Avian Remains from Khonkho Wankane,an Urban and Ceremonial Center in the Southern Lake Titicaca Altiplano

The urban and ceremonial center Khonkho Wankane flourished in the southern Lake Titicaca Altiplano region during the Late Formative prior to the rise of Tiwanaku as a dominant urban center. This site yielded an avifaunal number of identified specimens (NISP) = 631, with a minimum of 26 taxa represented among 539 skeletal elements or fragments. A total of 92 egg shell fragments also were recovered. The avifauna inhabited two major ecological zones: Lake Titicaca, its margins/wetlands and other water margins; and the dry Altiplano grassland (or puna). Lake/wetland taxa include Phoenicopterus chilensis (Chilean flamingo), Nycticorax nycticorax (night heron) and multiple taxa in the Order Anseriformes (ducks, geese, etc.) including Chloephaga melanoptera (Andean goose), Anas georgica (yellow‐billed pintail), A. flavirostris (speckled teal) and Oxyura jamaicensis (Andean ruddyduck). Also present are Fulica ardesiaca (Andean coot), Gallinago andina (puna snipe), Himantopus mexicanus (black‐necked stilt) and Charadrius alticola (puna plover). Puna taxa include Nothoprocta cf. ornata (ornate tinamou), Nothura cf. darwinii (Darwin's nothura), Metropelia sp. (ground doves), Athene cunicularia (burrowing owl), and multiple Passeriformes (songbirds, etc.). Taxa with a cosmopolitan distribution include Falco cf. femoralis (aplomado falcon) and Tyto alba (barn owl). Taphonomic analysis indicates that the avifauna were used in ceremonial contexts, including multiple Falco cf. femoralis burials, and for dietary and possibly tool‐making purposes. Other sources of introduction into the site deposits likely include natural mortality of taxa directly inhabiting the site area. These findings are compared to the avifauna recovered from the Formative site of Chiripa along the Lake Titicaca margin, which included a much higher proportion of lake bird fauna. At Khonkho Wankane, the importance of wetland avifauna may have been enhanced by the alteration of the local environment to include qochas, or artificial reservoirs/ponds, and continued despite reliance upon multiple domesticated plant and mammal species. Copyright © 2011 John Wiley & Sons, Ltd.     

Incipient urbanism at the Early Andean center of Khonkho Wankane,Bolivia

Abstract

For decades archaeologists have recognized that Khonkho Wankane was an important monumental site in the Bolivian Andes, and most have interpreted it as a Tiwanaku regional center. A decade of research indicates that Khonkho Wankane was an important Late Formative center that engendered Tiwanaku’s Middle Horizon urbanism and ritual-political centrality. It was neither a Tiwanaku regional center nor Tiwanaku’s second city. Furthermore, recent research at Khonkho suggests a critique of the “one center, one polity” model of political complexity. Rather than being the singular center of a spatially bounded multi-community polity, Khonkho was one of multiple interacting Late Formative centers in an emergent macro-community that gave rise to Tiwanaku urban centrality.     

Hunting,Herding, and Caravanning: Osteometric Identifications of Camelid Morphotypes at Khonkho Wankane,Bolivia

Khonkho Wankane is a ceremonial center located in the southern Lake Titicaca Basin, Bolivia. During the Late Formative period (AD 1–500), its residents practiced agropastoral lifeways and participated in the rise of the state at Tiwanaku. Like at many Andean sites, bones from the family Camelidae are the most abundant large mammal in domestic contexts. Identifying camelid morphotypes represented by these bones carries far‐reaching implications for understanding past hunting, herding, and caravanning practices, and their roles in larger social and economic webs. Identifications were based on a locally focused reference collection, including llamas (Lama glama) from the immediate vicinity of the site, as well as Andean guanacos (Lama guanicoe), a much smaller morphotype than the Patagonian guanacos used in many osteometric studies. Multivariate statistical analyses and incisor morphology identified all four camelid. Different analyses suggest that the crux of osteometry lies in the reference collection, not the statistical test. An additional, very large morphotype likely corresponds to a castrated llama, the preferred cargo animal among modern drovers. The presence of these animals is interpreted as evidence that groups hunted vicuña (Vicugna vicugna) and guanaco, which are not currently present around the site, herded llamas and alpacas (Vicugna pacos), and perhaps organized caravans with castrated llamas. Copyright © 2013 John Wiley & Sons, Ltd.     

