Getting “Out of Africa”: Sea Crossings,Land Crossings and Culture in the Hominin Migrations

Palaeoanthropologists and archaeologists have advanced a wide range of explanatory narratives for the various movements of Homo erectus/Homo ergaster, and the first modern Homo sapiens, “Out of Africa”—or even back again. The application of Occam's razor—a parsimonious approach to causes—gives a more cautious approach. There is nothing in the available evidence that would require the ability for a human water crossing from Africa before the later Pleistocene, whether across the Strait of Gibraltar, the Sicilian Channel or the southern Red Sea (Bab el-Mandab). A parsimonious narrative is consistent with movements across the Sinai peninsula. The continuous arid zone from northern Africa to western Asia allowed both occupation and transit during wet phases of the Pleistocene; there is no requirement for a “sponge” model of absorption followed by expulsion of human groups. The Nile Valley as a possible transit route from East Africa has a geological chronology that could fit well much current evidence for the timing of human migration. The limited spatial and temporal opportunities for movements “Out of Africa,” or back again, also puts particular difficulties in the way of the gene flow required for the multiregional hypothesis of the development of modern Homo sapiens.     

The southern dispersal hypothesis and the South Asian archaeological record: Examination of dispersal routes through GIS analysis

This research advances a model for coastal-based dispersals into South Asia during oxygen isotope stage (OIS) 4. A series of GIS-based analyses are included that assess the potential for expansions into the interior of South Asia, and these results are compared with known archaeological signatures from that time period. The results suggest that modern Homo sapiens could have traversed both the interior and coastlines using a number of routes, and colonized South Asia relatively rapidly. Use of these routes also implies a scenario in which modern H. sapiens, by either increased population growth or competitive ability, may have replaced indigenous South Asian hominin populations.     

Midden or Molehill: The Role of Coastal Adaptations in Human Evolution and Dispersal

Coastal adaptations have become an important topic in discussions about the evolution and dispersal of Homo sapiens. However, the actual distribution and potential relevance of coastal adaptations (broadly, the use of coastal resources and settlement along shorelines) in these processes remains debated, as is the claim that Neanderthals exhibited similar behaviors. To assess both questions, we performed a systematic review comparing coastal adaptations of H. sapiens during the African Middle Stone Age (MSA) with those of contemporaneous Neanderthals during the European Middle Paleolithic. In both species, systematic use of marine resources and coastal landscapes constitutes a consistent behavioral signature over?~?100,000 years (MIS 6–3) in several regions of Africa and Europe. We found more similarities than differences between Neanderthals and modern humans, with remaining disparities all in degree rather than kind. H. sapiens exploited a wider range of marine resources—particularly shellfish—more intensively. MSA shellfish-bearing sites are also more often associated with intense occupations on coastal landscapes, and more evidence of complex material culture such as shell beads. In terms of broader ramifications, Pleistocene coastal adaptations are best conceived of as an ‘add-on’ to previous adaptive strategies, complementing more frequently exploited inland resources and landscapes. Still, Neanderthals and modern humans increased their dietary breadth and quality, and added options for occupation and range expansion along coastlines. Potential evolutionary implications of these multi-generational behaviors include higher intakes of brain-selective nutrients as a basis for neurobiological changes connected to increased cognitive capacities, but also greater reproductive success, dispersal abilities and behavioral flexibility. Whether gradual differences between modern humans and Neanderthals stimulated different evolutionary trajectories is a question for future research.

     

New radiometric ages for the Fauresmith industry from Kathu Pan,southern Africa: Implications for the Earlier to Middle Stone Age transition

The Fauresmith lithic industry of South Africa has been described as transitional between the Earlier and Middle Stone Age. However, radiometric ages for this industry are inadequate. Here we present a minimum OSL age of 464 ± 47 kyr and a combined U-series–ESR age of 542₋₁₀₇⁺¹⁴⁰ kyr for an in situ Fauresmith assemblage, and three OSL ages for overlying Middle and Later Stone Age strata, from the site of Kathu Pan 1 (Northern Cape Province, South Africa). These ages are discussed in relation to the available lithostratigraphy, faunal and lithic assemblages from this site. The results indicate that the Kathu Pan 1 Fauresmith assemblage predates transitional industries from other parts of Africa e.g. Sangoan, as well as the end of the Acheulean in southern Africa. The presence of blades, in the dated Fauresmith assemblages from Kathu Pan 1 generally considered a feature of modern human behaviour ( McBrearty and Brooks, 2000, The revolution that wasn't: a new interpretation of the origin of modern human behavior, J. Human Evolution 39, 453–563),-provides evidence supporting the position that blade production in southern Africa predated the Middle Stone Age and the advent of modern Homo sapiens.     

