Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Thomas H. Rich [trich@museum.vic.gov.au] Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia. *These authors contributed equally to this work. ?Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia. ?Also affiliated with: Earth and Planetary Sciences, Western Australian Museum, Welshpool, Western Australia 6101, Australia. §Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne, Victoria 3800, Australia.     

Utilizing U–Pb CA-TIMS dating to calibrate the Middle to Late Jurassic spore-pollen zonation of the Surat Basin,Australia to the geological time-scale

Wainman, C.C., Hannaford, C., Mantle, D. & McCabe, P.J., April.2018. Utilizing U–Pb CA-TIMS dating to calibrate the Middle to Late Jurassic spore-pollen zonation of the Surat Basin, Australia to the geological time-scale. Alcheringa XX, xx-xx.

Spore-pollen palynostratigraphy is commonly used to subdivide and correlate Jurassic continental successions in eastern Australia and thus aid the construction of geological models for the petroleum and coal industries. However, the current spore-pollen framework has only been tenuously calibrated to the geological time-scale. Age determinations are reliant on indirect correlations of ammonite and dinoflagellate assemblages from New Zealand, the North West Shelf of Australia and Southeast Asia to the standard European stages. New uranium-lead chemical abrasion thermal ionization mass spectrometry (U–Pb CA-TIMS) dates from 19 tuff beds in the Middle–Upper Jurassic Injune Creek Group of the Surat Basin enables regional spore-pollen palynostratigraphic zones to be precisely dated for the first time. These results show the base of the APJ4.2 and APJ4.3 subzones are similar in age to previous estimates (Middle Jurassic, Bathonian) from indirect palynostratigraphic correlation. However, the base of the APJ5 Zone and the APJ6.1 Subzone may be somewhat younger than previously estimated, possibly by as much as 2.5 and 4.2 Myrs, respectively. The continued utilization of U–Pb CA-TIMS dates will further refine the absolute ages of these zones, improve the inter- and intra-basinal correlation of Middle–Upper Jurassic strata in eastern Australian basins and greatly enhance intercontinental correlations.

Carmine Christopher Wainman [carmine.wainman@adelaide.edu.au] and Peter James McCabe [peter.mccabe@adelaide,edu.au] Australian School of Petroleum, University of Adelaide, SA, 5005, Australia; Carey Hannaford [carey.hannaford@mgpalaeo.com.au] and Daniel Mantle [dan.mantle@mgpalaeo.com.au] MGPalaeo Pty Ltd, 5 Arvida Street, Malaga, WA, 6090, WA, Australia.     

A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida

Zhang, Y., He, W.H., Shi, G.R., Zhang, K.X. & Wu, H.T., 26.2.2015. A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida. Alcheringa 39, xxx–xxx. ISSN 0311-5518.

As the third and last part of a systematic palaeontological study of the brachiopod fauna from the Permian–Triassic boundary section at Zhongzhai in Guizhou Province (South China), this paper reports 15 species (including three new species: Tethyochonetes minor sp. nov., Neochonetes (Zhongyingia) transversa sp. nov., Paryphella acutula sp. nov.) in Order Productida. In addition, the morphological features and definitions of several key Changhsingian brachiopod taxa (e.g., Paryphella and Oldhamina interrupta) are clarified and revised.

Yang Zhang* [zhangy05@163.com] and G.R. Shi [guang.shi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; Weihong He [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Huiting Wu [ht_wu415@163.com], Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China. *Also affiliated with: Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

A preliminary report on new Ediacaran fossils from Iran

Vickers-Rich, P., Soleimani, S., Farjandi, F., Zand, M., Linnemann, U., Hofmann, M., Wilson, S.A., Cas, R. &; Rich, T.H. November, 2017. A preliminary report on new Ediacaran fossils from Iran. Alcheringa 42, 231–244. ISSN 0311-5518.

Recent exploratory field mapping of marine sedimentary sequences in the Koushk Mine locality of the Bafq region in Central Iran, and on the northern slopes of the Elborz Mountains south of the Caspian Sea, has yielded large complex body and trace fossils of Neoproterozoic–early Cambrian age. The recovered specimens resemble the previously documented Precambrian discoidal form Persimedusites, and a the tubular morphotype Corumbella, which is a novel occurrence for Iran and otherwise only recorded before from Brazil and the western USA. Additional enigmatic traces can not yet be interpreted unequivocally, but suggest that future work may uncover more unusual Ediacaran fossils from various localities in Central Iran.