Mammalian Microfaunal Remains from Khonkho Wankane (Late Formative Period), Mollo Kontu (Middle Horizon Period) and Pukara de Khonkho (Late Intermediate Period) in the Bolivian Altiplano

Paleoecological and taphonomic analysis of the mammalian microfauna from three Bolivian altiplano sites is presented: Khonkho Wankane [Late Formative period; number of identified specimens (NISP) = 1019], Mollo Kontu (Middle Horizon period; NISP = 610) and Pukara de Khonkho (Late Intermediate period; NISP = 165). The following species, all of which are present in this region today, were identified: Akodon boliviensis (Bolivian grass mouse), Andinomys edax (Andean mouse), Auliscomys sublimis (Andean leaf‐eared mouse), Cavia porcellus (guinea pig/cuy), Chinchilla chinchilla (chinchilla), Ctenomys leucodon (white‐toothed tuco‐tuco), Eligmodontia puerulus (Andean gerbil mouse), Galea musteloides (common yellow‐toothed cavy), Lagidium viscacia (vizcacha), Neotomys ebriosus (Andean swamp rat),and Oligoryzomys destructor (destructive pygmy rice rat). Some direct human involvement with the deposition of some of the larger species (Cavia porcellus, Chinchilla chinchilla, Ctenomys leucodon and Lagidium viscacia) was noted, although C. porcellus was identified only from a single element from Mollo Kontu. Most deposition was likely due to natural in‐burrow deaths and other natural attrition of these species. No significant shift in environment during the occupation of these sites was detected. Copyright © 2012 John Wiley & Sons, Ltd.     

Isotopic evidence for Middle Horizon to 16th century camelid herding in the Osmore Valley,Peru

Archaeological and zooarchaeological data indicate that camelid pastoralism was a subsistence and economic mainstay of Middle Horizon and more recent cultures in the Osmore region of southern Peru. However, it is not known whether camelids were primarily herded in highland puna pastures or near lower elevation sites in the middle valley or along the coast. This research examines the elevation of archaeological camelid herding in the Osmore Valley using stable isotope analysis. Stable carbon (δ¹³C) and nitrogen (δ¹⁵N) isotope ratios were measured on 28 archaeological camelid bone samples from the Middle Horizon sites of Cerro Baúl and Cerro Mejía, the Late Intermediate site of Yaral and the late pre‐Hispanic to Colonial period site of Torata Alta. Twenty‐three archaeological camelids have δ¹³C and δ¹⁵N values similar to five modern camelids maintained in highland puna pastures. In contrast, three camelids from the high status Wari site of Cerro Baúl, and two from Yaral have unexpectedly high δ¹³C and/or δ¹⁵N values outside the expected range for camelids pastured in highland puna habitats. The results may be explained by differences in foddering practices, altitudinal herding range or climatic conditions. Strontium isotope ratios (⁸⁷Sr/⁸⁶Sr) were also assessed to infer pasture elevation using the camelid remains from Cerro Baúl (n = 11). One individual in this sub‐sample exhibits a non‐local ⁸⁷Sr/⁸⁶Sr value indicative of an origin in the highland puna region east or south of Lake Titicaca. It was not possible to further distinguish between camelids herded in lower to middle elevation habitats outside the Lake Titicaca basin using ⁸⁷Sr/⁸⁶Sr values. This study suggests that multiple isotope proxies may be used to identify animals primarily pastured in lowland coastal versus highland puna (> 4000 masl (meters above sea level)) habitats, but are less useful at distinguishing between animals pastured in lower to middle elevation settings. Copyright © 2010 John Wiley & Sons, Ltd.     

