Revised conodont and fusuline biostratigraphy of the Bamchi Formation (Pyongan Supergroup) at the Bamchi section,Yeongwol and the Carboniferous–Permian boundary in South Korea

Wang, Q., Wang, Y., Qi, Y., Wang, X., Choh, S.J., Lee, D.C. & Lee, D.J., November 2017. Yeongwol and the Carboniferous–Permian boundary in South Korea. Alcheringa 42, 245–258. ISSN 0311-5518

Six conodont and one fusuline zones are recognized on basis of a total of 25 conodont and 13 fusuline species (including seven unidentified species or species given with cf. or aff. in total) from the Bamchi Formation, Yeongwol, Korea. The conodont zones include the Streptognathodus bellus, S. isolatus, S. cristellaris, S. sigmoidalis, S. fusus and S. barskovi zones in ascending order, which can be correlated with the conodont zones spanning the uppermost Gzhelian to Asselian Age of the Permian globally. The fusuline zone is named the Rugosofusulina complicata–Pseudoschwagerina paraborealis zone. The co-occurrence of the conodont Streptognathodus isolatus (the Global Boundary Stratotype Section and Point index for the base of Permian) and Pseudoschwagerina (a Permian inflated fusuline) indicates that the Carboniferous–Permian boundary can be placed in the lower part of the Bamchi Formation in South Korea.

Qiulai Wang* [qlwang@nigpas.ac.cn] CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, East Beijing Road 39, Nanjing 210008, PR China; Yue Wang* [yuewang@nigpas.ac.cn] LPS, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, East Beijing Road 39, Nanjing 210008, PR China; Yuping Qi* [ypqi@nigpas.ac.cn] Xiangdong Wang* [xdwang@nigpas.ac.cn] CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, East Beijing Road 39, Nanjing 210008, PR China; Suk-Joo Choh [sjchoh@korea.ac.kr] Department of Earth and Environmental Sciences, Korea University, Seoul 02841, Republic of Korea; Dong-Chan Lee [dclee@chungbuk.ac.kr] Department of Earth Sciences Education, Chungbuk National University, Cheongju 28644, Republic of Korea; Dong-Jin Lee [djlee@andong.ac.kr] Department of Earth and Environmental Sciences, Andong National University, Andong 36729, Republic of Korea. *Also affiliated with: University of Chinese Academy of Sciences, 19A Yuquan Road, Shijingshan District, Beijing 100049, PR China.     

Palaeobiological features of the coralomorph Amsassia from the Late Ordovician of South China

Lee, M., Elias, R.J., Choh, S.-J. & Lee, D.-J., May 2018. Palaeobiological features of the coralomorph Amsassia from the Late Ordovician of South China. Alcheringa XXX, X–X. ISSN 0311-5518.

Amsassia yushanensis sp. nov. occurs in the Late Ordovician Xiazhen Formation at Zhuzhai, Jiangxi Province of southeastern China. This species is characterized by typical phacelocerioid organization of modules comparable with the other Amsassia species described in recent literature. Bipartite fission, in which a parent module divided into two parts, is by far the most common type of increase in this species; tripartite and quadripartite types of axial fission do occur but are relatively uncommon. Processes of module division are similar to those of A. shaanxiensis and A. koreanensis, and also occurred in tetradiids. In A. yushanensis, restoration of coralla was occasionally accompanied by recovery of a damaged or injured module or group of modules probably following an influx of sediment, as observed in some favositoid corals. Amsassia superficially resembles Lichenaria, a representative genus of the most primitive stock of tabulate corals of Ordovician age, and has likely been mistakenly identified as Lichenaria in the North China Platform. Available information suggests that the validity of a reported occurrence of Lichenaria in the South China Platform is also questionable.

Mirinae Lee [mirinae.lee@kopri.re.kr] Division of Polar Earth-System Sciences, Korea Polar Research Institute, 21990, Incheon, Republic of Korea; Robert J. Elias [Robert.Elias@umanitoba.ca] Department of Geological Sciences, University of Manitoba, Winnipeg, R3T 2N2, Canada; Suk-Joo Choh [sjchoh@korea.ac.kr] Department of Earth and Environmental Sciences, Korea University, Seoul, 02841, Republic of Korea; Dong-Jin Lee [djlee@anu.ac.kr] Department of Earth and Environmental Sciences, Andong National University, Andong, 36749, Republic of Korea and College of Earth Sciences, Jilin University, Changchun 130061, PR China.     

Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand)

Conran, J.G., Bannister, J.M. & Lee, D.E., 2013. Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand). Alcheringa 37, 496–509. ISSN 0311-5518.

Laurelia otagoensis sp. nov. Conran, Bannister & D.E. Lee (Laurales: Atherospermataceae) is described from the earliest Miocene Foulden Maar diatomite deposit, Otago, New Zealand. The new species is represented by mummified fossil leaves with well-preserved cuticle and associated clusters of achenes bearing persistent, long plumose styles. This basal angiosperm family is of significance because of its classic southern disjunctions and ecological importance in extant Gondwana-type rainforests, but has a very sparse fossil record. The present study describes one of very few convincing leaf fossils for Atherospermataceae and the only definitive fossil fruits. The presence of fossil Laurelia in Oligo–Miocene New Zealand combined with fossil leaf impressions from the late Eocene, Miocene dispersed cuticle and pollen from the Oligocene to Holocene shows that the family has had a long history in Cenozoic New Zealand. These new fossils also support palaeoclimatic data suggesting warmer conditions in the earliest Miocene of New Zealand.

John G. Conran [john.conran@adelaide.edu.au], Australian Centre for Evolutionary Biology & Biodiversity and Sprigg Geobiology Centre, School of Earth and Environmental Sciences, Benham Bldg DX 650 312, The University of Adelaide, SA 5005 Australia; Jennifer M. Bannister [jennifer.bannister@xtra.co.nz], Department of Botany, University of Otago, PO Box 56, Dunedin, New Zealand; Daphne E. Lee [daphne.lee@otago.ac.nz], Department of Geology, University of Otago, PO Box 56, Dunedin, New Zealand. Received 17.12.2012; revised 9.4.2013; accepted 15.4.2013.     

Paleocene flora from Seymour Island,Antarctica: revision of Dusén’s (1908) angiosperm taxa

The Paleocene flora from Seymour Island, Antarctica, is one of the most diverse floras of this age in the Southern Hemisphere. First collected on the Swedish South Polar Expedition (1901–1903), it was described by Dusén in 1908 as having 87 leaf taxa. Forty-seven angiosperm taxa were described and/or illustrated. Many species are based on single specimens, and the flora has not been re-examined in its entirety since it was first described. This study is the first reassessment of the flora updating the original research using current methodologies, and permitting evaluation of the flora in the context of modern ideas on plant evolution and palaeogeography. This paper continues the revision of the material first studied by Dusén; a previous paper described the ferns and gymnosperms; here we describe the angiosperms. The revision is based on the original collections held at the Swedish Museum of Natural History, together with the first major new collections held at the British Antarctic Survey. Among the taxa recognized by Dusén, we recognize only three entire-margined and 11 tooth-margined angiosperms. This revision to 14 species notably lacks the two tropical elements originally described from the flora, Mollinedia seymourensis and Miconiiphyllum austral. Hence, its status as a ‘Mixed Flora’ comes into question and influences climatic interpretations based on Gondwanan floras. Anne-Marie Tosolini^? [a.tosolini@unimelb.edu.au], Department of Earth Sciences, University of Leeds, Leeds LS2 9JT, UK, and Environmental Geoscience, Latrobe University, Victoria, 3086, Australia; David Cantrill [david.cantrill@rbg.vic.gov.au], National Herbarium of Victoria, Royal Botanic Gardens Melbourne, Private Bag 2000, Birdwood Ave, South Yarra, Victoria, 3141, Australia; Jane Francis [j.francis@earth.leeds.ac.uk], Department of Earth Sciences, University of Leeds, Leeds LS2 9JT, UK. ?Present address: School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia. Received 5.5.2012; revised 13.12.2012; accepted 7.1.2012.     

Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia

Benson, R.B.J., Fitzgerald, E.M.G., Rich, T.H. & Vickers-Rich, P., 2013. Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia. Alcheringa 37, 1–6. ISSN 0311-5518

We report a large plesiosaurian tooth from the freshwater early–middle Aptian (Early Cretaceous) Eumeralla Formation of Victoria, Australia. This, combined with records of smaller plesiosaurian teeth with an alternative morphology, provides evidence for a multitaxic freshwater plesiosaurian assemblage. Dental and body size differences suggest ecological partitioning of sympatric freshwater plesiosaurians analogous to that in modern freshwater odontocete cetaceans. The evolutionarily plastic body plan of Plesiosauria may have facilitated niche differentiation and helped them to exclude ichthyosaurs from freshwater environments during the Mesozoic. However, confirmation of this hypothesis requires the discovery of more complete remains.

Roger B.J. Benson [roger.benson@earth.ox.ac.uk], Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK; Erich M.G. Fitzgerald [efitzgerald@museum.vic.gov.au], Thomas H. Rich [trich@museum.vic.gov.au], Museum Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia; Thomas H. Rich and Patricia Vickers-Rich [pat.rich@monash.edu], School of Geosciences, Monash University, Clayton, Victoria 3800, Australia. Received 30.10.2012; revised 27.1.2013; accepted 31.1.2013.     

New damselflies (Odonata: Zygoptera: Hemiphlebiidae,Dysagrionidae) from mid-Cretaceous Burmese amber

Zheng, D., Zhang, Q., Nel, A., Jarzembowski, E.A., Zhou, Z., Chang, S.-C. &; Wang, B., May 2016. New damselflies (Odonata: Zygoptera: Hemiphlebiidae, Dysagrionidae) from mid-Cretaceous Burmese amber. Alcheringa XX, xxx–xxx. ISSN 0311-5518

Two damselflies, Burmahemiphlebia zhangi gen. et sp. nov. and Palaeodysagrion cretacicus gen. et sp. nov., are described from the mid-Cretaceous Burmese amber. Burmahemiphlebia zhangi is the first record of Hemiphlebiidae from this amber, although the family was cosmopolitan during the Mesozoic. It can be readily distinguished from all other members of Hemiphlebiidae in having very short MP and CuA veins, and in its rectangular discoidal cell. The new fossils support the view that hemiphlebiid damselflies were one of the dominant groups of Zygoptera during the Mesozoic. Palaeodysagrion cretacicus is the first dysagrionid damselfly from Burmese amber and the second Mesozoic representative of this predominantly Paleogene group. It differs from other members of Dysagrionidae in having a unique elongate discoidal cell. These new finds increase the diversity of damselflies in mid-Cretaceous Burmese amber.

Daran Zheng* [dranzheng@gmail.com], Su-Chin Chang [suchin@hku.hk], Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Qingqing Zhang [qqzhang@nigpas.ac.cn], Edmund A. Jarzembowski? [edj@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Zhicheng Zhou [172692308@qq.com], The PLA Information Engineering University, 62 Kexue Ave, Gaoxin District, Zhengzhou 450002, Henan, PR China. *Also affiliated with State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China. ?Also affiliated with Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. ?Also affiliated with Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China.     

A new dysagrionid damselfly (Odonata: Zygoptera: Palaeodysagrion) from mid-Cretaceous Burmese amber

AUTHOR, N. &; AUTHOR, P. December 2017. Article title. Alcheringa 42, 301-305. ISSN 0311-5518. Dysagrionidae are common in Paleogene sedimentary rocks, but rarely recorded in the Mesozoic. This family, however, is diverse in Burmese amber. A new dysagrionid damselfly, Palaeodysagrion youlini Zheng, Chang &; Chang sp. nov., is described here based on a well-preserved specimen (holotype) in Burmese amber. The new damselfly provides wing apex and body characters for Palaeodysagrion. It differs from Palaeodysagrion cretacia in having Arc slightly distal of Ax2, the midfork slightly basal of the nodus, Cr and Sn almost perpendicular to RA and RP and in having a simple wing system. This is the fourth dysagrionid damselfly described from the Burmese amber.

