Reconnaissance of Prehistoric Sites in the Red Sea Coastal Region of the Sudan,NE Africa

This paper reports results of a recent Stone Age-focused archaeological survey in the Red Sea coastal region of the Republic of Sudan, northeast Africa. Bifaces (handaxes) are the most conspicuous artifact class encountered during the survey and are characteristic of the Acheulean technocomplex. Other recorded artifact types include points, scrapers, and prepared core products referable to the Nubian and recurrent Levallois methods. Most of the artifact-bearing localities lie landward—outside of the coastal margin—thus, the evidence does not signify direct coastal adaptation per se. Our preliminary findings suggest that multiple Pleistocene-age hominin settlements tied to a terrestrial niche existed in the region. The western margin of the Red Sea occupies a pivotal location, linking the Horn of Africa and the Levant, two vital regions in human evolutionary research. Thus, the Stone Age data from the Sudan region has direct relevance for assessing hominin dispersal routes out of Africa.     

Did Modern Human Dispersal Take a Coastal Route into India? New Evidence from Palaeolithic Surveys of Kachchh,Gujarat

The Indian Ocean coastline is argued to have been a critical route of modern human dispersal from Africa, introducing Late Palaeolithic industries into South Asia, but a dearth of evidence has prevented a direct evaluation of this. Kachchh (Gujarat, India), located immediately east of the Indus Delta, is an important setting to appraise the Palaeolithic occupation of the western Indian coastline. Targeted survey of Late Pleistocene sediments there found widespread evidence for occupation of Kachchh during the Late Pleistocene: Middle Palaeolithic and possibly Late Acheulean lithic artifacts. The conspicuous absence of Late Palaeolithic industries in these Late Pleistocene deposits, with their presence only noted in Holocene contexts, does not support the model of modern human expansions into India with these industries via the coastal route. We evaluate these results in the context of current debates regarding Late Pleistocene hominin demography, adaptation, and expansions.     

Why is there a lack of Mode 3 Levallois technologies in East Asia? A phylogenetic test of the Movius–Schick hypothesis

The ‘Movius–Schick hypothesis’ claims that an absence of Levallois (Mode 3) technologies in East Asia is due to the lack of a strong ancestral Acheulean (Mode 2) tradition in that region. Hence, this hypothesis is based on the assumption that similarities between Acheulean handaxes and Levallois cores can be interpreted as being phylogenetically homologous (i.e. due to common technological ancestry) as opposed to being homoplasic (i.e. due to convergent technological evolution). Here, the phylogenetic basis of this hypothesis is tested using a formal cladistic procedure. Under the framework of an ‘iterative’ approach to phylogenetic analysis, a series of post-hoc tests and re-evaluations of results follow the initial cladistic analysis. Results of these combined analyses indicate that morphological similarities between Mode 2 Acheulean handaxes and Mode 3 Levallois cores can, most parsimoniously, be seen as phylogenetically homologous. Hence, these results support the tenets of the ‘Movius–Schick’ hypothesis in suggesting that a lack of Levallois industries in East Asia may be due to the paucity of an ancestral Acheulean tradition in that region. The implications of these phylogenetic analyses for the concept of Palaeolithic ‘Modes’ are discussed. It is further suggested that low demographic levels (i.e. small effective population sizes) in East Asia may have constrained the technological phylogenetic trajectory of East Asia compared with that seen in other regions of the Old World during the Lower Palaeolithic. In addition, it is hoped that several methodological issues discussed here will contribute to the growing field of cultural phylogenetic analysis.     

The Lower Paleolithic of the Arabian Peninsula: Occupations,Adaptations, and Dispersals

To understand major processes of human evolution during the Plio- Pleistocene, it is necessary to consider the available evidence from key regions of the Old World. The Arabian peninsula is often depicted as a key geographic route for hominin dispersals in Out of Africa models, although the available primary evidence is rarely cited. And yet, significant Lower Paleolithic assemblages have been identified in many parts of the peninsula, including in areas near the Strait of Bab al Mandab. The presence of Oldowan-like and Acheulean assemblages may reflect at least two major dispersals outside of Africa. Acheulean localities are particularly abundant on the Arabian peninsula, and variations in stone tool manufacturing techniques and tool-type frequencies may reflect temporal changes in assemblages. Although there is good potential to chronometrically date sites in the Arabian peninsula, absolute dating methods have not been adequately applied, most temporal reconstructions relying on typotechnological change. If the Arabian peninsula is to provide solid evidence for understanding hominin adaptation and dispersal patterns, future fieldwork in secure depositional contexts needs to be conducted to overcome current limits in dating and environmental reconstructions.     