Provisioning Inka feasts at Tiwanaku, Bolivia: the geographic origins of camelids in the Pumapunku complex

While political integration can be achieved by many means, here we focus on the use of feasting and statecraft in the Inka Empire of the Andean Late Horizon (c. AD 1400-1532) in South America. In order to examine Inka political integration of the Lake Titicaca Basin of Bolivia, we examine paleomobility and paleodiet through radiogenic strontium and stable oxygen and carbon isotope data in archaeological camelid remains from the site of Tiwanaku. Mean radiogenic strontium isotope values from all archaeological camelid enamel and bone samples is ⁸⁷Sr/⁸⁶Sr = 0.70998 ± 0.00179 (1σ, n = 48), mean stable oxygen isotope values from a sub-set of archaeological camelid enamel and bone samples is δ¹⁸O_{carbonate (VPDB)} = −10.0‰ ± 2.6‰ (1σ, n = 18) and mean stable carbon isotope values from a sub-set of archaeological camelid enamel and bone samples is δ¹³C_{carbonate (VPDB)} = −9.0‰ ± 1.7‰ (1σ, n = 18). While many camelids consumed in these feasting events were likely local to the Lake Titicaca Basin, others came from a variety of different geologic zones, elucidating our understanding of Inka statecraft and the role of feasting in political integration in empires in the past.     

A new species of triplinerved Laurophyllum from the Eocene of Nerriga,New South Wales

A new fossil triplinerved dicotyledon leaf, Laurophyllum acrocryptocaryoides Conran & Christophel (Lauraceae) from Eocene deposits at Nerriga is described. The fossil is similar to the previously described L. acrodromum Hill, also from this site, but differs in leaf shape and cuticular features. L. acrocrytocaryoides also resembles some members of the extant genus Cryptocarya and its relationship to extant taxa is discussed.     

Acanthodian,onychodontid and osteolepidid fish from the middle-upper Taemas Limestone (Early Devonian), Lake Burrinjuck,New South Wales

A meagre ischnacanthid acanthodian fauna, known from fragmentary dentigerous jawbones from Lake Burrinjuck, Yass district, New South Wales, consists of three taxa: Taemasacanthus narrengullenensis, Taemasacanthus cooradigbeensis and an indeterminate ischnacanthid. The fossils are from three Early Devonian (Emsian) units at Taemas: the Bloomfield Limestone Member, the Currajong Limestone Member and the Warroo Limestone Member, which constitute the middle-upper parts of the Taemas Limestone (Murrumbidgee Group); and from Unit 6 of the ‘Upper Reef formation’ at Wee Jasper. Whereas ischnacanthid acanthodian remains are increasingly rarer in the higher levels of the Taemas Limestone, remains of onychodontid and osteolepidid fishes are relatively abundant; Onychodus yassensis sp. nov., Onychodus sp. and isolated osteolepidid teeth are locally common.     

The Location of Lake Titicaca’s Coastal Area During the Tiwanaku and Inca Periods: Methodology and Strategies of Underwater Archaeology

For more than 30 years, numerous research projects have revealed the dense and complex human settlement of the lacustrine basin of Lake Titicaca in Bolivia and Peru. Physical evidence of such establishments has been discovered in plains, valleys, and highlands connected to the lake. These remains confirm human occupation and development in this environment, particularly during the Tiwanaku (AD 500-1150) and Inca (AD 1400-1532) Periods. The research project discussed in this paper includes consideration of submerged areas through underwater archaeology. This investigation involves analysis of two areas that have evidence of ancient human occupation but are poorly documented: the coastal and lacustrine regions. Due to its dominance in the landscape, Lake Titicaca has always been a major feature in the life and identity of populations of this vicinity. These inhabitants have developed socio-economic and ritual behaviours directly associated with the lake that have left cultural and material prints that are the foci of the present study.     

Fossil fruit identification by SEM investigations, a tool for palaeoenvironmental reconstruction of Dmanisi site, Georgia

Archaeological excavations and multidisciplinary studies at the site of Dmanisi (Lesser Caucasus, Georgia) have provided an assemblage of lithic artefacts and numerous palaeontological and human remains in a volcano-sedimentary context dating to the early Lower Pleistocene. The sieving of archaeological sediments from Dmanisi also yielded mineralised botanical macro-remains that were analysed according to a particular protocol. The use of scanning electron microscopy (SEM) allowed the comparison between fossil fruit remains and modern specimens, leading to a precise identification of the earlier. Seed remains recognised by this method belonging to several taxa from the Boraginaceae family as well as to the genus Celtis L., of the Ulmaceae family. The ecological requirements of these taxa suggest the presence of xeric conditions, a result that is in accordance with data obtained by faunal and stratigraphic studies.     

