Human consumption of tortoises at Level IV of Bolomor Cave (Valencia, Spain)

The acquisition and consumption of small prey in the pre-Upper Palaeolithic is a highly debated topic at present. For some authors, the systematic obtaining of these animals is only part of the subsistence strategies used by anatomically modern Humans. However, the consumption of small prey dates back to the Plio-Pleistocene chronologies in some sites. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of tortoise consumption is documented not as common for these periods. However, Level IV of Bolomor Cave has clear diagnostic elements to document the acquisition and use of tortoises (Testudo hermanni) for food in the form of: (1) cutmarks on limb bones and ventral surface of the carapace and plastron; (2) presence of burning on tortoise skeleton and shell; (3) elements of anthropogenic breakage on carapace and plastron: percussion pits, percussion notches and impact flakes; and (4) human toothmarks on limb bones. This paper tries to examine the possible patterns in the tortoise consumption sequence from Level IV of Bolomor Cave and improves data on the butchery process and tortoise consumption in the Late Middle Pleistocene.     

Middle Pleistocene bird consumption at Level XI of Bolomor Cave (Valencia, Spain)

The consumption of small prey dates back to the Plio-Pleistocene chronologies in some African sites. However, the systematic acquisition and consumption of small prey in the pre-Upper Palaeolithic times is still a highly debated topic in Europe. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of bird consumption is not as common for these periods. Nevertheless, Level XI (MIS 6) of Bolomor Cave has clear diagnostic elements to document the acquisition and use of birds (Aythya sp.) for food in the form of: (1) cutmarks on bones of both the front and hind limb; (2) presence of burning patterns on the extremities of the bones (areas of the skeleton with less meat); and (3) human toothmarks on limb bones. The capture of birds is classified as quick-flying game in the archaeological sites. The acquiring of fast-running (mostly lagomorphs) and quick-flying small prey requires a sophisticated technology and involves obtaining and processing ways different from those used for large- and medium-sized animals. From this perspective, the aim of this paper is to examine possible patterns in the processing sequence of birds from Level XI of Bolomor Cave and to improve the data on their butchery and human consumption in the Middle Pleistocene of Iberian Peninsula.     

A new experimental study on percussion marks and notches and their bearing on the interpretation of hammerstone-broken faunal assemblages

Preliminary experiments on bone breakage have shown the potential utility of quantifying some of its diagnostic features (percussion marks, percussion notches) for taphonomic inferences about human and carnivore involvement in bone breakage in faunal assemblages. The present study increases the range of experiments undertaken to understand the identification of hammerstone percussion (dynamic loading) and its differences from carnivore bone breakage through tooth pressure (static loading). This study contributes to a better understanding of frequencies of percussion marks, uncovers and quantifies those percussion marks that lack key diagnostic features to be identifiable and that could be mistaken with carnivore tooth marks, quantifies notch types, shows different size ranges for notches on bones from small and large fauna than previously reported, quantifies the proportion of notches bearing percussion marks, and introduces new size data for percussion (impact) flakes and percussion marks. Furthermore, all these variables have been applied to a dual experimental set: one experiment using non-modified hammerstones and another based on the use of modified hammerstones. Results vary considerably according to hammerstone type. Some of these taphonomic variables increase the range of equifinality when identifying marks and notches created by different human and non-human agents. This calls for further caution when using isolated variables and features rather than a holistic approach to make taphonomic inferences.     

Cut marks on the Middle Pleistocene elephant carcass of Áridos 2 (Madrid,Spain)

Áridos 1 and Áridos 2 (Madrid, Spain) are two Middle Pleistocene sites belonging to the isotopic stages 9–11. Both places contain partial carcasses of Elephas (Paleoxodon) antiquus associated to Acheulian stone tools. In this work, the taphonomic study of the elephant remains of Áridos 2 is presented. This study has documented several cut marks on different bones, which indicate bulk flesh and viscerae extraction by Middle Pleistocene hominins. Several arguments are provided to support that at least some of the cut marks were made with handaxes, further suggesting that some of these artifacts were butchering tools in this stage of human evolution. Although cut marks on elephant carcasses have been documented at some Middle Pleistocene sites, very few have been published in detail to allow consideration of their status as hominin-imparted marks. By doing so, the present study provides more evidence of large carcass exploitation by hominins during this period.     