Patricia Vickers-Rich* [prich@swin.edu.au, pat.rich@monash.edu], Faculty of Science, Swinburne University of Technology, Melbourne (Hawthorn), Victoria 3122, Australia; Sara Soleimani [sara_soleimani@yahoo.com], Palaeontology Department, Geological Survey of Iran, Tehran, Iran; Farnoosh Farjandi [farnooshfarjandi@gmail.com], Department of Geochemical Exploration, Geological Survey of Iran, Tehran, Iran; Mehdi Zand [zand.mehdi.geo@gmail.com], Geology Department, Bafq Mining Company, Koushk Mine, Yazd, Iran. Ulf Linnemann [ulf.linnemann@senckenberg.de], and Mandy Hofmann [mandy.hofmann@senckenberg.de], Senckenberg Naturhistorische Sammlungen, Dresden, Museum für Mineralogie und Geologie, Sektion Geochronologie, Koenigsbruecker Landstrasse 159, D-01109, Dresden, Germany; Siobhan A. Wilson [sasha.wilson@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Raymond Cas [ray.cas@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Thomas H. Rich? [trich@museum.vic.gov.au], Museum Victoria, Exhibition Gardens, PO Box 666, Melbourne, Victoria, 3001 Australia. *Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; School of Environmental Sciences, Deakin University, Melbourne (Burwood), Victoria, Australia 3125; Palaeontology Department, Museum Victoria, Carlton Gardens, PO Box 666, Melbourne, Victoria 3001, Australia. ?Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Faculty of Science, Swinburne University of Science and Technology, Melbourne (Hawthorn), Victoria 3122, Australia.     

An Early Permian brachiopod–gastropod fauna from the Calytrix Formation,Barbwire Terrace,Canning Basin,Western Australia

Taboada, A.C., Mory, A.J., Shi, G.R., Haig, D.W. & Pinilla, M.K., 12.11.2014. An Early Permian brachiopod–gastropod fauna from the Calytrix Formation, Barbwire Terrace, Canning Basin, Western Australia. Alcheringa 39, xxx–xxx. ISSN 0311-5518

A small brachiopod–gastropod fauna from a core close to the base of the Calytrix Formation within the Grant Group includes the brachiopods Altiplecus decipiens (Hosking), Myodelthyrium dickinsi (Thomas), Brachythyrinella narsarhensis (Reed), Neochonetes (Sommeriella) obrieni Archbold, Tivertonia barbwirensis sp. nov. and the gastropod Peruvispira canningensis sp. nov. The fauna has affinities with that of the late Sakmarian?early Artinskian Nura Nura Member directly overlying the Grant Group in other parts of the basin but, as with all lower Cisuralian (and Pennsylvanian) glacial strata in Western Australia, its precise age remains poorly constrained, especially in terms of correlation to international stages. Although the Calytrix fauna lies within the Pseudoreticulatispora confluens Palynozone, the only real constraint on its age (and that of the associated glacially influenced strata) is from Sakmarian (Sterlitamakian) and stratigraphically younger faunas. A brief review of radiometric ages from correlative strata elsewhere in Gondwana shows that those ages need to be updated. The presence of Asselian strata and the position of the Carboniferous?Permian boundary remain unclear in Western Australia.

Arturo César Taboada [ataboada@unpata.edu.ar], CONICET-Laboratorio de Investigaciones en Evolución y Biodiversidad (LIEB), Facultad de Ciencias Naturales, Sede Esquel, Universidad Nacional de la Patagonia ‘San Juan Bosco’, Edificio de Aulas, Ruta Nacional 259, km. 16,5, Esquel U9200, Chubut, Argentina; Arthur Mory [arthur.mory@dmp.wa.gov.au], Geological Survey of Western Australia, 100 Plain Street, East Perth, WA 6004, School of Earth and Environment, The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia; Guang R. Shi [grshi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; David W. Haig [david.haig@uwa.edu.au], School of Earth and Environment (M004), The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia; María Karina Pinilla [mkpinilla@fcnym.unlp.edu.ar], División Paleozoología Invertebrados, Museo de Ciencias Naturales de La Plata, Paseo del Bosque s/n, 1900 La Plata, Buenos Aires, Argentina.     