The Middle Stone Age south of the Limpopo River

Current interest in the origins of anatomically modernHomo sapiens has focused attention on early modern human remains and related archaeological materials associated with the southern African Middle Stone Age. While the anatomically modern status and a Last Interglacial or later age for the human fossils enjoy general support, issues related to the definition of the Middle Stone Age, its dating, and the interpretation of human behavior lack consensus. Available evidence suggests that the anatomically modern human skeleton appeared well before many aspects of the subsistence and symbolic behavior that characterize recent foragers and that Middle Stone Age technology persisted longer in southern Africa than its northern hemisphere counterpart.     

Population relationships of later Pleistocene hominids: A multivariate study of available crania

The Neanderthal populations of the Upper Pleistocene have been regarded by a number of anthropologists as direct ancestors to modern man. Results of multivariate analyses conducted in this study suggest that early Neanderthal crania were morphologically more like modern Homo sapiens than were the later Neanderthals. The implications of these results are discussed in the light of archaeological evidence and comparative studies of Pleistocene crania from outside Europe.     

Lower and middle paleolithic of northern Asia: Achievements,problems, and perspectives

Considerable progress has been made in recent years in the study of the Lower and Middle Paleolithic of northern Asia. There is growing evidence for initial human occupation before 700,000 years ago—as early as elsewhere in Asia—and for a very early adaptation to the arctic desert environment. New models of Lower Paleolithic settlement involve expansion and reduction of occupation in response to climatic variation, rather than simple colonization followed by steady occupation. The Middle Paleolithic of northern Asia is better documented, including actual finds of archaicHomo sapiens. The transition to the Upper Paleolithic seems to involve the survival of earlier cultural traits, but the mechanisms and processes are not well understood. Further significant knowledge concerning these periods awaits the development of common methodologies for classification and analysis.     

Pleistocene connexions between Africa and Southwest Asia: an archaeological perspective

This paper investigates the archaeological evidence for direct prehistoric connexions between Southwest Asia and Africa during the Palaeolithic period. The importance of the ‘Levantine Corridor’ bridging the two continents is stressed in describing the dispersal ofHomo erectus. The continental bridge could have been through the Nile Valley during the Late Acheulian and the emergence of earlyHomo sapiens. Upper Palaeolithic and especially Epi-Palaeolithic dispersal during the terminal Pleistocene are discussed. The role of Egyptian-Levantine connexions in the context of the complex situation which led to sedentism is stressed.     

Pleistocene Water Crossings and Adaptive Flexibility Within the Homo Genus

Pleistocene water crossings, long thought to be an innovation of Homo sapiens, may extend beyond our species to encompass Middle and Early Pleistocene Homo. However, it remains unclear how water crossings differed among hominin populations, the extent to which Homo sapiens are uniquely flexible in these adaptive behaviors, and how the tempo and scale of water crossings played out in different regions. I apply the adaptive flexibility hypothesis, derived from cognitive ecology, to model the global data and address these questions. Water-crossing behaviors appear to have emerged among different regional hominin populations in similar ecologies, initially representing nonstrategic range expansion. However, an increasing readiness to form connections with novel environments allowed some H. sapiens populations to eventually push water crossings to new extremes, moving out of sight of land, making return crossings to maintain social ties and build viable founder populations, and dramatically shifting subsistence and lithic provisioning strategies to meet the challenges of variable ecological settings.

     

The Upper Palaeolithic Lithic Industry of Nazlet Khater 4 (Egypt): Implications for the Stone Age/Palaeolithic of Northeastern Africa

Between Marine Isotope Stage (MIS) 4 and 2, Northeast Africa witnessed migrations of Homo sapiens into Eurasia. Within the context of the aridification of the Sahara, the Nile Valley probably offered a very attractive corridor into Eurasia. This region and this period are therefore central for the (pre)history of the out-of-Africa peopling of modern humans. However, there are very few sites from the beginning of the Upper Palaeolithic that document these migration events. In Egypt, the site of Nazlet Khater 4 (NK4), which is related to ancient H. sapiens quarrying activities, is one of them. Its lithic assemblage shows an important laminar component, and this, associated with its chronological position (ca. 33 ka), means that the site is the most ancient Upper Palaeolithic sites of this region. The detailed study of the Nazlet Khater 4 lithic material shows that blade production (volumetric reduction) is also associated with flake production (surface reduction). This technological duality addresses the issue of direct attribution of NK4 to the Upper Palaeolithic.     

The significance of the fossil hominid skull from Petralona,Greece

Evidence is accumulating that the hominid cranium found in the Petralona cave in 1960 is associated with cave deposits of middle Pleistocene age. If this is so, the fossil is the most complete middle Pleistocene cranium yet discovered and provides important morphological, metrical and radiographic information on the possible evolutionary transition from Homo erectus to Homo sapiens. The classification of the specimen is discussed and it is suggested that a grade system within Homo spaiens should be erected. The Petralona fossil would be allocated to Homo sapiens grade 1 rather than to Homo erectus or to a subspecies of Homo sapiens.     