The Lower Acheulian site of Ambrona,Soria (Spain): ages derived from a combined ESR/U-series model

The Lower Paleolithic sites of Ambrona and Torralba (Soria, Spain) are often associated with each other because of their proximity although they do not represent the same morphostratigraphic unit. Systematic and extensive excavations were conducted at these sites by an American team during the 1960s and 1980s. Recently (1989–2000), a Spanish team has reactivated the research by initiating a new interdisciplinary project. In the central sector and west of the Ambrona site a stratigraphy has been established with six units from AS1 at the base to AS6 at the surface, corresponding to fluvial–lacustrine deposits in which abundant fauna associated with Acheulian artefacts were found. The most abundant species were elephant, horse, deer and aurochs. Other species like carnivores occur in small numbers only. The lithic assemblage is relatively sparse and consists of a few bifaces, choppers and several flake tools distributed throughout levels AS1 to AS5. The fauna and the lithic industry therefore document human presence at Ambrona during the Middle Pleistocene. In order to verify the presumed antiquity of the Ambrona deposits, horse teeth sampled in levels AS1, AS2 and AS6 were analysed using combined ESR/U-series (US) methods. The samples from the lower levels (AS1 and AS2) underwent a very recent uranium uptake as indicated by the U-series dates, ranging between 5 and 18 ka. In contrast, a more conventional postmortem uranium uptake was observed in the upper level samples (AS6) in the enamel and dentine tissues, while a light U-leaching was observed in the cementum. The combined ESR/U-series dates obtained on these samples suggest a minimum age of approximately 350 ka, contemporaneous of OSI 9 or the end of OSI 11, for the Ambrona site.     

Interpreting pachyderm single carcass sites in the African Lower and Early Middle Pleistocene record: A multidisciplinary approach to the site of Nadung’a 4 (Kenya)

Nadung’a 4 is one of the single carcass pachyderm sites recorded in East Africa during the Lower and Early Middle Pleistocene. The site has yielded an abundant lithic assemblage in close association with the partial carcass of an elephant. Conjoined pedological, geoarchaeological, spatial, technological, and taphonomical analyses have been carried out to address the relationship between hominids and elephant. The resulting data are consistent with a non-fortuitous association between both categories of remains. The lithic artefacts do not match a classical Acheulean tool-kit, as would be expected for the time period ascribed to the site, and the functional patterns inferred from their analysis make this site radically different from other purported butchery sites. The implications of these original features are discussed.     

Lithic refitting and movement connections: the NW area of level TD10-1 at the Gran Dolina site (Sierra de Atapuerca,Burgos, Spain)

The first lithic refit studies were carried out at the end of 19th century, but the method was not considered an area of real interest to archaeology until quite recently. Today, lithic refitting is applied in a multitude of areas of enquiry including lithic technology, intra-site and inter-site spatial distribution, archaeostratigraphy and formation processes. In this paper, we present a refit study of the lithic materials recovered in the base of the NW sector of level TD10 at the site of Gran Dolina (Sierra de Atapuerca, Spain). We aim to provide information about spatial distribution through identifying activity areas and the internal connections between those areas. Our work resulted in various refits whose connections reveal the movements of the pieces and/or knappers that once occupied the site. Our results also show the importance of studying the set of materials as a whole and the bias involved in analyzing only a small sample. The refits, connections and directions of movement allow us to infer areas in which an activity took place, but these results must be confirmed in future works covering the entire lithic assemblage of TD10.     

