A biometric technique for assessing prehistoric freshwater mussel population dynamics (family: Unionidae) in north Texas

Biometry is useful for a suite of analyses relevant to zooarchaeology, which include analysis of mortality profiles, taphonomy, paleoecology, among other purposes. Here, size-age prediction models are developed for 10 freshwater mussel species found in north Texas to develop a method for studying paleoenvironmental conditions. Shell length is often used to evaluate the structures of modern mussel populations, but its use on paleozoological specimens is not feasible because complete shells rarely preserve. Instead, pallial line-to-lateral teeth length (PLL) and pseudocardinal teeth-to-pallial line length (PSP) are evaluated as proxy measures for shell length. Linear regression models based on PLL and PSP using modern mussels demonstrate that they are accurate proxies of shell length for multiple species from a variety of habitats. In addition, ontogenetic-age structures for a modern sample and for two late Holocene assemblages from north Texas are developed using PLL and PSP. These techniques are useful tools for evaluating past ecological conditions of freshwater mussel populations when large samples are available for study, which expands analytical potential of zooarchaeological studies of prehistoric unionid remains.     

Harvesting the Seashores in the Late Mesolithic of Northwestern Europe: A View From Brittany

The shell middens of Brittany provide the last evidence of a Mesolithic way of life along the French Atlantic façade. This is partly a result of Holocene marine transgressions that prevent easy access to earlier coastal settlements. Nevertheless, the dependence on the sea seen in the Late Mesolithic seems to be a consequence of a long-established exploitation system. Stable carbon and nitrogen isotope signatures in human bone reflect a dominance of marine protein, while the zooarchaeological components of shell middens show a high species richness of exploited marine resources. Paleoenvironmental reconstruction suggests that more or less the whole range of resources exploited was accessible in the immediate vicinity of the sites. Seasonal aspects of the utilised and potentially available subsistence resources, along with stable isotope and lithic data, raise the possibility of restricted mobility for these populations, within relatively limited territories. The impression of extreme dependence of these coastal populations on the seashore might have been a key factor in their final disappearance, whether this is viewed as replacement or acculturation. Indeed, the Mesolithic communities of Brittany could have been caught between rising sea-levels and the arrival of Neolithic communities from the east and the south.     

Shell morphometry of seven limpet species from coastal shell middens in southern Africa

Measurements of shell parts and features (elements) of marine limpets can be used to derive morphometric equations for estimating total shell lengths. This is demonstrated for seven limpet species commonly found on the southern African coast. The equations can be used to reconstruct whole shell lengths for highly fragmented limpet samples in prehistoric shell middens. A linear regression model is based on measurements of all shell elements, resulting in high coefficients of determination with excellent predictive power in most cases. These morphometric equations would enable archaeologists to derive more metrical information from fragmentary archaeological material than was previously the case. We also present a case study where morphometric equations of two limpet species are applied to an archaeological sample from the South African west coast for the purpose of investigating possible biases in limpet shell preservation. We conclude that small whole limpet shells survive longer than the bigger ones in this particular case, but that many more such case studies need to be conducted in order to fully understand differential preservation of southern African limpet shells in archaeological sites.     

The exploitation of marine resources at Saruq al‐Hadid: Insights into the movement of people and resources in Bronze and Iron Age south‐eastern Arabia

Marine resources were an integral and consistent component of subsistence strategies employed in south‐eastern Arabia throughout late prehistory. Of particular interest is the movement of these resources from the coast to interior sites and the implications of this movement for transhumance and trade in the region during this period. Marine species were frequently identified in the faunal assemblage from the inland site of Saruq al‐Hadid, dating from the Bronze Age to Early Iron Age (c.2000–c.800 BCE). This included marine fish species, along with two cormorant species (Phalacrocorax sp.) and several fragments of dugong (Dugong dugon). Twenty‐seven families of marine shell were also identified in the remains recovered from the site. The presence of these remains at this inland site demonstrates that resources were frequently moved from the coast to the interior throughout Saruq al‐Hadid’s occupation, indicative of their enduring significance in subsistence strategies employed at the site. This paper presents the results of zooarchaeological analysis of these remains and discusses the significance of their presence at Saruq al‐Hadid, with reference to subsistence, craft production and intra‐regional exchange during the Bronze and Iron Ages.     