The phylogenetic significance of the Middle Pleistocene Narmada hominin cranium from central India

A parsimony analysis of the Narmada cranium from central India and a number of other demes and species of Homo concludes that the Narmada hominin shares a closer relationship with the European Steinheim specimen, than with Asian H. erectus or H. pekinensis. This suggests that the population represented by the Narmada cranium is likely to have had its origins in Europe rather than in Asia. Overall the available evidence supports an ‘Out of Africa’ scenario, as the early Asian hominins belong to a distinct clade which has no extant descendants and thus appear to represent an evolutionary ‘dead end’. The later African and European hominins are defined by a clade including early H. sapiens from Africa. The main difference between these two distinct clades is that the H. erectus lineage is defined by increasing degrees of neuro‐orbital disjunction associated with increased anterior cranial base extension, while the lineage leading to early H. sapiens is characterized by the opposite condition of reduced neuro‐orbital disjunction associated with increased anterior cranial base flexion. While there is also evidence of differential patterns of head and neck musculature between these two clades, they are of secondary importance. Preliminary dating of bovid remains found in association with the hominin cranium by the gamma spectrometric U‐series dating technique suggests a tentative minimum age of not less than 236,000 years. This is in agreement with evidence from biostratigraphical studies of the Boulder Conglomerate, which place these deposits in the Middle Pleistocene. Copyright © 2004 John Wiley & Sons, Ltd.     

Did Modern Human Dispersal Take a Coastal Route into India? New Evidence from Palaeolithic Surveys of Kachchh,Gujarat

The Indian Ocean coastline is argued to have been a critical route of modern human dispersal from Africa, introducing Late Palaeolithic industries into South Asia, but a dearth of evidence has prevented a direct evaluation of this. Kachchh (Gujarat, India), located immediately east of the Indus Delta, is an important setting to appraise the Palaeolithic occupation of the western Indian coastline. Targeted survey of Late Pleistocene sediments there found widespread evidence for occupation of Kachchh during the Late Pleistocene: Middle Palaeolithic and possibly Late Acheulean lithic artifacts. The conspicuous absence of Late Palaeolithic industries in these Late Pleistocene deposits, with their presence only noted in Holocene contexts, does not support the model of modern human expansions into India with these industries via the coastal route. We evaluate these results in the context of current debates regarding Late Pleistocene hominin demography, adaptation, and expansions.     

New Human Remains of Archaic Homo sapiens and Lower Palaeolithic Industries from Visogliano (Duino Aurisina, Trieste, Italy)

New human remains have been found during recent excavations conducted in the lower section of Visogliano dolina (Trieste, Italy). They are represented by teeth and a fragment of particularly robust mandible that for their morphological and morphometrical characters can be generically attributed to archaic Homo sapiens. The lithic industry is formed by flakes, carinated points, scrapers, choppers and chopping tools and is characteristic of the Italian Pebble Culture. Sediments, pollen and faunal remains indicate that this part of the deposit was formed during interglacial climatic conditions in the early, but not initial phase of Middle Pleistocene (isotopic stadium 13 or 11).     

Taxonomic reassignment of the Paleolithic human navicular from Cueva de los Torrejones (Guadalajara,Spain)

Evidence of human activity and hominin remains are very scarce inland on the Iberian Peninsula. This fact raises the issue of the scarcity of evidence that Paleolithic Homo sapiens occupied this area outside of the littoral margins (Atlantic, Cantabrian, and Mediterranean coasts). Here, we comparatively describe a human right adult navicular bone recovered in the Cueva de los Torrejones site, located in the village of Tamajón (Guadalajara, Spain). This fossil was preliminarily established as belonging to Homo cf. neanderthalensis, due to the late Pleistocene faunal association, mainly because of the presence of Crocuta crocuta and Panthera pardus. The metrical and morphological study of the navicular T93-S3.27 from Cueva de los Torrejones clearly differentiates it from Neandertals and their ancestors, the hominins from Sima de los Huesos, allowing for this fossil to be taxonomically assigned with confidence as H. sapiens. The navicular from the Cueva de los Torrejones is absolutely and relatively medio-laterally narrow with a low wedging index as those of fossil and modern H. sapiens, and clearly different of Neandertals. The increased discoveries and publications of new naviculars belonging to genus Homo, together with the findings of P. pardus and C. crocuta in more recent chronologies in the Iberian Peninsula, are compatible with this reevaluation. We propose a probable chronology for this fossil between 12 and 15 ka and ca. 25 ka, based on the biostratigraphy and the oldest presence of H. sapiens in the Iberian Peninsula. This work confirms the human presence within the Iberian Peninsula during the Upper Paleolithic and reopens the question of the peopling of the inner Peninsula during this period.     