New material referable to Wakaleo (Marsupialia: Thylacoleonidae) from the Riversleigh World Heritage Area,northwestern Queensland: revising species boundaries and distributions in Oligo/Miocene marsupial lions

Gillespie, A.K., Archer, M., Hand, S.J. & Black, K.H., 2014. New material referable to Wakaleo (Marsupialia: Thylacoleonidae) from the Riversleigh World Heritage Area, northwestern Queensland: revising species boundaries and distributions in Oligo/Miocene marsupial lions. Alcheringa 38, 513–527. ISSN 0311–5518.

New material of Wakaleo oldfieldi and W. vanderleueri from the Miocene freshwater limestones of the Riversleigh World Heritage Area, northwestern Queensland, is described. This material includes the first known upper dentition of W. oldfieldi and dentaries of both species bearing the previously undescribed and morphologically distinct M₃. Previously, the two species were distinguished only by size differences in P3 and the size of P₃ relative to M1. Wakaleo oldfieldi exhibits a more plesiomorphic M₃ that retains a well-developed talonid basin in contrast to W. vanderleueri, which has lost this structure. The phyletic succession and geological occurrences of Wakaleo species make this genus an important taxon in biochronological analyses of Australian Cenozoic assemblages. At Riversleigh, W. oldfieldi is found in deposits allocated to Faunal Zone B and Faunal Zone C, which are regarded as early and middle Miocene in age, respectively. The presence of this species in the Kutjamarpu Local Fauna of central Australia suggests that fauna may be of a similar age. Broader faunal correlations have suggested Faunal Zone C correlates with the middle Miocene Bullock Creek Local Fauna, which contains the more derived W. vanderleueri. Based on stage-of-evolution arguments, W. oldfieldi should occur in older deposits than those yielding W. vanderleueri. The presence of both species of Wakaleo in Faunal Zone C assemblages at Riversleigh suggests that current presumptions about the contemporaneity of the many Faunal Zone C Sites should be examined more rigorously.

Anna K. Gillespie [a.gillespie@unsw.edu.au], Michael Archer [m.archer@unsw.edu.au], Suzanne J. Hand [s.hand@unsw.edu.au] and Karen H. Black [k.black@unsw.edu.au] School of Biological Earth and Environmental Science, UNSW 2052, Sydney, Australia. Received 3.1.2014, revised 21.2.2014, accepted 21.3.2014.     

How diverse were early animal communities? An example from Ediacara Conservation Park,Flinders Ranges,South Australia

Coutts, F.J., Gehling, J.G. & García-Bellido, D.C., August 2016. How diverse were early animal communities? An example from Ediacara Conservation Park, Flinders Ranges, South Australia. Alcheringa 40, xxx–xxx. ISSN 0311-5518

Fossils of the Ediacara biota record the earliest evidence of animal communities and, as such, provide an invaluable glimpse into the abiotic and biotic processes that helped shape the evolution of complex life on Earth. A diverse community of Ediacaran macro-organisms is preserved with high resolution in a fossil bed recently excavated from north Ediacara Conservation Park (NECP) in the Flinders Ranges, South Australia. Many of the commonly described Ediacaran taxa from the Flinders Ranges are represented on the bed surface and include: Parvancorina, Rugoconites, Spriggina, Dickinsonia, Tribrachidium, Kimberella, Charniodiscus and Yorgia, including two new taxa. Numerous additional fossil-bed fragments from the same locality were analysed that preserve a similar suite of taxa and shared sedimentology. On all surfaces, preserved microbial mat appeared complex, both in topography and in texture, and the unique combination of fine grainsize, high diversity and trace fossils provide insights into the palaeoecology of the ancient organisms that lived during the Ediacaran Period some 550 Ma. Several trace fossils are overlapped by body fossils, indicative of successive events, and complex organismal behaviour. The complexity of this fossil surface suggests that the primordial community was relatively mature and possibly at late-stage succession.

Felicity J Coutts [felicity.coutts@adelaide.edu.au], School of Biological Sciences, University of Adelaide, Adelaide 5000, South Australia, Australia. James G Gehling [jim.gehling@samuseum.sa.gov.au], South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia. Diego C. García-Bellido [diego.garcia-bellido@adelaide.edu.au], South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia.     

Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted,early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia

Chamberlain, P.M., Travouillon, K.J., Archer, M. & Hand, S.J., November 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa 40, XX–XX. ISSN 0311-5518.

A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.