The Shell-Bearing Archaic in the Middle Cumberland River Valley

The Middle Cumberland River Valley (MCRV) of Tennessee comprises a unique regional environment that has continually supported human occupation along the natural river levees and adjacent terrace landforms since the Late Pleistocene. Over thousands of years Archaic period inhabitants of the MCRV harvested the invertebrate species that populated the streams and waterways of the region, using them for subsistence and raw materials and taking an active role in managing the riverine resources. The cumulative result of this process appears in the archaeological record as abundant and often-dense deposits of invertebrate zooarchaeological remains. However, few formal archaeological investigations have been conducted on Archaic shell-bearing sites in the region. In this field report we present initial results of site file analysis, radiocarbon dating, and species composition research in order to introduce the MCRV manifestation of the cultural phase traditionally known as the Shell Mound Archaic.     

Species specific marine radiocarbon reservoir effect: a comparison of ΔR values between Patella vulgata (limpet) shell carbonate and Gadus morhua (Atlantic cod) bone collagen

A number of recent studies of the UK coastal environment have assessed the ¹⁴C marine radiocarbon reservoir effect (MRE) via quantification of ΔR values for several periods throughout the Holocene using marine mollusc shells. However, none have employed fish bone as the marine sample, and the importance of being able to use this material as a reliable dating tool is evident when considering the boom in the British fish trade during the first millennium AD, the so-called ‘fish event horizon’, and the corresponding volume of fish remains that appear in the archaeological record from this time. This study compares ΔR values derived using the multiple paired sample approach employing burnt cereal grain (Hordeum sp.) as the terrestrial sample and either fish bone [North Sea cod (Gadus morhua)] or marine shell [limpet (Patella vulgata)] as the marine sample. The results show a general trend of increasing ΔR for the fish bone compared to shell, however, the differences are not statistically significant when the standard error for predicted values is used as the measure of variability in the ΔR values.     

MARINE RESOURCES IN THE MESOLITHIC AND NEOLITHIC AT THE GROTTA DELL’UZZO (SICILY): EVIDENCE FROM ISOTOPE ANALYSES OF MARINE SHELLS*

Oxygen isotopes in shell carbonate samples from the marine rocky‐shore intertidal gastropod Monodonta turbinata (Born) are investigated in both modern analogue specimens and in archaeological specimens from the Grotta dell’Uzzo (Sicily). Variations in shell edge values of δ¹⁸O in living specimens collected monthly over two years are closely correlated with monthly seawater temperatures measured at the time of collection, showing that the species can be used for palaeoseasonality studies. Analyses of shell edge δ¹⁸O values in archaeological specimens, from Mesolithic through to early Neolithic phases at the Grotta dell’Uzzo, enabled the inference of various seasons of collection of shellfish and how such seasonality varies between the different phases of occupation. Interesting similarities and differences exist between the seasons of marine shellfish exploitation and the seasons inferred from the vertebrate zooarchaeological assemblages. A major inference drawn from the analyses and discussion is that the exploitation of all marine resources (fish and shellfish) increased in the later Mesolithic and early Neolithic periods.     

The Distribution and Research Value of Archaeological Mussel Shell: An Overview from Mississippi

Abstract

Archaeological mussel shell rarely has been employed beyond paleodietary analysis in eastern North America, and most published work is site-specific in nature. Broad-scale syntheses of data on freshwater mussel shells are needed to realize the research potential of this common archaeological material. Such syntheses should also reveal gaps in our knowledge and perhaps illuminate reasons for why such gaps exist. In this paper, data on freshwater mussel shells from sites in Mississippi are gathered, and broad patterns in temporal and spatial distributions as well as site structure are discussed. The spatial data clearly suffer from survey, recovery, analytical and reporting biases, while data on temporal trends and site structure are lacking or are available at only gross levels. Given that shell data can be used in a number of research areas (e.g., biogeography, paleoenvironment, chemical sourcing, measurement of environmental disturbance), such unsystematic treatment of shell remains is unwarranted and should be corrected.     