The reconstruction of past environments through small mammals: from the Mousterian to the Bronze Age in El Mirón Cave (Cantabria,Spain)

El Mirón is a large cave in the Cantabrian Cordillera of northern Spain that presents a long archaeostratigraphic sequence radiocarbon-dated by over 60 assays to between 41,000 and 2000 BP. The sediments, collected from four areas within the cave and sieved-washed with fine wire meshes, contain microvertebrate remains of fish, frogs, lizards, birds and mammals, of which the latter are most abundant. Preliminary taphonomic analysis suggests that the microvertebrates were naturally collected by owls and (less) small carnivores. Small mammal assemblages are useful for palaeoenvironmental reconstruction because they are linked to particular habitats and are sensitive to environmental changes. The small mammals from El Mirón are ideal for this because sample sizes are large, bone preservation is good, and the stratigraphic sequence is long. In this paper we reconstruct the late Quaternary environments in the Cantabrian region of Spain using small-mammal assemblages from El Mirón Cave. On the basis of the ecologic adaptations of this suite of fauna, the majority still extant, we have identified seven habitat types, which are plotted through time. The evolution of the small mammal assemblages at El Mirón reveals seven major climatic shifts that correspond closely to the climatic changes recognized in the Iberian Peninsula during the last 41 kyr.     

Introduction to the project and excavation of Diepkloof Rock Shelter (Western Cape,South Africa): a view on the Middle Stone Age

This introduction presents the background to the present research project at Diepkloof Rock Shelter, initiated in 1998. It is followed by a series of original papers that were presented in November 2010 at the join 13th PAA Congress (Panafrican Association of Prehistory and Associated Disciplines) and 20th conference of SAfA (Society of Africanist Archaeologists) at the University Cheikh Anta Diop in Dakar (Senegal). These papers were presented in a “Symposium on the MSA sequence of Diepkloof Rock Shelter: a view on the cultural evolution of southern African modern humans” organized by Pierre-Jean Texier, Guillaume Porraz, John Parkington and Jean-Philippe Rigaud. This series of papers is a first attempt at a multidisciplinary reconstruction of the way Middle Stone Age people inhabited the site of Diepkloof and the way they interacted with their environment. The resultant narrative outlines artifactual change through the sequence and discusses the factors that might underlie it.     

Sedimentological and archaeological fabrics in Palaeolithic levels of the South-Eastern Pyrenees: Cova Gran and Roca dels Bous Sites (Lleida, Spain)

The comparative study of sedimentary fabrics in relation to archaeological fabrics in various levels of the sites of Cova Gran de Santa Linya (Middle and Early Upper Palaeolithic) and the Roca dels Bous (Middle Palaeolithic) has made it possible to analyse the formation dynamics of the deposits and determine the extent of disturbance of the archaeological levels by natural processes. To achieve this, diagrams and two- and three-dimensional indices were calculated from the azimuth and dip angle of natural clasts and artefacts. The results indicate that the sedimentary levels were formed mainly by planar fabric shape gravitational processes (low depositional angles), which differ notably from the archaeological fabrics they contain, characterised by a greater degree of isotropy. This difference in the fabrics would reflect human activity and indicates that the archaeological levels both in Cova Gran and Roca dels Bous have not been subject to significant natural modifications, and are preserved in situ.     

The hunted hunter: the capture of a lion (Panthera leo fossilis) at the Gran Dolina site,Sierra de Atapuerca,Spain