Daran Zheng* [dranzheng@gmail.com], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210,008, PR China; Su-Chin Chang [suchin@hku.hk]*, Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Bo Wang? [bowang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210,008, PR China. *Also affiliated with: Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China. ?Also affiliated with: Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100,101, PR China.     

Artinskian (Early Permian) fusuline fauna from the Rongma area in northern Tibet: palaeoclimatic and palaeobiogeographic implications

Zhang, Y.C., Wang, Y., Zhang, Y.J. & Yuan, D.X., 2013. Artinskian (Early Permian) fusuline fauna from the Rongma area in northern Tibet: palaeoclimatic and palaeobiogeographic implications. Alcheringa 37, 529–546. ISSN 0311–5518.

A fusuline fauna consisting of ten species in five genera from the Qudi Formation of the Jiaomuri and Gangtangcuo sections in the Rongma area, northern Tibet, is described. This fauna contains a few typical Peri-Gondwanan fusuline species such as Pseudofusulina pamirensis, Neodutkevitchia insignis and N. sp. cf. N. tumidiscula. The fauna is dated as Artinskian based on the presence of both genera Chalaroschwagerina and Praeskinnerella. The Peri-Gondwanan fusuline fauna in the Qiangtang Block is grouped palaeobiogeographically into the Southern Transitional Zone, which is located in the southern part of the Peri-Gondwanan blocks during the late Sakmarian and Artinskian. Moreover, the occurrence of the Peri-Gondwanan fusuline fauna, the lithological transition from glacimarine deposits to carbonates, and the Peri-Gondwanan fusulines’ apparent southeastward migration from the Kalmard Block of central Iran during the Artinskian, are interpreted here to be the result of global warming after the peak of the Late Palaeozoic Ice Age.

Yi-chun Zhang [geozyc@yahoo.com], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China; Yue Wang [yuewang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China; Yu-jie Zhang [zebiac@163.com], Chengdu Center, China Geological Survey, 2 Renming Road North, Chengdu 610081, PR China; Dong-xun Yuan [yuanzi55@163.com], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, China; Department of Earth Sciences, Nanjing University, 22 Hankou Road, Nanjing 210008, PR China. Received 12.3.2013; revised 2.5.2013; accepted 12.5.2013.     

A new species of the aphid family Burmitaphididae (Hemiptera: Sternorrhyncha: Aphidomorpha) from Upper Cretaceous Burmese amber

Liu, X., Qiao, G.X., Yao, Y. & Ren, D., 28 March 2019. A new species of the aphid family Burmitaphididae (Hemiptera: Sternorrhyncha: Aphidomorpha) from Upper Cretaceous Burmese amber. Alcheringa 43, 455–460. ISSN 0311-5518

A new species of extinct aphids is reported based on a fossil specimen with a relatively complete body and broken wings from Upper Cretaceous Burmese amber. Vasteantenatus reliquialaus sp. nov. (Hemiptera: Aphidomorpha: Burmitaphididae) differs from other burmitaphidids in having antennae distinctly longer than the body. The diagnosis of Burmitaphididae is emended, and a key to all species of the family is provided

Xue Liu [katharina0521@foxmail.com], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China; Gexia Qiao [qiaogx@ioz.ac.cn], Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, No. 1 Beichen West Road, Chaoyang District, Beijing, 100101, China; Yunzhi Yao* [yaoyz100@126.com], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China; Dong Ren [rendong@mail.cnu.edu.cn], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China.     

First bryozoan fauna from the middle Miocene of Central Java,Indonesia

Di Martino, E., Taylor, P.D., Fernando, A.G.S., Kase, T. & Yasuhara, M. 3 June 2019, 2019. First bryozoan fauna from the middle Miocene of Central Java, Indonesia. Alcheringa 43, 461–478. ISSN 0311-5518.