Shifts Along a Spectrum: A Longitudinal Study of the Western Eurasian Realized Climate Niche

ABSTRACT

Climate niches that modern humans and earlier hominin ancestors occupied have changed dramatically over time, but the extent of those changes has gone largely undocumented. This study investigates the manner in which the realised hominin climate niche has expanded, contracted, or stayed stationary across four time periods (Last Interglacial, Last Glacial Maximum, Mid-Holocene, and 1950–2000) in Western Eurasia. Using spatially gridded general circulation model data and site locations this study examines climate variables from archaeological sites and current Western Eurasian cities to describe both the regional Western Eurasian fundamental and realised climate niches. Changes between the three prehistoric periods and modern-day time period are analysed by calculating each realised niche breadth, overlap, position, and variance. Results indicate that as global temperatures cooled from the Last Interglacial to Last Glacial Maximum, populations expanded their climate niche breadth beyond that of earlier Neanderthal groups, shifting toward regions with less seasonal variation. Conversely, Mid-Holocene humans, who saw the proliferation of both agriculture and population, contracted their realised climate niche space. The contraction and expansion of realised climate niche space illustrates how hominins have evolved the capacity to shift their niche through changes to their subsistence strategy and adaptations to overall climatic conditions.     

Pleistocene Water Crossings and Adaptive Flexibility Within the Homo Genus

Pleistocene water crossings, long thought to be an innovation of Homo sapiens, may extend beyond our species to encompass Middle and Early Pleistocene Homo. However, it remains unclear how water crossings differed among hominin populations, the extent to which Homo sapiens are uniquely flexible in these adaptive behaviors, and how the tempo and scale of water crossings played out in different regions. I apply the adaptive flexibility hypothesis, derived from cognitive ecology, to model the global data and address these questions. Water-crossing behaviors appear to have emerged among different regional hominin populations in similar ecologies, initially representing nonstrategic range expansion. However, an increasing readiness to form connections with novel environments allowed some H. sapiens populations to eventually push water crossings to new extremes, moving out of sight of land, making return crossings to maintain social ties and build viable founder populations, and dramatically shifting subsistence and lithic provisioning strategies to meet the challenges of variable ecological settings.

     

旧石器时代亚欧大陆高纬度地区人群的扩散及其对中国旧石器文化格局的影响

欧亚大陆高纬度地区气候与资源环境迥异于人类的起源地——热带非洲。人类需要具备更多的文化适应手段才能适应高纬度地区的生存。在旧石器时代早中期,人类向该地区的迁移扩散极为有限,而旧石器时代晚期,随着技术、社会组织等文化层面的进步,人类开始广泛向高纬度地区扩散,打破了东亚地区与欧亚大陆西侧的文化隔离屏障,对中国旧石器时代西北、东北及华北地区的文化格局产生了不同程度的影响。本文拟通过人类适应的视角,结合考古材料,分析人类向欧亚大陆高纬度地带的扩散历程及由此而引发的人类迁徙、交流对中国旧石器时代文化面貌格局产生的具体影响。     

The Muddle in the Middle Pleistocene: The Lower–Middle Paleolithic Transition from the Levantine Perspective

The terms Lower Palaeolithic and Middle Palaeolithic represent research constructs within which cultural evolution and prehistoric hominin behaviours can be studied, with the transition usually understood as marking a watershed in our evolution: an adaptation with a million-year record of success that gives way to something new. The interpretation of the Lower Palaeolithic Acheulian technocomplex is usually understood as a period of cultural stasis that extends over much of Africa and Eurasia, principally associated with Homo erectus. Those innovations that can be observed occur widely separated from one another in space and time. Yet a closer and more detailed examination of the Middle Pleistocene records from East Africa, southern Africa, Europe and the Levant reveals significant variation in cultural repertoires. A kind of paradox emerges, in which an Old World Lower Palaeolithic, apparently lacking an overall dynamic of distinctive and directed change in terms of cumulative variation over time, nevertheless culminates in a transition which sees the universal appearance of the Middle Palaeolithic. The two main hypotheses that have been advanced to explain the global transition, which happens essentially synchronously, appear mutually exclusive and contradictory. One view is that altered climatic-environmental constraints enabled and encouraged an ‘Out-of-Africa’ dispersal (or dispersals) of a new type of genus Homo. This cultural replacement model has been challenged more recently by the alternative hypothesis of accumulating but unrelated and temporally non-linked regional, and in fact potentially autochthonous, processes. The Levant, by virtue of its position bridging Africa and Eurasia (thus being the region into which any out-of-Africa groups would have had first to disperse into), must be seen as a critical region for assessing the relative merits of these competing hypotheses. This paper deals with the Lower–Middle Paleolithic boundary in the Levant within a long temporal perspective. The Middle Pleistocene record in the Levant enables us to examine the amplitude of variation within each techno-complex, as well as to question whether there are diachronic changes in the amplitude of techno-typological variations as well as changes in the manner by which they appear in the record. The results carry significant implications for understandings of demographic and societal processes during the Lower–Middle Paleolithic transition in the Levant.     