Many Pleistocene caves and rock shelters contain evidence of carnivore and human activities. For this reason, it is common to recover at these sites faunal remains left by both biological agents. In order to explain the role that carnivores play at the archaeological sites it is necessary to analyse several elements, such as the taxonomical and skeletal representation, the age profiles, the ratio of NISP to MNI, the anthropogenic processing marks on the carcasses (location and purpose of cutmarks and burning and bone breakage patterns), carnivore damage (digested bones, location and frequencies of toothmarks and bone breakage), length of the long bones, frequencies of coprolites and vertical distribution of the faunal remains, inter alia. From this, the documentation of carnivores in a faunal assemblage with a clear anthropogenic component can be understood from three main phenomena: (1) the carnivores as accumulators and the use of the site as a den; (2) carnivores as scavengers of hominid refuse and; (3) carnivores as hominids’ prey. Of these three phenomena, the last one is the least documented at the Middle Pleistocene sites. From this perspective, here we present the case of the anthropogenic use of a lion (Panthera leo fossilis) from level TD10-1 of Gran Dolina (MIS 9, Sierra de Atapuerca, Burgos, Spain). The lion bone remains show signs of direct interaction between this big cat and human groups that occupied Gran Dolina in these chronologies. From this perspective, the aim of this paper is to contribute to the knowledge of the role developed by large carnivores in the anthropogenic contexts and to provide data on human use of these predators at the European Middle Pleistocene sites.     

Digital imaging technology and experimental archeology: a methodological framework for the identification and interpretation of fire modified rock (FMR)

This paper presents the results of a series of experiments for the identification and analysis of fire modified rock (FMR). FMR is a common but frequently overlooked artifact type. Experiments were conducted simulating the effects of different hypothetical burning scenarios on rocks similar to those found in a South African Middle Stone Age site. A digital imaging method was then used to quantify FMR color values, designed to limit intra-analyst bias. Statistical tests and a blind test suggest that unburned rocks and experimental FMR can be separated statistically based on physical appearance. Two burning scenario models, based on measured experimental data were applied to archaeological FMR from a South African Middle Stone Age site named Pinnacle Point 5-6 (PP5-6) and show that the archaeological samples are not statistically different from a simulated campfire and possibly a lithic raw material heat-treatment fire.     

Pitted stone cobbles in the Mesolithic site of Font del Ros (Southeastern Pre-Pyrenees,Spain): some experimental remarks around a controversial tool type

The presence of cobbles with activity-related marks in the Mesolithic site of Font del Ros (Berga, Spain), and in particular one group of artefacts – pitted stones – raises problematic issues associated with the characterization of percussion activities. Although these artefacts have generated an extensive bibliography on ethological, ethnographic, ethnoarchaeological and archaeological levels, various questions persist in relation to their possible contextual function. In this paper we present the results of an experimental programme in which three types of activities that could create pitted stones are reproduced: bipolar knapping of vein quartz, hazelnut cracking, and hazelnut grinding. The aim of this experimental programme is to describe marks and use-wear traces related to such activities.     

A new saucrosmylid lacewing (Insecta,Neuroptera) from the Middle Jurassic of Daohugou,Inner Mongolia,China

Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A., 2014. A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa 38. ISSN 0311-5518.

A new genus and new species of Saucrosmylidae (Insecta, Neuroptera) are described (Daohugosmylus castus) based on a well-preserved hindwing from the Middle Jurassic of Daohugou, Inner Mongolia, China. Daohugosmylus gen. nov. is distinguished by a large and nearly semi-circular hindwing, relatively wide R1 space possessing several rows of cells, anteriorly bent Rs, dense crossveins over the entire wing, and smooth outer margin.

Qing Liu (corresponding author) [qingliu1029@gmail.com], Haichun Zhang [hczhang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], Yan Fang [yanfang@nigpas.ac.cn], Daran Zheng [dranzheng@gmail.com], Qi Zhang [zhqi1105@126.com] and Edmund A Jarzembowski [jarzembowski2@live.co.uk], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China; secondary address of Daran Zheng & Qi Zhang, University of Chinese Academy of Sciences, Beijing, 100049, PR China; and Ed Jarzembowski, Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. Received 13.11.2013; revised 20.1.2014; accepted 21.1.2014.     