Philippa M. Chamberlain [philippa.chamberlain@uq.net.au], School of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia; Kenny J. Travouillon [kenny.travouillon@museum.wa.gov.au; kennytravouillon@hotmail.com], Western Australian Museum, Locked Bag 49, Welshpool DC, WA, 6986, and School of Earth Sciences, University of Queensland, St Lucia, Queensland, 4072, Australia; Michael Archer [m.archer@unsw.edu.au] and Suzanne J. Hand [s.hand@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, New South Wales, 2052, Australia.     

First record of Homoctenus (Tentaculitoidea,Homoctenida) from the Late Devonian of northwest Peninsular Malaysia

Meor, H.A.H., Yong, A.M., Zakaria, M.Z.Z. & Ghani, A.A., 2.6.2015. First record of Homoctenus (Tentaculitoidea, Homoctenida) from the Late Devonian of northwest Peninsular Malaysia. Alcheringa 39, 550–558. ISSN 0311-5518.

The homoctenid tentaculitoid genus Homoctenus is reported for the first time from Peninsular Malaysia. The fossils derive from the Upper Devonian Sanai Limestone, exposed in the state of Perlis, northwest Peninsular Malaysia. The fossils are closely related to Homoctenus tenuicinctus tenuicinctus and are described as Homoctenus sp. cf. H. tenuicinctus. The homoctenids were recovered from an interval containing a rich conodont assemblage, indicating a Frasnian age (Palmatolepis linguiformis Zone).

Meor Hakif Amir Hassan [meorhakif@um.edu.my], Yong Adilah Mustafa [yongadilah@gmail.com], Mohamad Z.Z. Zakaria [zulhafiszariq@siswa.um.edu.my], Azman A. Ghani [azmangeo@um.edu.my], Geology Department, University of Malaya, 50603 Kuala Lumpur, Malaysia. Received 12.4.2015; revised 27.5.2015; accepted 2.6.2015.     

Reappraisal of Austrosaurus mckillopi Longman, 1933 from the Allaru Mudstone of Queensland,Australia’s first named Cretaceous sauropod dinosaur

Poropat, S.F., Nair, J.P., Syme, C.E., Mannion, P.D., Upchurch, P., Hocknull, S.A., Cook, A.G., Tischler, T.R. &; Holland, T. XX.XXXX. 2017. Reappraisal of Austrosaurus mckillopi Longman, 1933 Longman, H.A., 1933. A new dinosaur from the Queensland Cretaceous. Memoirs of the Queensland Museum 10, 131–144. [Google Scholar] from the Allaru Mudstone of Queensland, Australia’s first named Cretaceous sauropod dinosaur. Alcheringa 41, 543–580. ISSN 0311-5518

Austrosaurus mckillopi was the first Cretaceous sauropod reported from Australia, and the first Cretaceous dinosaur reported from Queensland (northeast Australia). This sauropod taxon was established on the basis of several fragmentary presacral vertebrae (QM F2316) derived from the uppermost Lower Cretaceous (upper Albian) Allaru Mudstone, at a locality situated 77 km west-northwest of Richmond, Queensland. Prior to its rediscovery in 2014, the type site was considered lost after failed attempts to relocate it in the 1970s. Excavations at the site in 2014 and 2015 led to the recovery of several partial dorsal ribs and fragments of presacral vertebrae, all of which clearly pertained to a single sauropod dinosaur. The discovery of new material of the type individual of Austrosaurus mckillopi, in tandem with a reassessment of the material collected in the 1930s, has facilitated the rearticulation of the specimen. The resultant vertebral series comprises six presacral vertebrae—the posteriormost cervical and five anteriormost dorsals—in association with five left dorsal ribs and one right one. The fragmentary nature of the type specimen has historically hindered assessments of the phylogenetic affinities of Austrosaurus, as has the fact that these evaluations were often based on a subset of the type material. The reappraisal of the type series of Austrosaurus presented herein, on the basis of both external morphology and internal morphology visualized through CT data, validates it as a diagnostic titanosauriform taxon, tentatively placed in Somphospondyli, and characterized by the possession of an accessory lateral pneumatic foramen on dorsal vertebra I (a feature that appears to be autapomorphic) and by the presence of a robust ventral mid-line ridge on the centra of dorsal vertebrae I and II. The interpretation of the anteriormost preserved vertebra in Austrosaurus as a posterior cervical has also prompted the re-evaluation of an isolated, partial, posterior cervical vertebra (QM F6142, the ‘Hughenden sauropod’) from the upper Albian Toolebuc Formation (which underlies the Allaru Mudstone). Although this vertebra preserves an apparent unique character of its own (a spinopostzygapophyseal lamina fossa), it is not able to be referred unequivocally to Austrosaurus and is retained as Titanosauriformes indet. Austrosaurus mckillopi is one of the oldest known sauropods from the Australian Cretaceous based on skeletal remains and potentially provides phylogenetic and/or palaeobiogeographic context for later taxa such as Wintonotitan wattsi, Diamantinasaurus matildae and Savannasaurus elliottorum.