A 3D morphometric analysis of surface geometry in Levallois cores: patterns of stability and variability across regions and their implications

Levallois cores and products were manufactured by hominin populations distributed across wide regions of Africa and Eurasia. Levallois technology remains an important focus for research in Palaeolithic archaeology, yet quantitative morphological comparisons of Levallois core morphology from different regions remain rare. Here, utilizing Levallois cores from Africa, the Near East, Europe, and the Indian subcontinent, patterns of morphological variability in the shape of the Levallois flaking surface and core outline (margin) shape were examined for patterns of variability and stability across regions using 3D geometric morphometrics. The multivariate statistical shape analyses undertaken revealed a clear pattern: that is, the greatest levels of shape variability in Levallois cores is evident in the form of their outline (planform) shape. Conversely, the geometrical relationship between the margin of the Levallois cores and their topological surface morphology was relatively uniform. This pattern of variability was evident in terms of variation both across regions and between cores from the same locality. These results indicate that the outline form of such cores was a less important variable than the geometric/topological properties of the surface morphology and, in particular, the relationship between the margin of the core and those variables. These results have implications for why it has been reported that replicating such cores in modern experiments is a particularly difficult task. The specific interrelationship between the geometric properties of the core and the core margin provide further evidence that Levallois core technology would be unlikely to emerge from the context of opportunistic migrating platform reduction strategies (such as those seen in many Mode 1 industries). If, as is widely suggested, Levallois cores were deliberate products in Pleistocene contexts, these results also hint that relatively sophisticated means of social transmission (i.e. teaching) may have been required to sustain their production over time and space.     

The Possible Time and Region of Host Switches of Ancient Malaria Parasites with Reference to the Pliocene–Quaternary Archaeological Sites in Africa

About 96% of all malaria deaths occur in Africa, and the malignant falciparum malaria also originated on the continent. Although falciparum malaria only appeared in the Holocene period, it can be hypothesized that the transfer of malaria parasites from other primates to humans occurred several times in history parallel to human evolution. This study develops the model that examines the possible coexistence of the potential original host apes, human ancestors, and the diverse anopheline mosquito species; and how, where, and when the host switch of these parasites from great apes to humans occurred. Based on the Pliocene-early Pleistocene archaeological sites, it was found that certain early hominin populations could have lived in malaria areas where the anopheline mosquito fauna was moderately diverse. The people of the Lupemban Culture, as well as the Greenlandian and Northgrippian human populations of East and West-Central Africa, lived close to the high diversity of anopheline fauna and the territories of such great apes as Gorilla gorrilla. African mid-Holocene cultures likely came in contact with gorilla populations — the original hosts of Plasmodium falciparum — along the coasts of the Gulf of Guinea and the East African Rift Valley during their migration to southern Africa. The host switch of the ancestor of the falciparum malaria parasite likely occurred in these regions.

     

The application of biometry and LA-ICP-MS to provenance isolated bones: a study of hominin remains from Oumm Qatafa Cave,Judean Desert