Context, curation, and bias: an evaluation of the Middle Paleolithic collections of Combe-Grenal (France)

The site of Combe-Grenal is arguably the reference site for the Mousterian of southwest France. Bordes excavated the site over a period of 13 years and generated a large collection of lithics and fauna from a deep series of 64 Lower and Middle Paleolithic levels. Though Bordes only partially published his work at the site, its long stratigraphic sequence and particularly its collections have been subjected to numerous studies over the years and have figured prominently in several long-running debates concerning the Mousterian. Recently, the authors undertook a project to inventory and analyzed these collections along with the documentation of their archaeological context. In so doing, however, a number of serious problems became apparent with the collection in its current state, problems that cannot easily be resolved given the state of the available documentation. These problems have important implications not only for results of studies based on these collections, but more generally, they also point to problems in terms of how archaeological collections are curated.     

A new record of an aristonectine elasmosaurid (Sauropterygia,Plesiosauria) from the Upper Cretaceous of New Zealand: implications for the Mauisaurus haasti Hector, 1874 hypodigm

O’Gorman, J.P., Otero, R.A. & Hiller, N., 2014. A new record of an aristonectine elasmosaurid (Sauropterygia, Plesiosauria) from the Upper Cretaceous of New Zealand: implications for the Mauisaurus haasti Hector, 1874 Hector, J., 1874. On the fossil Reptilia of New Zealand. Transactions and Proceedings of the New Zealand Institute 6, 333–358. [Google Scholar] hypodigm. Alcheringa 38, 504–512. ISSN 0311-5518

An indeterminate aristonectine elasmosaurid is recorded from a lower Maastrichtian bed of the Conway Formation, Waipara River, South Island, New Zealand. The described specimen (CM Zfr 104), previously considered part of the hypodigm of Mauisaurus haasti, came from the upper part of the Alterbidinium acutulum biozone, the same zone from which the only well-known aristonectine from New Zealand, Kaiwhekea katiki, is recorded. The cervical vertebrae of CM Zfr 104 have the same distinctive features (i.e., with extremely broad rather than long centra) as those from previously recorded juvenile aristonectines from Argentina, Chile and Antarctica. This new record is congruent with the biogeographic relationships of Cretaceous marine amniotes from the Weddellian Palaeobiogeographic Province (i.e., Patagonia, western Antarctica, New Zealand and southeastern Australia). Therefore, this type of vertebra is regarded as a distinctive feature of the Weddellian aristonectine elasmosaurids.

José P. O’Gorman [joseogorman@fcnym.unlp.edu.ar], División Paleontología Vertebrados, Museo de La Plata, Universidad Nacional de La Plata, Paseo del Bosque s/n., B1900FWA, La Plata, Argentina; [CONICET (Consejo Nacional de Investigaciones Científicas y Técnicas)]; Rodrigo A. Otero [paracrioceras@gmail.com], Red Paleontológica U-Chile. Laboratorio de Ontogenia y Filogenia, Departamento de Biología, Facultad de Ciencias, Universidad de Chile, Av. Las Palmeras 3425, Santiago, Chile; Norton Hiller, [norton.hiller@canterbury.ac.nz], Department of Geological Sciences, University of Canterbury, PB 4800, Christchurch, 8001, New Zealand and Canterbury Museum, Rolleston Avenue, Christchurch, New Zealand, 8013.     

Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia,China

SHI, C.F., WANG, Y.J., YANG, Q. & REN, D., September 2012. Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China. Alcheringa 36, 311–320. ISSN 0311-5518.

A new grammolingiid genus, Chorilingia containing four new species (C. euryptera, C. parvica, C. translucida and C. peregrina) is described and illustrated from the Jiulongshan Formation at Daohugou, Inner Mongolia, China. The new genus is differentiated mainly on the first branch of vein Rs (Rs1) separating distal to the forks of both CuA and CuP. A key to species of the genus is provided.     