The fish of Lake Titicaca: implications for archaeology and changing ecology through stable isotope analysis

Research on past human diets in the southern Lake Titicaca Basin has directed us to investigate the carbon and nitrogen stable isotopes of an important dietary element, fish. By completing a range of analyses on modern and archaeological fish remains, we contribute to two related issues regarding the application of stable isotope analysis of archaeological fish remains and in turn their place within human diet. The first issue is the potential carbon and nitrogen isotope values of prehistoric fish (and how these would impact human dietary isotopic data), and the second is the observed changes in the fish isotopes through time. Out of this work we provide quantitative isotope relationships between fish tissues with and without lipid extraction, and a qualitative analysis of the isotopic relationships between fish tissues, allowing archaeologists to understand these relationships and how these values can be applied in future research. We test a mathematical lipid normalization equation to examine whether future researchers will need to perform lipid extraction procedures for Lake Titicaca fish. We also analyze a number of aquatic plants to better understand the range of isotopic signatures of the Lake Titicaca ecosystem. We use these data to better understand prehistoric human diet and the role that fish may have played in the past as well as potential changes in local lake ecology through time.     

Discovery of late Changhsingian (latest Permian) brachiopod Attenuatella species from South China

Cretaceous polar dinosaur faunas were taxonomically diverse, which suggests varied strategies for coping with the climatic stress of high latitudes. Some polar dinosaurs, particularly larger taxa such as the duckbill Edmontosaurus Lambe, 1917 Lambe, L. M. 1917. A new genus and species of crestless hadrosaur from the Edmonton Formation of Alberta. Ottawa Naturalist, 31: 65–73.  [Google Scholar], were biomechanically and energetically capable of migrating over long distances, up to 2600 km. However, current evidence strongly suggests many polar dinosaurs (including sauropods, large and small theropods, and ankylosaurs of New Zealand) overwintered in preference to migration. Certain groups also appear more predisposed to overwintering based on their physical inability (related to biomechanics, natural history, or absolute size) to migrate, such as ankylosaurs and many small taxa, including hypsilophodontids and troodontids. Low-nutrient subsistence is found to be the best overwintering method overall, although the likelihood that other taxa employed alternative means remains plausible. Despite wide distribution of some genera, species-level identification is required to assess the applicability of such distributions to migration distances. Presently, such resolution is not available or contradicts the migration hypothesis.     

The record of Araucariaceae macrofossils in New Zealand

Pole, M., December, 2008. The record of Araucariaceae macrofossils in New Zealand. Alcheringa 32, 405–426. ISSN 0311-5518.

The Araucariaceae have a long record in New Zealand, extending back to the Jurassic at least, and Araucaria extends back to at least the Late Cretaceous. This paper reviews the macrofossil record of the family and presents new information based largely on the leaf cuticle record. Agathis, which is the only genus of the family currently growing in New Zealand, has no record before the Cenozoic. All specimens previously identified from pre-Cenozoic strata clearly belong to other taxa or do not show characteristic features of the genus. Araucariaceae macrofossils are virtually ubiquitous in the Cretaceous assemblages of New Zealand but are conspicuous by their absence or rarity in Palaeocene deposits. Their demise may be an expression of events at the Cretaceous–Palaeogene boundary.     

Permian bivalve molluscs from the Gai-As Formation,northern Namibia: systematics,taphonomy and biostratigraphy

Fossil bivalves from two horizons in the Gai-As Formation of NW Namibia are tentatively correlated with mid-Permian taxa of the Passa Dois Group of Brazil, supporting the concept that the Paraná Basin extended into Africa. The Namibian fauna includes a new genus and species, Huabiella compressa, which was previously confused with Brazilian taxa. The taphonomy of the bivalve-rich strata indicates deposition under the influence of episodic events, such as storms. The Gai-As Formation directly overlies the mesosaurid-bearing deposits of the Huab Formation, indicating a significant unconformity when compared with the more complete succession of the Passa Dois Group, Paraná Basin, Brazil. The studied bivalve assemblages are no younger than 265±2.5 Ma (mid-Permian), based on U/Pb radiometric dating of zircons from tuffs.     

Nothofagus macrofossils from the Tertiary of Tasmania

Leaves assignable to Nothofagus from two fossil deposits in Tasmania represent the first macrofossils of this genus from the Tertiary in southeastern Australia. One fossil species, N. johnstonii, is closely related to the extant Australian species N. cunninghamii while the other fossil species, N. tasmanica, has very close affinites with the extant Australian species N. moorei. All four of these species are closely interrelated. The pollen type produced by the fossil species is unknown, since all three types are present in the microfloras. However, both N. cunninghamii and N. moorei produce N. menziesii-type pollen. The macrofossils confirm the conclusion from pollen studies that evolution in Nothofagus has occurred very slowly.     