Three hominin phalanges were recently identified in old faunal collections attributed to the Late Acheulean (Layer D2) dated to ca. 213 Kya, from Oumm Qatafa Cave (Judean Desert), a site excavated in 1928–1949. In terms of general appearance (colour, patina and adhering sediment), these specimens resembled the fauna with which they were found, but the likelihood of stratigraphic problems especially in old excavations, the presence of 4th millennium b.c. burials in the topmost Layer A at the site and the absence of any mention of Late Acheulean hominin remains in publications relating to the site prompted us to verify that the phalanges were indeed in situ. Osteometric examination showed the phalanges to be indistinguishable from those of Middle Paleolithic Levantine anatomically modern humans (AMHS) as well as Upper Paleolithic and recent populations, thus contributing little to the resolution of their provenance. To further investigate this issue, we compared the elemental composition of the phalanges to that of fauna from the same and overlying archaeological layers using non-destructive laser ablation–inductively coupled plasma mass spectrometry (LA-ICP-MS). The results showed a close resemblance in elemental composition between the phalanges and fauna from Layer D2, implying that they are in situ. This would indicate either an early occurrence of AMHS in the region or the presence of an ancestral archaic Homo. We propose that LA-ICP-MS offers a useful minimally invasive method for provenancing isolated human and faunal remains.     

Anatomy,behavior, and modern human origins

The fossil record suggests that modern human morphology evolved in Africa between 150,000 and 50,000 years ago, when the sole inhabitants of Eurasia were the Neanderthals and other equally nonmodern people. However, the earliest modern or near-modern Africans were behaviorally (archaeologically) indistinguishable from their nonmodern, Eurasian contemporaries, and it was only around 50,000-40,000 years ago that a major behavioral difference developed. Archaeological indications of this difference include the oldest indisputable ornaments (or art broadly understood); the oldest evidence for routine use of bone, ivory, and shell to produce formal (standardized) artifacts; greatly accelerated variation in stone artifact assemblages through time and space; and hunting-gathering innovations that promoted significantly larger populations. As a complex, the novel traits imply fully modern cognitive and communicative abilities, or more succinctly, the fully modern capacity for Culture. The competitive advantage of this capacity is obvious, and preliminary dates suggest that it appeared in Africa about 50,000 years ago and then successively in western Asia, eastern Europe, and western Europe, in keeping with an African origin. Arguably, the development of modern behavior depended on a neural change broadly like those that accompanied yet earlier archaeologically detectable behavioral advances. This explanation is problematic, however, because the putative change was in brain organization, not size, and fossil skulls provide little or no secure evidence for brain structure. Other potential objections to a neural advance in Africa 50,000-40,000 years ago or to the wider Out-of-Africa hypothesis, include archaeological evidence (1) that some Neanderthals were actually capable of fully modern behavior and (2) that some Africans were behaviorally modern more than 90,000 years ago.     

Is the Soanian techno-complex a Mode 1 or Mode 3 phenomenon? A morphometric assessment

The Soanian is traditionally seen as one of the major (non-Acheulean) Palaeolithic techno-complexes of the Indian subcontinent. Over several decades comparisons of Soanian assemblages have been made with the non-bifacial industries of East Asia and north-west Europe. The chronological status and typo-technological relationship(s) of the Soanian to other Palaeolithic industries have been the subject of much debate. When first named and described the Soanian was considered to contain evidence of Mode 3 Levallois-style core reduction. However, in recent years, the potential Mode 3 component of the Soanian has largely been ignored, and the techno-complex is described under various guises as a core/flake or ‘Mode 1’ techno-complex. Here, a comparative morphometric assessment of selected Soanian cores and other Palaeolithic nuclei is undertaken, to test the hypothesis that this industry contains a definite Mode 3 Levallois element. Discriminant Function Analyses (DFA) of morphometric variables provide robust evidence that at least part of the Soanian techno-complex contains Mode 3 Levallois cores. The implications of these analyses for the relationship between the Soanian and the Acheulean, and the relevance of the Soanian in considerations of the Movius Line are also discussed.     

Pole to Pole. Archaeology and Adaptation in the Middle Pleistocene at Opposite Ends of the Acheulean World

The concept of cultural evolution in the Acheulean is linked to a number of important anatomical, phylogenetic, adaptive, cultural and cognitive research questions. Yet there are very few studies which are able to demonstrate advancements in material culture (technological and/or conceptual) supported by large bodies of empirical data. Derek Roe's work at Olduvai Gorge is one of the few exceptions and here, in my opinion, raw material considerations affect the evidence of increasing diachronic sophistication (Leakey and Roe 1994). Otherwise, evidence for such material culture evolution is largely anecdotal. This study takes large bodies of handaxe data from South Africa and Britain, and using dated assemblages, wherever possible, explores the idea of hominins paying increasing attention to shape and morphological regularity (not symmetry) over time. The results give no reason to believe such a diachronic trend exists. These data are set against the latest research in chronostratigraphic studies and hominin taxonomy.     