A palaeodietary investigation of carbon (C/C) and nitrogen (N/N) in human and faunal bones from the Copper Age cemeteries of Varna I and Durankulak,Bulgaria

Stable isotope analyses have been applied to human and faunal bone collagen from the Varna I and Durankulak cemeteries to explore palaeodietary adaptations in the Neolithic and Eneolithic (Copper Age). The results suggest both populations primarily utilised terrestrial, C₃-based diets, despite their proximity to the Black Sea. The wider δ¹⁵N range of the Durankulak humans likely indicates the differential utilisation of terrestrial meat sources, which is probably related to the degree to which primary and/or secondary ovicaprid products were consumed, particularly since ovicaprid δ¹⁵N values differ from other herbivores. The isotopic distribution of Varna I reflects a linear relationship between δ¹⁵N and δ¹³C, suggesting that a minority of individuals enriched in both isotopic parameters supplemented their diets with marine resources. These burials include the well known ‘chieftain’ (burial 43) and show notable material wealth by way of grave goods. At the population level, however, there is no significant correlation between stable isotope values and material wealth at Varna I, a fact with implications for theories regarding emergent social/economic hierarchies in Balkan prehistory. Five burials at Durankulak were found to have relatively enriched δ¹³C and δ¹⁵N values with respect to the rest of the population. These burials reflect a prominently marine-based or mixed terrestrial C₃-based diet that included C₄ inputs, possibly from millet, for which the limitations of stable isotope analysis on bulk collagen are not able to differentiate. AMS dating has shown that these burials belong to a much later period.     

The Supply of Glass at Portus Ilicitanus (Alicante,Spain): A Meta‐Analysis of HIMT Glasses

Portus Ilicitanus (Picola, Alicante) was the main sea harbour of the Roman Colonia Iulia Ilici Augusta and as such played a crucial role in the supply of fundamental commodities to the Iberian Peninsula. Excavations yielded large quantities of glass in fourth‐ and early fifth‐century contexts. Elemental analysis of 60 samples by laser ablation – inductively coupled plasma – mass spectrometry (LA–ICP–MS) confirmed that the glasses were imported from the Eastern Mediterranean. A majority of the glasses correspond to the HIMTa primary production group, which originates from Egypt. The statistical evaluation of published data of 589 HIMT glasses further revealed differential distribution patterns of the HIMTa and HIMTb subtypes between the Eastern and Western Mediterranean, suggesting chronological trends that are linked to wider geopolitical changes. This demonstrates the need for systematic large‐scale approaches to identify supply patterns and possible factors underlying geographical differences and/or chronological developments.     

Shell-gathering from mangroves and the seasonality of the Southeast Asian Monsoon using high-resolution stable isotopic analysis of the tropical estuarine bivalve (Geloina erosa) from the Great Cave of Niah, Sarawak: methods and reconnaissance of molluscs of early Holocene and modern times

This paper describes a reconnaissance, high-resolution, study of stable isotopes (δ¹⁸O and δ¹³C) from modern shells of the estuarine bivalve Geloina erosa, and those dated to the early Holocene that were harvested by people from mangroves near the Great Cave of Niah on the north coast of Borneo. This reconnaissance study provides high-resolution palaeoclimatic-palaeohydrological information concerning early human activity in the region and the past character of the Southeast Asian Monsoon. Laser ablation continuous flow isotope ratio mass spectrometry (LA-CF-IRMS) on modern shells of Geloina erosa revealed ‘saw-tooth’ stable isotopic profiles that bear a close resemblance to peaks and troughs of trends in recent local rainfall, including the 1998 El Niño drought, highlighting the potential of Geloina erosa for reconstructing seasonality of the Southeast Asian Palaeomonsoon. LA-CF-IRMS analysis of prehistoric shells of Geloina erosa held in the Harrisson Archives of the Sarawak Museum revealed cyclical shifts in δ¹⁸O with similar amplitudes of variation as found in modern shells of Geloina erosa. As at the present day, this probably reflects the changing seasonality of the monsoon rains with shifts to negative δ¹⁸O values during periods of high runoff. Lighter mean δ¹⁸O values of the prehistoric shells, however, may indicate a greater annual surplus of rainfall and possibly consistent with the early Holocene strengthening of the summer monsoon at that time. The similarity of the last growth increment δ¹⁸O of the prehistoric shells to their mean δ¹⁸O profile values suggests that gathering took place during times of moderate runoff.