Stephen F. Poropat* [sporopat@swin.edu.au; stephenfporopat@gmail.com] Department of Chemistry and Biotechnology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Jay P. Nair [j.nair@uq.edu.au; jayraptor@gmail.com] School of Biological Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Caitlin E. Syme [caitlin.syme@uqconnect.edu.au] School of Biological Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Philip D. Mannion [philipdmannion@gmail.com] Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, UK; Paul Upchurch [p.upchurch@ucl.ac.uk] Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK; Scott A. Hocknull [scott.hocknull@qm.qld.gov.au] Geosciences, Queensland Museum, 122 Gerler Rd, Hendra, Queensland 4011, Australia; Alex G. Cook [alex.cook@y7mail.com] School of Earth Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia; Travis R. Tischler [travisr.tischler@outlook.com] Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia; Timothy Holland [drtimothyholland@gmail.com] Kronosaurus Korner, 91 Goldring St, Richmond, Queensland 4822, Australia. *Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia.     

Oldest meiolaniid turtle remains from Australia: evidence from the Eocene Kerosene Creek Member of the Rundle Formation,Queensland

Poropat, S.F., Kool, L., Vickers-Rich, P. &; Rich, T.H., September 2016. Oldest meiolaniid turtle remains from Australia: evidence from the Eocene Kerosene Creek Member of the Rundle Formation, Queensland. Alcheringa 41, XX–XX. ISSN 0311-5518.

Fossil meiolaniid turtles are known only from South America and Australasia. The South American record is restricted to the Eocene, and comprises two genera: Niolamia and Gaffneylania. The Australasian meiolaniid record is more diverse, with three genera known (Ninjemys, Warkalania and Meiolania); however, the oldest known specimens from this continent are significantly younger than those from South America, deriving from upper Oligocene sediments in South Australia and Queensland. Herein, we describe the oldest meiolaniid remains found in Australasia to date. The specimens comprise a posterior peripheral, a caudal ring, and an osteoderm, all of which derive from the middle–upper Eocene Rundle Formation of The Narrows Graben, Gladstone, eastern Queensland. Despite their fragmentary nature, each of these specimens can be assigned to Meiolaniidae with a high level of confidence. This is particularly true of the partial caudal ring, which is strongly similar to those of Niolamia, Ninjemys and Meiolania. The extension of the Australasian meiolaniid record to the Eocene lends strong support to the hypothesis that these turtles arose before South America and Australia detached from Antarctica, and that they were consequently able to spread across all three continents.

Stephen F. Poropat*^? [stephenfporopat@gmail.com], Australian Age of Dinosaurs Natural History Museum, The Jump-Up, Winton, Queensland 4735, Australia; Lesley Kool*^? [koollesley@gmail.com] and Thomas H. Rich [trich@museum.vic.gov.au], Melbourne Museum, 11 Nicholson St, Carlton, Victoria 3053, Australia; Patricia Vickers-Rich [pat.rich@monash.edu], Monash University, Wellington Rd, Clayton, Victoria 3800, Australia. *These authors contributed equally to this work. ^?Also affiliated with Monash University, Wellington Rd, Clayton, Victoria 3800, Australia.     

A forewing of the Jurassic dragonfly Austroprotolindenia jurassica Beattie & Nel (Odonata: Anisoptera) from the Talbragar Fish Bed,New South Wales,Australia

Nel, A., Frese, M., McLean, G. & Beattie R., May 2017. A forewing of the Jurassic dragonfly Austroprotolindenia jurassica from the Talbragar Fish Bed, New South Wales, Australia. Alcheringa 41, 532–535. ISSN 0311-5518.

The discovery of a well-preserved dragonfly forewing in the Upper Jurassic Talbragar Fish Bed near Gulgong and attributed to Austroprotolindenia jurassica Beattie & Nel allows this taxon to be placed in Protolindeniidae. It extends the palaeogeographical distribution of this family, previously known only from the Jurassic of Europe, to Australia.