Despite the publication of several taxonomic studies during the last few years, our knowledge of bryozoans from the diversity hotspot of the Indo-West Pacific remains seriously deficient. Here we describe 11 bryozoan species, comprising two anascan- and nine ascophoran-grade cheilostomes, from the middle Miocene (Langhian–Serravallian) of Sedan in Central Java, Indonesia. Three ascophoran-grade cheilostomes, Characodoma multiavicularia sp. nov. Di Martino & Taylor, Stenosipora? cribrata sp. nov. Di Martino & Taylor and Lacrimula patriciae sp. nov. Di Martino & Taylor, are described as new species. All of the three extant species have an Indo-Pacific distribution today and two are here reported as fossil for the first time. Four species are left in open nomenclature, either because of the scarcity of available material or the absence of crucial morphological features. Two of these, placed in Cosciniopsis and Actisecos, are likely to be new, while Discoporella sp. represents the easternmost known record for this common free-living genus. A single species, Lacrimula asymmetrica Cook & Lagaaij, was already known from the early Miocene of the same region, although from a site further to the east.

Emanuela Di Martino* [e.di-martino@nhm.ac.uk], Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK; Paul D. Taylor [p.taylor@nhm.ac.uk], Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK; Allan Gil S. Fernando [agsfernando@yahoo.com], National Institute of Geological Sciences, The University of Philippines, Diliman, Quezon City, Philippines; Tomoki Kase [neritopsis@gmail.com], National Museum of Nature and Science, Tsukuba, Japan; Moriaki Yasuhara [yasuhara@hku.hk], School of Biological Sciences, Swire Institute of Marine Science, University of Hong Kong, Kadoorie Biological Sciences Building, Pokfulam Road, Hong Kong SAR, PR China     

The first damsel-dragonfly (Odonata: Isophlebioidea: Campterophlebiidae) from the Middle Jurassic of Shaanxi Province,northwestern China

ZHENG, D., DONG, C., WANG, H., YE, Y., WANG, B., CHANG S-C. & ZHANG, H., May 2017. The first damsel-dragonfly (Odonata: Isophlebioidea:Campterophlebiidae) from the Middle Jurassic of Shaanxi Province, northwestern China. Alcheringa 41, 509–513. ISSN 0311-5518.

Campterophlebiidae is the most diverse family of fossil odonatans in China with ten genera recovered mostly from Middle Jurassic strata of Inner Mongolia. We describe a well-preserved campterophlebiid damsel-dragonfly from the Middle Jurassic Yanan Formation in Shanxi Province, northwestern China. This discovery adds to the diversity of Campterophlebiidae and identifies a new Middle Jurassic insect fossil locality in China. Within Campterophlebiidae, the new taxon most closely resembles Ctenogampsophlebia from the Middle Jurassic of Inner Mongolia but differs from other genera in having vein AA with four parallel posterior branches uncrossed in the anal triangle.

Daran Zheng [dranzheng@gmail.com] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Chong Dong [cdong@nigpas.ac.cn], He Wang [hwang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Yifei Ye [459353251@qq.com] Shannxi Non-ferrous Yulin Coal Co., Ltd, Yulin, PR China; Bo Wang [bowang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China; Su-Chin Chang [suchin@hku.hk] Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China.     

On the integration of Ordovician conodont and graptolite biostratigraphy: new examples from Gansu and Inner Mongolia in China

Wang, Z.H., Bergström, S.M., Zhen, Y.Y., Chen, X. & Zhang, Y.D., 2013. On the integration of Ordovician conodont and graptolite biostratigraphy: New examples from Gansu and Inner Mongolia in China. Alcheringa 37, 510–528. ISSN 0311-5518.

Few Ordovician successions in the world contain both biostratigraphically highly diagnostic conodonts and graptolites permitting an integration between standard biozones based on these fossil groups. The Sandbian Guanzhuang section in the vicinity of Pingliang in the Gansu Province has an outstanding graptolite record through most of the Nemagraptus gracilis and Climacograptus bicornis graptolite biozones. Calcareous interbeds in the succession yield biostratigraphically important conodonts, including some species used for biozonations in Baltoscandia and the North American Midcontinent. Likewise, the middle–upper Darriwilian Dashimen section in the Wuhai region of Inner Mongolia hosts both diverse graptolites of the Pterograptus elegans, Didymograptus murchisoni and lowermost Nemagraptus gracilis biozones, and conodonts of Midcontinent and Baltoscandic types. The distribution patterns of these index fossil groups provide an unusual opportunity to closely correlate conodont and graptolite biozones in the middle to upper Darriwilian to Sandbian interval. For instance, the base of the C. bicornis Biozone is approximately coeval with the base of the Baltoscandic B. gerdae Subbiozone and a level near the middle of the North American P. aculeata Biozone.