Large shell middens in Lamberts Bay,South Africa: a case of hunter–gatherer resource intensification

Enormous shell middens (“megamiddens”) dating to c. 3000–2000 BP along the West Coast of South Africa were explained initially within a framework based on the Kalahari San ethnography. The overwhelming dominance of marine shell and low densities of artefacts and vertebrate remains were seen as merely the reflection of processing localities. A far more complex picture has emerged recently, with a trend of raising population densities and longer residential permanence between 3500 and 2500 BP. Successive reformulations of the foraging ecology of both marine and terrestrial prey are apparent, with isotopic values from human skeletons and quantified dietary remains showing increased marine food consumption during the megamidden period when compared to other stages. Shellfish collection shifted from a mix of limpets, whelks and black mussels before 3000 BP to the intensified collection of the later from about 2600 BP, with local impact on marine fauna becoming evident at this time. Coastal groups became also increasingly less reliant on large mobile game and more so on small territorial bovids and tortoises as from 3500 BP, with strongest emphasis on this foraging behaviour after 2700 BP. Read against palaeoenvironmental data, this reconstruction is consistent with hunter–gatherer resource intensification models world-wide.     

ZOOMORPHIC EFFIGY PENDANTS: AN EXAMINATION OF STYLE,MEDIUM, AND DISTRIBUTION IN THE CADDO AREA

Abstract

Caddo zoomorphic effigy pendants were crafted from a wide variety of raw material, such as freshwater mussel shell, Busycon perversum (whelk) marine shell, animal bone, and various types of soft stone. They have been documented at contemporaneous Late Caddo (A.D. 1400–1700) sites along the Red River in northeast Texas, southwest Arkansas, and northwest Louisiana; the Black Bayou, Big Cypress Bayou, and upper Sabine River basins in northeast Texas; the Ouachita River in south-central Arkansas; and the Arkansas River in central Arkansas and eastern Oklahoma. This paper describes the development of a comprehensive corpus that investigates stylistic groups, differential raw material or medium utilized, and the spatial distribution of zoomorphic effigy pendants throughout the Caddo area. Results reveal north-south heterogeneity, suggesting the presence of broader traditional cultural narratives associated with a complex idea that is manifest regionally in distinct stylistic forms and linked to Caddo beliefs concerning Beneath World themes. Additionally, this stylistic and spatial analysis offers an additional dataset to further explore social, political, and economic linkages among and between contemporary Caddo groups around A.D. 1500.     

The role of shellfish in hunter–gatherer societies during the Early Upper Palaeolithic: A view from El Cuco rockshelter,northern Spain

The role of coastal resources in the subsistence strategies of Palaeolithic human populations has only recently become an important topic in Old World archaeology. Information on the exploitation of these resources, both as foodstuffs and symbolic elements, can be used to infer the emergence of modern human behaviour as well as to track the diversification and intensification of human diet over time. The excavations carried out at El Cuco rockshelter, located in northern Spain have provided evidence for the exploitation of marine resources during the Early Upper Palaeolithic. The accumulation of Patella shells at El Cuco provides the largest accumulation and the first clear evidence of collection and consumption of molluscs during the Aurignacian on the Atlantic Façade of Europe. A deposit of ornamental shells appeared in a very homogeneous context dated to the Gravettian, suggesting that the shells belonged to the same item. The analysis of this evidence has allowed us to conclude that marine resources were systematically used at least from the beginning of the Upper Palaeolithic in the region. However, a comparison with the Mesolithic shows that intensive shellfish gathering did not occur until the end of the Upper Palaeolithic. Regarding the ornaments, it is interpreted that the identified shell beads were used as social or personal status markers.     

Oxygen isotopic composition of limpet shells from the Beagle Channel: implications for seasonal studies in shell middens of Tierra del Fuego

Shell middens distributed along the coastal areas of the Beagle Channel (Tierra del Fuego) testify the primary role of marine resources in the successful social dynamic of hunter–gatherer–fisher societies (HGF) in extreme high latitude environmental conditions. Intertidal and subtidal limpet species are frequently identified taxa of these archaeological deposits. We investigated whether shell δ¹⁸O of the limpet species Nacella deaurata is a valuable record of local sea surface temperature (SST) and ambient water δ¹⁸O (δ¹⁸Ow) fluctuations, and in turn can be used to estimate the season of its exploitation by Holocene HGF. Sequential shell δ¹⁸O of modern species track seasonal changes of SST and δ¹⁸Ow. The comparison of shell δ¹⁸O profiles of N. deaurata and Nacella magellanica (a coexisting species) suggests the occurrence of distinct growth rate and physiological adaptations between species. Shell δ¹⁸O of zooarchaeological specimens of N. deaurata from Lanashuaia II suggests that animals were exploited predominantly in winter and that experienced similar environmental conditions of present day at ∼1320 years BP. Results also indicate that N. deaurata is a valuable candidate for studying past seasonal changes in SST and δ¹⁸Ow in this southernmost part of South America.     

Applied Zooarchaeology of Freshwater Mussel (Bivalvia: Unionidae) Shell from Golson (22HU508), A Deasonville-Period Site on the Yazoo River,Mississippi

Excavations at Golson (22HU508), an archaeological site on the Yazoo River in western Mississippi, USA, produced an assemblage of over 4000 freshwater mussel shells, 77 percent of which were identifiable to species. Although limited spatial sampling of the deposits limits what can be said about past mussel community characteristics, twelve new river records for mussel species are derived from the assemblage. This typical result emphasises the value of archaeological shell assemblages for conservation biology and provides a rationale for why sites containing shell may be considered significant based upon their potential contributions to environmental history.     

Middle Holocene fishing strategies in East Africa: zooarchaeological analysis of Pundo,a Kansyore shell midden in northern Nyanza (Kenya)

A zooarchaeological and taphonomic analysis is presented for the faunal remains from the Pundo shell midden in northern Nyanza province (Kenya), which contains Kansyore ceramics and dates to ca. 8000–7600 cal. BP. The faunal assemblage is overwhelmingly dominated by fish and molluscs, of which only the fish data are presented in detail. Taxonomic identifications suggest a diachronic change in relative species abundance, with cichlids becoming increasingly dominant, eventually comprising nearly 80% of the fish assemblage. The Pundo faunal data are compared with those from a number of other Kansyore sites. Pundo confirms that early Kansyore sites are purely forager sites, with domestic caprines only appearing at late Kansyore sites (post‐dating ca. 4400 cal. BP). Comparing ethological data for fish taxa found at Pundo and other lakeshore shell middens with those for fish taxa found at larger riverside sites, a seasonal round is suggested, in which short‐term fishing camps such as Pundo may have been occupied in the dry season, while riverside sites were probably occupied during the rainy season. Copyright © 2008 John Wiley & Sons, Ltd.     

Coastal Feasts: A Pleistocene Antiquity for Resource Abundance in the Maritime Deserts of North West Australia?

ABSTRACT

Located on the edge of Australia's North West continental shelf, Barrow Island is uniquely located to address a number of research questions, such as the antiquity and changing nature of Indigenous occupation, including shifting uses of regional economic resources in response to post-glacial sea-level rise. These questions are addressed from a range of archaeological, zooarchaeological, and geoarchaeological disciplines. Although only preliminary, results to date indicate the presence of marine resources dating to before sea-level stabilization (～ 7.5 ka) that contain both dietary and utilitarian species, including high-ranked species such as sea turtle. The marine assemblages reflect a variety of habitats and substrates with a 17,000-year record for the presence of a former tidal marsh or estuary. We also note recently obtained ¹⁴C and OSL dates that extend the dietary marine faunas and initial occupation to well before 41 ka. This demonstrates that consumption of coastal resources began prior to the Holocene, when we begin to observe more widespread evidence of marine resource exploitation in the broader Canarvon Bioregion of northwest Australia. This evidence supports arguments for further research to directly test both the productivity of, and human reliance on, marine habitats from initial occupation.     

Early Hominin Paleoecology

Abstract

This paper reports on the zooarchaeological and archaeobotanical remains from the initial season of excavations at the Norse period site at Undir Junkarinsfløtti in the Faroe islands. These remains represent the first zooarchaeological analysis undertaken for the Faroes and only the third archaeobotanical assemblage published from the islands. The excavated deposits are described and the key findings from the palaeoenvironmental remains highlighted within the context of the wider North Atlantic environmental archaeology of the Norse period.     