André Nel [anel@mnhn.fr], CNRS UMR 7205, CP 50, Entomologie, Muséum National d’Histoire Naturelle, 45 rue Buffon, F-75005, Paris, France; Michael Frese [michael.frese@canberra.edu.au], University of Canberra, Institute for Applied Ecology and Faculty of Education, Science, Technology and Mathematics, Bruce, ACT 2601, Australia; Graham McLean [mcleangg@bigpond.com], The Australian Museum, 1 William St., Sydney, NSW 2010, Australia; Robert Beattie [rgbeattie@bigpond.com], The Australian Museum, 1 William St., Sydney, NSW 2010, Australia.     

Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae,Caimaninae) in the Yecua Formation (late Miocene) of Bolivia

Tineo, D.E., Bona, P., Pérez, L.M., Vergani, G.D., González, G., Poiré, D.G., Gasparini, Z.N. & Legarreta, P., 1.10.2014. Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae, Caimaninae) in the Yecua Formation (late Miocene) of Bolivia. Alcheringa 39, xxx–xxx. ISSN 0311-5518

Outcrops of the Yecua Formation (late Miocene) are exposed for approximately 230 m along the La Angostura section of the Piraí River (50 km southwest of Santa Cruz de la Sierra). These reveal massive (argillic palaeosols) and laminated (quiet-water lacustrine and marsh settings) mudstones interbedded with thin sandstones containing microfossils, molluscs and vertebrate remains. Significantly, the succession hosts a giant crocodylian, Mourasuchus (Alligatoridae, Caimaninae), which is represented by both skull and postcranial fragments found in association with freshwater turtles and fishes. Mourasuchus was distributed widely from the middle Miocene of Colombia to upper Miocene of Venezuela, Brazil and Argentina, suggesting connections between major fluvial systems and an active mechanism for dispersal of South American freshwater vertebrates during the Miocene.

David Eric Tineo [tineo.d.e@gmail.com] and Daniel Gustavo Poiré [dgpoire@yahoo.com.ar], CONICET—Centro de Investigaciones Geológicas, Universidad Nacional de La Plata. Calle 1 (644), B1900FWA, La Plata, Argentina; Paula Bona [paulabona26@gmail.com] and Zulma Gasparini [zgaspari@fcnym.unlp.edu.ar], CONICET—División Paleontología Vertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Leandro Martín Pérez [pilosaperez@gmail.com] CONICET—División Paleozoología Invertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Gustavo Dardo Vergani [gvergani@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina; Gloria González Rigas [ggonzalez@pluspetrol.net]—Pluspetrol Bolivia Corporation SA, Av. Grigotá esq. Las Palmas, Santa Cruz de la Sierra, Bolivia; Pablo Legarreta [plegarreta@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina.     

Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan,southern Guangxi,South China

Yang, T.L., He, W.H., Zhang, K.X., Wu, S.B., Zhang, Y., Yue, M.L., Wu, H.T. & Xiao, Y.F., November 2015. Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan, southern Guangxi, South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

The Talung Formation (latest Permian) and basal part of Luolou Formation (earliest Triassic) of the Dongpan section have yielded 30 bivalve species in 17 genera. Eight genera incorporating 11 species are systematically described herein, including three new species: Nuculopsis guangxiensis, Parallelodon changhsingensis and Palaeolima fangi. Two assemblages are recognized, i.e., the Hunanopecten exilis–Euchondria fusuiensis assemblage from the Talung Formation and the Claraia dieneri–Claraia griesbachi assemblage from the Luolou Formation. The former is characterized by abundant Euchondria fusuiensis, an endemic species, associated with other common genera, such as Hunanopecten, which make it unique from coeval assemblages of South China. A palaeoecological analysis indicates that the Changhsingian bivalve assemblage at Dongpan is diverse and represented by various life habits characteristic of a complex ecosystem. This also suggests that redox conditions were oxic to suboxic in deep marine environments of the southernmost Yangtze Basin during the late Changhsingian, although several episodes of anoxic perturbations and declines in palaeoproductivity saw deterioratation of local habitats and altered the taxonomic composition or population size of the bivalve fauna.