The Middle Stone Age human remains from Diepkloof Rock Shelter (Western Cape,South Africa)

In the course of recent excavations at Diepkloof Rock Shelter (South Africa), three human remains were found in the Middle Stone Age layers. These human remains are two pedal phalanges (intermediate and distal) from a fifth ray, which belong to the same individual, and a deciduous first lower molar. The layers in which they were found represent the end of the Howiesons Poort and the beginning of post-Howiesons Poort occupation, and are radiometrically dated to MIS 3, between 45 and 65 kyr ago. They are thus very close in time to the assumed period of anatomically modern humans (AMH) dispersal out of Africa into Eurasia, a period for which very few remains are available in the African fossil record. Here, we describe these new remains and compare their morphology and dimensions to samples of African and Eurasian Pleistocene and Holocene humans. Our results show that the human remains from Diepkloof Rock Shelter fall well within the range of variation of AMH. The phalanges are of very small size and indistinguishable from those of recent humans. In addition, our analysis of the dm₁ underlines a large size variation at the end of the MIS 4/beginning of the MIS 3 in the Western Cape. The discovery of human remains at Diepkloof Rock Shelter, albeit limited to three small bones, enriches a poorly represented chronological period of the African hominin fossil record.     

Acheulean variation and selection: does handaxe symmetry fit neutral expectations?

It has been suggested that the property of symmetry observed in Acheulean handaxes was selected for functional, adaptive or social and aesthetic reasons. However, selectionist accounts of variation may be contrasted with the approach taken by population geneticists to molecular variation. Population geneticists always first assume a neutral pattern of variation for molecular data, and only look to non-neutral (e.g., selective) scenarios for pattern and variation in the face of strong evidence against this null model of neutral expectation. Here, using a combination of cultural transmission theory, morphometrics, and the principles of population genetics, (null) neutral expectations for Acheulean handaxe symmetry are tested. The results of the analyses are inconsistent with a null hypothesis of neutral expectation for patterns of handaxe symmetry variation. Rather, the results imply that the property of symmetry in Acheulean handaxes was subject to selection for functional, adaptive or social reasons.     

Craniometric evidence for South African Later Stone Age herders and hunter–gatherers being a single biological population

Later Stone Age (LSA) hunter–gatherers and herders co-existed in South Africa during the last 2000 years. In spite of being the focus of intensive research over the years, the biological status and origins of the herders are still unclear. Did they represent a genetically distinct immigrant population who remained separate from the indigenous hunter–gatherers, or where they indigenous hunter–gatherers who took up herding after contact with herders, probably in northern Botswana? Here, this issue is investigated using craniometric data collected on a large sample of individually dated human crania from coastal LSA context. Mahalanobis distances (D), calculated from the raw metric data, show that there was a small increase in inter-individual craniofacial variation after the introduction of herding at ca. 2000 BP. Here it is argued that this small increase in variation is neither consistent with a large-scale immigration of genetically distinct herders into South Africa, or the long-term co-existence of two genetically distinct populations. Two alternative explanations fit the data better: (1) herding entered South Africa via the small-scale immigration of genetically distinct herders; and (2) local hunter–gatherer populations adopted herding after coming in contact with herders in northern Botswana. While small-scale immigration would not have had a major influence on the local gene pool, it would have increased variation to some extent as immigrants mixed with local populations. If small-scale external gene flow was not a factor in the introduction of herding, secular issues related to the introduction of herding could explain the increased variation in post-2000 BP populations.     

Flakes Crossing the Straits? <Emphasis Type="Italic">Entame</Emphasis> Flakes and Northern Africa–Iberia Contact During the Acheulean

The entame core method was defined after studying the Acheulean bifaces from the site of Ternifine, Algeria. This specialized core method for the production of larges flakes (>10 cm) used for biface blanks involves the detachment of primary large flakes from skillfully selected quartzite cobbles. While technologically simple, a competent selection of raw material and dexterous detachment of the flake resulted in a blank perfectly suitable for the production of bifaces, with minimal further shaping required. This core method resulted in high frequencies of entame blanks in the Ternifine lithic assemblages, as well as from the Iberian Peninsula Acheulean, but not in assemblages from other large flake Acheulean sites. It is suggested that the frequent use of the entame core method common to both North Africa and the Iberian Peninsula indicates similarity in lithic tradition during the Acheulean. This may support the view of North African origin for the Iberian Acheulean during the Middle Pleistocene.