The lower paleolithic of the Near East

The Near East forms the geographic crossroads between Africa, Asia and Europe and was certainly a main route for the dispersal of Homo erectusinto Eurasia. The study of Lower Paleolithic sites in this region and in the neighboring Caucasus area sheds some light on several potential colonization events. Sites such as Ubeidiya (Jordan Valley) and Dmanisi (Georgia) suggest the early sorties took place around 1.4-1.0 Ma. Despite the lack of radiometric dates, sequences of raised beaches, marine deposits, river terraces, and paleolake formations have enabled various investigators to identify several series of major aggradation and erosion periods within the Pleistocene. Lithic assemblages derived from a few systematic excavations and collections from stratigraphically dated outcrops led to a threefold subdivision of the Acheulian sequence into the Lower, Middle, and Upper Acheulian. The study of nonbiface assemblages, however, has not resolved the question of whether these assemblages deserve inclusion as separate entities or should be viewed as sites within the Acheulian settlement pattern. While the typotechnological definitions of each major phase can be compared to what is known from other regions of the Old World, the Acheulo-Yabrudian (or the Mugharan Tradition) is seen as a local entity. Rare human remains and scarce data concerning subsistence activities do not warrant a comparative discussion with what is known from African and some European sites.     

Middle pleistocene adaptations in Central Europe

The biological evolution documented in the fossil and archaeological record of Central Europe between about 700,000 and 100,000 years ago reflects the transition fromHomo erectus to the earliestHomo sapiens. These populations created different types of industries, ranging from assemblages of small artifacts and industries with simple pebble tools to standardized units such as the Evolved Acheulean and the Early Mousterian. The industries of the Last Interglacial (Taubachian) show some regression in technological standardization and a reappearance of small artifacts. Other aspects of cultural adaptation (settlements, dwellings, hunting activity, symbolic and aesthetic expressions) are also summarized. It is suggested that the impact of climate and climatic changes on human settlement and behavior was greater in Central Europe (where two glaciations advanced from both north and south) than in the Mediterranean and Western Europe. The general trend of cultural veolution was toward greater technological and social complexity, but its course was not unilinear.     

The origins of lithic projectile point technology: evidence from Africa,the Levant,and Europe

Projectile weaponry is a human cultural universal, but its origins and antiquity remain poorly understood. Stone- and bone-tipped projectile weapons have long been treated as emergent features of the "Upper Paleolithic" behavioral revolution. Recently it has been proposed that projectile technology was in widespread use among Homo sapiens populations in Africa during Middle Stone Age (MSA) times. One obstacle to researching the origins of projectile point technology is that the criteria archaeologists employ for recognizing plausible and implausible stone projectile points are largely subjective (overall tool shape, microwear traces). Tip cross-sectional area (TCSA) is a ballistically significant dimension that works well at discriminating North American stone projectile points (spearthrower dart tips and arrowheads) from spear points. This paper compares the TCSA values of ethnographic North American stone projectile points to hypothetical Middle and Upper Paleolithic stone projectile points from Africa, the Levant, and Europe. The results of this comparison do not support the hypothesis of widespread use of stone-tipped projectiles in Africa, the Levant, or Europe prior to 40 Ka. In the New World and in Australia, where we have the richest ethnographic record of stone projectile point use, these implements are largely employed in big-game hunting and in warfare. One or both of these factors may have played a role in the widespread adoption of stone projectile point technology after 40 Ka.     

Lithostratigraphy of Border Cave, KwaZulu, South Africa: a Middle Stone Age sequence beginning c. 195,000 b.p.

Border Cave is well-known for its Middle Stone Age (MSA) sequence and associated hominids, as well as for the earliest demonstrable Later Stone Age (LSA) (c. 38,000 b.p.) strata in southern Africa. Detailed lithostratigraphic and sedimentological study permits identification of 8 Pleistocene sedimentary cycles, including 6 major cold phases and 2 intervening weathering horizons. The 2 youngest cold phases are associated with the LSA and have 8 ¹⁴C dates 38,600-13,300 b.p. By gauging sedimentation rates in finer and coarser sediments, duration of sedimentary breaks, and allowing for differential compaction, the excellent radiocarbon framework provided by 28 available ¹⁴C dates can be extrapolated to the 6 cold intervals and 2 palaeosols that are older than 50,000 b.p. These clearly span oxygenisotope stages 4, 5 and 6, placing the base of the MSA deposits at c. 195,000 b.p., Homo sapiens sapiens at c. 90,000–115,000 b.p. and the sophisticated, microlithic “Howieson's Poort” industry at 95,000 b.p. These results require radical reassessment of the age and nature of the MSA complex and of the earliest evolution of anatomically-modern people.