Zhi-hao Wang [zhwang@nigpas.ac.cn] Xu Chen [xu1936@gmail.com], and Yuan-dong Zhang [ydzhang@nigpas.ac.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; Stig M. Bergström [stig@geology.ohio-state.edu], School of Earth Sciences, Division of Earth History, The Ohio State University, Columbus, OH 43210, USA; Yong Yi Zhen [yongyi.zhen@austmus.gov.au], Australian Museum, 6 College Street, Sydney NSW 2010, Australia.     

Radialimbricatus,a new ichnogenus from the lacustrine sandstone facies of the Lower Cretaceous Jinju Formation,South Korea

Lee, D.-C. February 2017. Radialimbricatus, a new ichnogenus from the lacustrine sandstone facies of the Lower Cretaceous Jinju Formation, South Korea. Alcheringa XX, XXX–XXX.

The trace fossil Radialimbricatus igen. nov. with type species of R. bitoensis isp. nov. occurs in river-mouth-bar sandstone beds of a lacustrine sequence of the Lower Cretaceous Jinju Formation in South Korea. Radialimbricatus is a horseshoe-shaped or elliptical, convex hyporelief structure characterized by imbrication of traces with radial ridges and a median furrow. These features differentiate Radialimbricatus from arthropod ichnofossils such as Rusophycus and Cruziana, possibly cnidarian ichnofossils such as Astropolichnus and vermiform organism ichnofossils such as Oldhamia. Cross-sectional features of R. bitoensis, including upward mud protrusions at lateral margins, demonstrate that R. bitoensis can be formed as an open epigenic furrow at the water/mud interface or an endogenic burrow along the sand/mud interface at the same time. Analysis of discrete and imbricating horseshoe-shaped traces indicates that the tracemaker maintained an obligate inclined posture even when forming trails. The radial ridges and pit along the median furrow are interpreted to reflect feeding behaviour, and the imbrication to result from unidirectional and more or less regular movements to next feeding sites. The tracemaker is thought to have been an organism with an elliptical body outline, radially arranged body parts at the base of or around the body (probably for collecting and transporting foods and moving to feeding sites) and a mouth near the centre of the body, which is consistent with body plan features of arthropods, cnidarians and annelids.

Dong-Chan Lee [dclee@chungbuk.ac.kr], Department of Earth Sciences Education, Chungbuk National University, Cheongju, 28644, South Korea.     

Maradidae: a new family of vombatomorphian marsupial from the late Oligocene of Riversleigh,northwestern Queensland

Black, K., March 2007. Maradidae: a new family of vombatomorphian marsupial from the late Oligocene of Riversleigh, northwestern Queensland. Alcheringa 31, 17-32. ISSN 0311-5518.

Marada arcanum gen. et sp. nov. is described from the late Oligocene Hiatus Site, Riversleigh World Heritage Property, northwestern Queensland. Although known from only a single dentary, it is assigned to a new family Maradidae, based on a unique combination of both plesiomorphic and apomorphic features. Of the known vombatomorphians, Marada is most similar to primitive wynyardiids and diprotodontoids (palorchestids and diprotodontids). Further clarification of the phylogenetic position of Maradidae within Vombatomorphia requires discovery of upper dentitions and crania.

Karen Black [k.black@unsw.edu.au], Vertebrate Palaeontology Laboratory, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, 2052, Australia; received 17.1.2005, revised 1.6.2005.     