Tinglu Yang [yang@geology.so], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Weihong He* [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Shunbao Wu [shbwu@cug.edu.cn], Yang Zhang [zhangy05@163.com], Mingliang Yue [812182779@qq.com], Huiting Wu [ht_wu415@163.com] and Yifan Xiao [shadowyi@sohu.com], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

Probable oribatid mite (Acari: Oribatida) tunnels and faecal pellets in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation,central-western Queensland,Australia

Fletcher, T.L. & Salisbury, S.W., XX.XX. 2014. Probable oribatid mite (Acari: Oribatida) tunnels and faecal pellets in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, central-western Queensland, Australia. Alcheringa 38, 541–545. ISSN 0311-5518.

Tunnels and faecal pellets likely made by oribatid mites have been found in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, central-western Queensland, Australia. Although this is the first identified and described record of oribatid mites in the Mesozoic of Australia, other published, but unassigned material may also be referable to Oribatida. Current understanding of the climatic significance of mite distribution is limited, but the presence of oribatids and absence of xylophagus insects in the upper portion of the Winton Formation are consistent with indications that the environment in which this unit was deposited was relatively warm and wet for its palaeolatitude. Such traces may provide useful and durable proxy evidence of palaeoclimate, but more detailed investigation of modern taxa and their relationship to climate is still needed.

Tamara L. Fletcher [t.fletcher1@uq.edu.au] and Steven. W. Salisbury, [s.salisbury@uq.edu.au] School of Biological Sciences, The University of Queensland, Australia, 4072. Received 28.1.2014; revised 1.4.2014; accepted 3.4.2014.     

Pliocene Mollusca (Bivalvia,Gastropoda) from the Sørsdal Formation,Marine Plain,Vestfold Hills,East Antarctica: taxonomy and implications for Antarctic Pliocene palaeoenvironments

Quilty, P.G., Darragh, T.A., Gallagher, S.J. & Harding, L.A. July 2016. Pliocene Mollusca (Bivalvia, Gastropoda) from the Sørsdal Formation, Marine Plain, Vestfold Hills, East Antarctica: taxonomy and implications for Antarctic Pliocene palaeoenvironments. Alcheringa 40, XXX–XXX. ISSN 0311-5518.

Pliocene shallow-water marine sediments at Marine Plain (centred on 68°37.7?S; 78°07.8?E) and covering approximately 10 km² in the Vestfold Hills, East Antarctica, have yielded six species of gastropods, and 11 species of bivalves from two beds within the Sørsdal Formation. Most of the material is close to in situ but some specimens have been disturbed from their life position; there is no evidence of significant transport. The gastropods include Nacella concinna (Strebel, 1908), Falsimargarita parvispira Quilty, Darragh, Gallagher & Harding sp. nov., indeterminate species of trochids and naticids, Chlanidota (Chlanidota) sp. cf. C. signeyana Powell, 1951, and two species of Trophon/Trophonella. Bivalves include Ennucula sp. aff. E. grayi (d’Orbigny, 1846), Aequiyoldia defossata Quilty, Darragh, Gallagher & Harding, sp. nov., ‘Pectunculina’ sp., Lissarca sp., Austrochlamys anderssoni (Hennig, 1911), Ruthipecten campestris Quilty, Darragh, Gallagher & Harding sp. nov., Adamussium necopinatum Quilty, Darragh, Gallagher & Harding sp. nov., Limatula (Antarctolima) sp. cf. L. hodgsoni (Smith, 1907), Cyclocardia magna Quilty, Darragh, Gallagher & Harding sp. nov., ?Hiatella sp. cf. H. arctica (Linnaeus, 1767) and Laternula elliptica (King, 1832). Preservation varies considerably owing to recrystallization, dissolution or distortion through compaction, so several species are left in open nomenclature. Oxygen isotope data indicate that water temperature was 4–7.5°C at the time of shell growth. Many species or species groups are now extinct or have migrated away from the Antarctic to the sub-Antarctic region. An Antarctic mollusc fauna has been characteristic of the region for much of the Cenozoic.

Patrick G. Quilty [p.quilty@utas.edu.au], Discipline of Earth Sciences, University of Tasmania, Private Bag 79, Hobart, Tasmania 7001, Australia; Thomas A. Darragh [tdarragh@museum.vic.gov.au], Museum Victoria, GPO Box 666 Melbourne, Victoria 3000, Australia; Stephen J. Gallagher [sjgall@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Lucy A. Harding [harding.lucy.a@edumail.vic.gov.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia.