The extinct large cranes of the North-West palaearctic

Bones of a large crane, comparable in size with the Sarus Crane, Grus antigone, have been found in the Late Pleistocene of Britain and France, neolithic of Germany, and Bronze and Iron Age sites in Britain. They occur together with bones of the Common Crane, G. grus. On zoogeographical grounds these are unlikely to be referable to G. antigone, and the name G. primigenia Milne-Edwards 1896 is available for the species.G. melitensis, a crane of similar large size from the Eemian interglacial of Malta differs in having a narrower tendinal bridge on the distal tibiotarsus, and a small slender coracoid that suggests that the powers of flight may have been reduced. On present evidence it has no link with the previous species other than that of size.     

Population relationships of later Pleistocene hominids: A multivariate study of available crania

The Neanderthal populations of the Upper Pleistocene have been regarded by a number of anthropologists as direct ancestors to modern man. Results of multivariate analyses conducted in this study suggest that early Neanderthal crania were morphologically more like modern Homo sapiens than were the later Neanderthals. The implications of these results are discussed in the light of archaeological evidence and comparative studies of Pleistocene crania from outside Europe.     

Getting “Out of Africa”: Sea Crossings,Land Crossings and Culture in the Hominin Migrations

Palaeoanthropologists and archaeologists have advanced a wide range of explanatory narratives for the various movements of Homo erectus/Homo ergaster, and the first modern Homo sapiens, “Out of Africa”—or even back again. The application of Occam's razor—a parsimonious approach to causes—gives a more cautious approach. There is nothing in the available evidence that would require the ability for a human water crossing from Africa before the later Pleistocene, whether across the Strait of Gibraltar, the Sicilian Channel or the southern Red Sea (Bab el-Mandab). A parsimonious narrative is consistent with movements across the Sinai peninsula. The continuous arid zone from northern Africa to western Asia allowed both occupation and transit during wet phases of the Pleistocene; there is no requirement for a “sponge” model of absorption followed by expulsion of human groups. The Nile Valley as a possible transit route from East Africa has a geological chronology that could fit well much current evidence for the timing of human migration. The limited spatial and temporal opportunities for movements “Out of Africa,” or back again, also puts particular difficulties in the way of the gene flow required for the multiregional hypothesis of the development of modern Homo sapiens.     

Pleistocene frogs from the Darling Downs,southeastern Queensland,and their palaeoenvironmental significance

Systematic collecting from fluvial late Pleistocene deposits from the Darling Downs, southeast Queensland, Australia, has led to the recovery of the first fossil frogs from the region, ail from the Myobatrachidae, a family of ground dwelling and burrowing frogs. The most common species recovered, Limnodynastes tasmaniensis, is extant on the Darling Downs. The fossil taxa include species whose extant populations inhabit arid zones(Limnodynastes sp. cf. L. spenceri), montane forests (Kyarranus spp.), and open woodlands (Neobatrachus sudelli), and indicate the existence of a mosaic of habitats during the Pleistocene. The absence of the Hylidae (tree frogs), a family common throughout the Darling Downs today, may be explained by a taphonomic bias that favours non-arboreal forms. Alternatively, hylids may have been rare or absent on the Darling Downs during the Pleistocene.     

First report of a South American short-faced bears' den (Arctotherium angustidens): palaeobiological and palaeoecological implications

Here we report the first example of associated short-faced bear fossils from South America. The specimens represent three individuals referable to the Ensenadan (early to middle Pleistocene) species Arctotherium angustidens (Ursidae, Tremarctinae), the giant South American short-faced bear. Although the fossil record of short-faced bears in South America is very rich, they have not previously been recorded in association. These three individuals were found in a cave during quarry exploitation. We suggest that these bears represent the first record of a family group and open the discussion about cave utilization and hibernation or torpor by South American short-faced bears.     

New Human Remains of Archaic Homo sapiens and Lower Palaeolithic Industries from Visogliano (Duino Aurisina, Trieste, Italy)

New human remains have been found during recent excavations conducted in the lower section of Visogliano dolina (Trieste, Italy). They are represented by teeth and a fragment of particularly robust mandible that for their morphological and morphometrical characters can be generically attributed to archaic Homo sapiens. The lithic industry is formed by flakes, carinated points, scrapers, choppers and chopping tools and is characteristic of the Italian Pebble Culture. Sediments, pollen and faunal remains indicate that this part of the deposit was formed during interglacial climatic conditions in the early, but not initial phase of Middle Pleistocene (isotopic stadium 13 or 11).     

Isotopic analysis of the ecology of herbivores and carnivores from the Middle Pleistocene deposits of the Sierra De Atapuerca,northern Spain

Carbon and oxygen isotope values reveal resource partitioning among the large mammal fauna from three contemporaneous Middle Pleistocene hominid-bearing localities within the Sierra de Atapuerca (northern Spain). Carbon isotope values sampled from the tooth enamel of fauna present during Atapuerca Faunal Unit 6 show that a C₃-dominated ecosystem surrounded the area where fossils were preserved during this time. For the herbivores, Fallow deer isotope values are significantly different from Red deer and horses and show that this species did not forage in open environments at this locality. Red deer and horses show similar feeding strategies with less negative carbon values implying use of more open environments for these taxa. For the carnivores, carbon isotope values for Ursus deningeri are significantly different from either lions (Panthera leo) or foxes (Vulpes vulpes) and support the contention that this species is herbivorous. Special metabolic mechanisms involved in hibernation in U. deningeri might also have influenced its isotope values. The carbon isotope values of remaining carnivores were similar and suggest that each was typically a generalist carnivore, eating a wide variety of prey items. While the isotopic results generally correspond to ecology indicated by previous techniques, this study shows that isotope analyses can provide further insights that alternate techniques do not provide. Isotope analyses can help elucidate the ancient ecology of taxa present in the Sierra de Atapuerca during the Middle Pleistocene allowing for an accurate portrayal of the setting in which humans lived.     

Lagostomus maximus (Desmarest) (Rodentia,Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay

Ubilla, M. & Rinderknecht, A., April 2016. Lagostomus maximus (Desmarest) (Rodentia, Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The extant plains vizcacha, Lagostomus maximus, is described from the Late Pleistocene (Dolores Formation) of Uruguay based on an almost complete articulated skeleton. It is compared with the nominally extinct Pleistocene species of the genus. An AMS ¹⁴C taxon-age is determined for L. maximus at 11 879 ± 95 years BP (cal. BP 13 898–13 941). Lagostomus maximus is absent from modern mammal communities in Uruguay, and no Holocene evidence is available. Because L. maximus exhibits remarkable sexual and ontogenetic dimorphism, we examined a range of juvenile, sub-adult and adult male and female specimens. It is not possible to differentiate the articulated Pleistocene fossil from sub-adult specimens of L. maximus based on this sample. Moreover, the skull characteristics, including a broad vertical ramus of the zygomatic arch, semi-circular temporal crest and short robust sagittal crest, most closely resemble extant female individuals. Molar size is non-predictive for ontogenetic stage or body mass because it reaches stability during early adulthood. Other fragmentary skull remains are herein assigned to L. sp. cf. L. maximus. Lagostomus cavifrons from the Pleistocene of Argentina is considered synonymous with L. maximus. Its character states concur with the range of intraspecific variability and it has been established on a young adult or adult male individual. Likewise, the other Argentinean Pleistocene species based on incomplete mandibles, L. angustidens, L. striatus, L. heterogenidens, L. egenus, L. minimus and L. debilis, concur with ontogenetic morphs and are here assigned to Lagostomus sp. The ecological preferences of extant L. maximus infer open arid or semi-arid landscapes for the latest Pleistocene of southern Uruguay. This hypothesis is reinforced by the coeval presence of Microcavia, Galea and small camelids in the same strata. The Last Glacial Maximum likely promoted this environmental setting. Latest Pleistocene or early Holocene climatic change might have facilitated local extinctions and/or range shifts among this mammal fauna.

Martín Ubilla [martinubilla@gmail.com], Facultad de Ciencias, Universidad de la Republica, Iguá 4225, 11400, Montevideo, Uruguay; Andrés Rinderknecht [apaleorinder@yahoo.com], Museo Nacional de Historia Natural, Paleontología, 11000, Montevideo, Uruguay.     

Presence of the African Saber-toothed FelidMegantereon whitei(Broom, 1937) (Mammalia, Carnivora, Machairodontinae) in Apollonia-1 (Mygdonia Basin, Macedonia, Greece)

We have made a multivariate morphometric study of the fossils ofMegantereonfrom the Apollonia-1 site at the Mygdonia Basin (Macedonia, Greece), using discriminant functions. Results obtained indicate that these specimens belong to the African speciesMegantereon whiteiBroom, which is also found in the Lower Pleistocene of Dmanisi (East Georgia, Caucasus) and Orce (Southern Spain). Morphofunctional studies ofM. whiteisuggest that this saber-toothed felid was an ambush predator with great killing capability in relation to its food requirements, thus leaving enough meat to be scavenged by both hyaenas and hominids.M. whiteimay have played an important ecological role making the first dispersal ofHomoto Eurasia the Lower Pleistocene possible.     

Shellfishing and the Colonization of Sahul: A Multivariate Model Evaluating the Dynamic Effects of Prey Utility,Transport Considerations and Life-History on Foraging Patterns and Midden Composition

ABSTRACT

Archaeological evidence of shellfish exploitation along the coast of Sahul (Pleistocene Australia-New Guinea) points to an apparent paradox. While the continental record as a whole suggests that human populations were very low from initial colonization through early Holocene, coastal and peri-coastal sites dating to that time are dominated by small, low-ranked, littoral taxa to the near-complete exclusion of large, higher ranked, sub-littoral species, precisely the opposite of theory-based expectations, if human populations and predation rates were indeed as low as other data suggest. We present a model of shellfish exploitation combining information on species utility, transport considerations, and prey life-history that might account for this apparent mismatch, and then assess it with ethnographic and archaeological data. Findings suggest either that high-ranked taxa were uncommon along the Pleistocene coastlines of Sahul, or that abundant and commonly taken high-ranked prey are under-represented in middens relative to their role in human diets largely as a function of human processing and transport practices. If the latter reading is correct, archaeological evidence of early shellfishing may be mainly the product of subsistence activities by children and their mothers.     

MOLLUSC SHELL SIZES IN ARCHAEOLOGICAL CONTEXTS IN NORTHERN SPAIN (13 200 TO 2600 CAL BC): NEW DATA FROM LA GARMA A AND LOS GITANOS (CANTABRIA)

Shellfish metrical data are a source of information about the exploitation of marine resources in the past. In this study, we propose a methodological approach based on the size structures of different rocky intertidal gastropod species. Three limpet species (Patella vulgata, Patella intermedia and Patella ulyssiponensis) and the toothed topshell Osilinus lineatus are studied from two sites in Cantabrian Spain: La Garma A and Los Gitanos caves over a period of 10 000 years, covering the Pleistocene–Holocene transition. Data are also supplied about a further sea snail species, the periwinkle Littorina littorea (Upper Magdalenian). A reduction in size can be seen, between the upper Magdalenian and the late Neolithic, in the case of the first four species. The explanation for this decline is probably related to the climate change that occurred in the transition between the Late Pleistocene and the Holocene, but it is possible that human impact might also have influenced shell sizes in the Mesolithic and Neolithic.     

Relative and absolute dating of the human skull and skeleton from Galley Hill,Kent

Over a period of some 100 years it has gradually been accepted that the human skeletal remains found in the late nineteenth century in ancient gravels at Galley Hill, Kent are not those of early man, but are of much more recent date. This conclusion was finally verified by means of relative (N, F) and absolute (¹⁴C) dating. Here we present the results of further analysis of the remains from Galley Hill for their uranium content by a method having general application to the relative dating of fossil human and other mammalian bone.     

On the occurrence of the scimitar-toothed cat, Homotherium latidens (Carnivora; Felidae), at Kents Cavern,England

Teeth of Homotherium latidens recovered from late Pleistocene sediments, Kents Cavern, England have long been the source of controversy. H. latidens is conspicuously absent from other late Pleistocene cave deposits in Britain, and is widely thought to have been extirpated from the region during the isotope stage 10 glacial period. Here we present high spatial resolution analyses of fluorine and uranium uptake profiles in teeth of three species from the same cave. The H. latidens tooth is clearly distinguished from the unambiguously provenanced Late Pleistocene hyaena and Middle Pleistocene cave bear teeth. These results are consistent with the theory that the H. latidens teeth originated at an exogenous location, were probably transported to Kents Cavern as Palaeolithic trade goods, and were buried in Kents Cavern in Palaeolithic times.     

The Enisei River of Central Siberia in the Late Pleistocene

Recent Paleolithic work along the middle Enisei River of central Siberia has revealed a long history of occupation that almost certainly begins in the Middle Pleistocene. Although the evidence for the Lower Paleolithic is somewhat tentative, there is good reason to believe that hunter-gatherers had periodically occupied the middle Enisei before the last interglacial. The steppe environment of the region during the Upper Pleistocene was relatively bountiful; more than 200 Upper Paleolithic sites, both before and after the Last Glacial Maximum, have been located. The region appears to have been abandoned during the Last Glacial Maximum. Most of Soviet and Russian archaeological work has been guided by a cultural–historical orientation, but recently there has been increased interest in developing adaptationist and ecological research strategies. The middle Enisei and the wider central Siberian region are key to understanding early adaptations to the north and the dimensions of Paleolithic population movements.     

Akhalkalaki: The Taphonomy of An Early Pleistocene Locality in the Republic of Georgia

As part of our investigations into the potential of the Republic of Georgia for providing information on early hominin occupation of Eurasia, we report here on Akhalkalaki, a large late Early Pleistocene locality located along the lower slopes of a Miocene andesitic cone. Originally excavated in the 1950s as a palaeontological site, it was re-opened in the 1990s and stone tools were found associated with the fauna, suggesting that it is also an archaeological occurrence. Excavations in the 1950s and 1990s uncovered thousands of bones of an early Galerian fauna, including the remains of new species of Hippopotamus,Equus , and Canis (Vekua, 1962, 1987) and dominated by the remains of Equus süssenbornensis. We present the stratigraphy of the site, which together with faunal correlations and reversed paleomagnetics indicates an age most likely in the late Matuyama Chron, probably between 980,000 and 780,000 years ago. Taphonomic analysis suggests that the fauna was deposited and buried over a short time period, and was heavily modified by carnivores, but we cannot demonstrate involvement by hominins. Based on evidence of abundant krotovina (animal burrows filled with sediment) and the lack of definitive evidence for hominin modification to the bones, the stone tools at the site may have been mixed in with the older fauna. The taphonomic characteristics of the Akhalkalaki bone assemblage are not readily explained with reference to assemblage formation processes developed with actualistic studies that have been mostly conducted in Africa, including carnivore dens, predator arenas, human hunting and scavenging, mass deaths, or attritional bone deposition. Because of extreme anthropogenic modification of the present environments, the temperate setting, and the presence of mainly extinct taxa, local models based on actualistic studies cannot approximate the mammalian ecology reflected in the Akhalkalaki bone assemblage. A few comparisons are made with preliminary taphonomic observations from Dmanisi, an Early Pleistocene Homo ergaster site not far away.     

The oldest handaxes in Europe: fact or artefact?

Hominin presence is well documented in a number of Early Pleistocene and early Middle Pleistocene European localities. However, the evidence currently available indicates that Acheulean handaxes spread in the fluvial basins of Western Europe during MIS 11, ～400 kyr ago, associated with Homo heidelbergensis, although a number of early Middle Pleistocene Acheulean assemblages have been dated from MIS 16 onwards. For this reason, the magnetostratigraphic dating in Southeast Spain of two archaeological localities, the open-air site of Solana del Zamborino (SZ) and the rock-shelter site of Cueva Negra del Estrecho del Quípar (EQ), that put back the appearance of handaxes to the Early-Middle Pleistocene limit (Scott and Gibert, 2009) is of particular interest, as the new ages suggest that H. heidelbergensis was a contemporary of H. antecessor that had the ability to produce Levallois debitage and to control fire during the Early-Middle Pleistocene transition. However, we have detected a number of errors in the interpretation of the archaeological assemblage from the first site as well as striking discrepancies with the original faunal lists published for both localities, with several large mammal species that are omitted or arbitrarily changed to make the assemblages consistent with the new ages deduced from magnetostratigraphy. For this reason, we suggest that: (1) the finding of reverse polarity in the sediments sampled for paleomagnetism in SZ may simply record one of the polarity reversals that took place during the Brunhes Chron, although the use by Scott and Gibert (2009) of a composite stratigraphic column precludes correlating these levels with a specific reversal; and (2) the fauna and tools of EQ correspond to the late Middle Pleistocene sedimentary infillings of this karst site, while the samples taken for paleomagnetism belong to a previous sedimentary cycle during the Matuyama Chron. Such interpretations would be in better agreement with the age estimates provided by biostratigraphy and also with the currently accepted chronology for the appearance of Acheulean industries in Western Europe.     

The first record of palaeopathology from the zooarchaeological record of late Pleistocene,southwest Tasmania

Several bones from the Late Pleistocene archaeological site, Kutikina Cave, in southwest Tasmania, show pathological conditions. The majority of these specimens represent Bennett's or red‐necked wallaby (Macropus rufogriseus), with a single specimen of the Common wombat (Vombatus ursinus) recorded. While there has been extensive work in this region of Australia since the early 1980s, this is the first record of palaeopathology from the late Pleistocene of Tasmania, and the first from a human accumulation of predominately macropod material. The palaeopathology raises several questions concerning the mobility of these animals which might have increased their susceptibly to predation, while some of the effected elements are fragmented, suggesting that people utilised these bones despite their deformities. Copyright © 2008 John Wiley & Sons, Ltd.     

Pleistocene foraminifera from Botany Bay,New South Wales

Unconsolidated sediments from a borehole in Botany Bay have been analyzed for their foraminiferal content. Faunas from between 41 and 54 m below sea level are interpreted as Pleistocene in age. The foraminiferal assemblage from 54 m indicates marine conditions of moderately deep water (25–30 m), and a climate warmer than at present. The higher assemblages indicate a more brackish environment

The foraminifera, belonging to 123 species in 67 genera, are tabulated and 24 are discussed and illustrated. Three species, Oolina bifidocostata, Fissurina alatoquadrata and Elphidium botaniensis and one subpecies, Quinqueloculina affinis atrata, are described as new.     

Camelus from the upper pleistocene of Mount Carmel,Israel

A Camelus metapodial, found among the animal bones excavated by Garrod from the Levalloiso-Mousterian of the Tabun Cave (Mount Carmel, Israel), adds to the relatively small number of Camelus finds reported from the Pleistocene of the Near East. The mammal faunas associated with these finds suggest relatively dry open conditions, but not desert conditions. The Tabun specimen is of moderate size, in contrast to the very large Camelus from Far'ah II; two species of camel may thus have been present in the Near East during the Upper Pleistocene.     

Preliminary U-series and Thermoluminescence dating of excavated deposits in Liang Bua sub-chamber,Flores, Indonesia

We report the results of a test excavation of deposits in a limestone cave sub-chamber located beneath the main chamber of Liang Bua, Flores, Indonesia; the discovery site of the small hominin species, Homo floresiensis. Well-preserved remains of extinct Pleistocene fauna and stone artefacts have previously been identified on the surface of a sediment cone within the sub-chamber. Our excavation of the deposits, at the base of the sediment cone in the sub-chamber (to 130 cm depth) yielded only a few fragmentary bones of extant fauna. Uranium/Thorium (U-series or U/Th) dating of soda straw stalactites excavated from 20 to 130 cm in depth demonstrates that the excavated sediments were deposited during the Holocene. Red Thermoluminescence (TL) dating of the sediments at the base of the excavation (130 cm depth) indicates these sediments were last exposed to sunlight at 84 ± 15 ka (thousand years), similar to red TL ages of cave sediments from the main chamber. Together, these results indicate that the surface faunal remains, which are morphologically analogous to Pleistocene finds from the main chamber excavations, were transported to the sub-chamber relatively recently from the main chamber of Liang Bua and probably originated from conglomerate deposits at the rear of the cave and from deposits around the front entrance. There is no evidence for hominin occupation of the sub-chamber, instead it seems to have acted as a sink for cultural materials and fossil remains transported from the surface via sinkholes. Despite the small number of finds from the test excavation, it is possible that more extensive excavations may yield additional transported cultural and faunal evidence at greater depths.