Pliocene corals from the Togopi Formation of the Dent Peninsula,Sabah, northeastern Borneo,Malaysia

Saw, J.V.M., Hunter, A.W., Johnson, K.G. & Abdul Rahman, A.H.B., November 2018. Pliocene corals from the Togopi Formation of the Dent Peninsula, Sabah, northeastern Borneo, Malaysia. Alcheringa 43, 291–319. ISSN 0311-5518

The palaeobiology of the Malay Archipelago region remains poorly documented, despite its present-day significance as a modern global marine biodiversity hotspot. The Togopi Formation of the Dent Peninsula, situated in Borneo on the western Sulu Sea and eastern coast of Sabah, Malaysia, preserves Plio-Pleistocene sedimentary sequences interpreted to represent localized transgressive episodes, and which have a relatively high coral diversity. Fossil corals were sampled from three available quarries on the Dent Peninsula, the sediments of which have been previously dated as 4.5–3.4?Ma in age based on foraminiferal data and radiometric analyses. These Pliocene corals are identified here based on their macromorphology, micromorphology and microstructural characteristics. In total, this study describes 28 fossil coral taxa, with 16 genera recognized and 22 taxa identified to species level, 21 of which can be confidently assigned to extant species. These new data have resulted in revised stratigraphic ranges for eight of these species. As the most comprehensive systematic survey of corals from the Pliocene of the Indo-Pacific to date, this study indicates a high diversity of corals on the margin of the Sabah Sea, Borneo, at this time, including taxa found today, thus casting doubt on the local impact of the Plio-Pleistocene extinction previously reported from faunal analyses of the central Indo-Pacific.

Jasmin V.M. Saw [varnmay@yahoo.com] Department of Petroleum Geosciences, Universiti Teknologi PETRONAS, 32610 Seri Iskandar, Perak, Malaysia; Aaron W. Hunter* [awh31@cam.ac.uk] Department of Earth Sciences, University of Cambridge, Downing Street, Cambridge CB2 3EQ, UK; Kenneth G. Johnson [k.johnson@nhm.ac.uk] Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Abdul Hadi B Abdul Rahman [hadi_rahman@utp.edu.my] Department of Petroleum Geosciences, Universiti Teknologi PETRONAS, 32610 Seri Iskandar, Perak, Malaysia. *Also affiliated with: School of Earth Sciences, The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia.     

Dendroid multicellular thallophytes preserved in a Neoproterozoic black phosphorite in southern China

Du, W., Wang, X.L., Komiya, T., Zhao, R. & Wang, Y., April 2016. Dendroid multicellular thallophytes preserved in a Neoproterozoic black phosphorite in southern China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

A new form of dendroid multicellular thallophyte is documented in the Ediacaran Doushantuo phosphorite at Weng’an, Guizhou Province, southern China. The dendroid thallophytes have variable forms, possibly owing to heteromorphic variation. Many lateral branches extend from the upper portions of the main axes; the lateral branches bear terminal vegetative vesicles, reproductive vesicles, monosporangium-like discoidal vesicles and urn-shaped pseudoparenchymatous structures. The vegetative vesicles give rise to clavate pseudoparenchymatous structures, characterized by differentiation of the thallus medulla/cortex, which might represent an early stage of thallus development. An oogamous conceptacle arising from one carpogonial vesicle forms a highly specialized goblet-shaped conceptacle. The discovery of the new dendroid multicellular thallophytes provides not only the first fossil-based evidence of the morphological complexity and tissue differentiation in the Precambrian organisms but also insights into the life cycle of the Precambrian red algae.

Wei Du* [duwei@ea.c.u-tokyo.ac.jp], School of Earth Sciences and Resources, China University of Geosciences, Beijing, PR China; Xun Lian Wang [wxl@cugb.edu.cn], School of Earth Sciences and Resources, China University of Geosciences, Beijing, 100083, PR China; Tsuyoshi Komiya [komiya@ea.c.u-tokyo.ac.jp], Graduate School of Arts and Sciences, University of Tokyo, Tokyo 153-8902, Japan. Ran Zhao [zhao_ran13@eps.s.u-tokyo.ac.jp], Department of Earth and Planetary Science, University of Tokyo, Tokyo 113-0033, Japan. Yue Wang [gzyuewang@126.com], School of Resources and Environments, Guizhou University, Guiyang, 550025, PR China. *Also affiliated with Graduate School of Arts and Sciences, University of Tokyo, Tokyo 153-8902, Japan.