Phylogenetic implications of macropodid (Marsupialia: Macropodoidea) posteranial remains from Miocene deposits of Riversleigh,northwestern Queensland

A new sthenurine taxon, Rhizosthenurus flanneryi gen. et sp. nov. from middle late Miocene deposits of Riversleigh, northwestern Queensland, is established on the basis of material previously assigned to Bulungamayinae. Postcranial remains belonging to an indeterminate basal macropodid are described. Cladistic analysis of 50 discrete postcranial characters coded for 16 ingroup and two outgroup taxa suggests that R. flanneryi is the most plesiomorphic member of a clade containing the late Miocene macropodid Hadronomas puckridgi and crown-group sthenurines. The indeterminate basal macropodid is placed as the immediate sister taxon to the bulungamayine Ganguroo bilamina and a monophyletic clade containing macropodines and sthenurines. This arrangement supports consideration of bulungamayines as ancestral to other macropodids and suggests that macropodines and sthenurines are monophyletic. However, the possibility of a diphyletic origin for Macropodinae and Sthenurinae from within Bulungamayinae cannot be dismissed.     

Tertiary frogs from South Australia

Four species of hylid and leptodactylid frogs are reported from the Tertiary (mid-Miocene) Ngapakaldi fauna of the Etadunna Formation at Lake Palankarinna, South Australia. The species comprise Limnodynastes archeri sp. nov., Litoria sp. cf. caerulea (White), Litoria sp. indet. and Australobatrachus ilius Tyler. Previously the extant genera Limnodynastes and Litoria have been known only from Quaternary deposits.     

Tertiary megafossil flora of Maslin Bay,South Australia: a preliminary report

The Maslin Bay flora of South Australia is of lower Middle Eocene age and contains diverse, well preserved angiosperm assemblages. It has yielded 2700 specimens belonging to approximately 200 leaf taxa.

Physiognomic analysis (sensu Webb, 1959; Wolfe 1970) indicates greatest similarity to extant Simple Mesophyll Vine Forest and Complex Notophyll Vine Forest found currently in Queensland.

Preliminary taxonomic studies have confirmed the presence of taxa closely resembling Podocarpus (Podocarpaceae), Agathis (Araucariaceae), Fatsia (Araliaceae) and Banksia (Proteaceae). Microfloral analysis of the deposit confirms the presence of the latter three families, but suggests far greater occurrence of Proteaceae than the number of leaf specimens of this family indicates. Nothofagus pollen is the dominant type, although leaves of this genus are absent from collections. Based on sedimentation, physiognomy and systematic studies of leaves and fungi (Lange, 1969) it is concluded that the Maslin Bay region supported tropical to subtropical rain forest during the lower Middle Eocene.     

An investigation of the frequency of squatting facets in Later Stone Age foragers from South Africa

Distinctive morphological changes attributed to a habitual squatting posture were observed on the distal femur, distal tibia and on the talar neck of Later Stone Age (LSA) foragers. The frequency of squatting in LSA foragers (n = 56) was determined by the presence of anatomical features of joint hyperflexion. Three South African comparative skeletal groups from different time periods were also analysed: skeletal remains of early farming populations (n = 17), 18th century ‘Free Blacks’ and/or slaves from Cobern Street, Cape Town (n = 21), and a modern cadaver sample (n = 29). The results show that 28 out of 56 LSA foragers (50%) were habitual squatters; 13 out of 17 farmers (76.5%) and one of 21 individuals from the Cobern Street collection (4.8%) demonstrated squatting facets. No anatomical squatters were found in the modern cadaver sample. There was no significant sex difference between squatters and non‐squatters. Hence at least half of the LSA foragers and farmers were habitual squatters, according to the signs of joint hyperflexion. Squatting is a comfortable position for those used to it because the body weight is supported with minimal muscular activity. Two probable reasons are suggested for the difference in postures adopted by the different groups: (1) availability or lack of availability of furniture and (2) cultural and individual differences in resting posture. Copyright © 2005 John Wiley & Sons, Ltd.