Starch analysis reveals prehistoric plant translocations and shell tool use,Marquesas Islands,Polynesia

Starch analysis is proving particularly useful in tropical regions like the central Pacific where crop inventories are often dominated by starchy fruits and tubers and recovery of macrobotanical remains is rare. Analysis of 23 shell tools from the Marquesas Islands provides direct evidence for translocation of five traditional crop plants. Four taxa derive from the western Pacific, including Artocarpus altilis (breadfruit), Piper methysticum (kava), Colocasia esculenta (taro), and one or more species of Dioscorea (yam). The fifth taxon, the South American Ipomoea batatas (sweet potato), dates from the 14th century AD onward and constitutes the earliest record of this cultigen in the archipelago. The occurrence of sweet potato starch suggests that this crop plant may have been more important than usually is assumed, while the limited recovery of breadfruit starch, the main Marquesan food plant at western contact, requires further investigation. The starch residues also inform on tool use, demonstrating that shell tools ethnographically associated with specific crops (e.g., “breadfruit peelers”) had more generalised functions. This study is a further demonstration of the potential of starch analysis to provide important information on the history of crop introductions, on-site activities, and artefact use.     

Indirect Evidence of Falconry in Medieval Poland as Inferred from Published Zooarchaeological Studies

Although written sources indicate that falconry or hawking was very popular already in early medieval Poland, there is no hard zooarchaeological evidence that would support it. So far, only two studies based on animal remains suggested hawking in medieval Poland. The purpose of this study was to examine all published zooarchaeological reports from all archaeological sites in Poland and check for possible clues of hawking. Altogether, 281 remains of 12 species of diurnal birds of prey were recorded from 38 sites (49 time‐site units) in Poland beginning from the Middle Ages onward. The most frequently found was the goshawk Accipiter gentilis followed by the white‐tailed eagle Haliaeetus albicilla and the sparrowhawk Accipiter nisus. Clues of possible hawking include high status of sites, occurrence of species used in falconry outside their natural ranges, preponderance of females, the presence of sub‐adult specimens and others. It is suggested that surface damage observed on prey bones done by the hawks' beaks and claws may be an extra evidence of hawking. The present study strongly supports the notion that hawking was a very popular activity in medieval Poland. In the future, medieval bone assemblages should always be checked for clues of hawking. The present study may also be used as a source of raw data for other analyses because it includes detailed information on birds of prey from all zooarchaeological reports published so far in Poland. Copyright © 2015 John Wiley & Sons, Ltd.     

Taphonomic analysis of remains of mammals eaten by pumas (Puma concolor Carnivora, Felidae) in central Argentina

Prey remains recovered from puma (Puma concolor) scat were analyzed in order to identify taphonomic features produced on them by this predator. Mammal remains are abundant in puma scat, but recognition of bone remains decreases as prey size increases. Modifications produced by digestion are strong. However, micromammal bones show a gradation in preservation that includes well preserved specimens. Data presented here and their interpretation may possibly be extrapolated to zooarchaeological or paleontological assemblage.     

Human impacts on ancient shellfish: a 10,000 year record from San Miguel Island,California

We use measurements of more than 11,000 marine shells from 41 archaeological components to construct a 10,000 year record of human impacts on ancient mussel and abalone stocks on San Miguel Island, California. General reductions in the mean size of mussel and abalone shells gathered through time are attributed to growing human population and predation pressure. Based on comparison with historically documented changes in shellfish communities caused by the local extinction of sea otters in the 19th century, changes in mean shell size and the abundance of other shellfish species may have been facilitated by Native American predation on sea otters as early as 7500 years ago. Despite having measurable impacts on local ecosystems, Native Americans on San Miguel harvested huge quantities of shellfish throughout the Holocene. Such long-term harvests appear to have been sustained by an early emphasis on fishing at lower trophic levels, by periodically shifting village locations, and by intensifying the use of finfish and sea mammals through time. This pattern of “fishing up the food web” contrasts with many modern fisheries, suggesting that the study of ancient fisheries can help us better manage our own endangered coastal ecosystems.