Thambetolepis delicata gen. et sp. nov., an enigmatic fossil from the Early Cambrian of South Australia

The acid insoluble sclerites of Thambetolepis delicata gen. et sp. nov. of late Tommotian or immediately post-Tommotian age from near Ardrossan, South Australia, are described. These sclerites, as well as those of Sachites Meshkova 1969 and Halkieria Poulsen 1967, are from multi-sclerite animals similar to Wiwaxia Walcott 1911 together with which they constitute the new class Thambetolepidea. Biological affinities of these fossils are discussed in the light of the complex internal structure of the sclerites. Possible assignments to the opisthobranch gastropods, the polyplacophorans, or the annelids are discussed but the taxonomic position of Thambetolepis and the Thambetolepidea is unresolved.     

Notocarpos garratti gen. et sp. nov., a new Silurian mitrate carpoid from Victoria

Notocarpos garratti gen. et sp. nov. is described from the middle Ludlovian Humevale Formation of the Clonbinane district, Victoria. It is compared with similar anomalocystitid carpoids and is found to resemble most closely Allanicytidium flemingi Caster & Gill 1968 from the Early Devonian Reefton Beds of New Zealand. N. garratti provides evidence that anomalocystitids rested with the flattened thecal surface against the sea floor (i.e., an orientation opposite to that proposed by Jefferies, 1968). It is further suggested that the stele was adapted to provide a rearward mode of locomotion.     

Parotosuchus from the Early Triassic of Queensland and Western Australia

Three members of the labyrinthodont Family Capitosauridae are described. Two new species from the Arcadia Formation of the Rewan Group, Queensland, show affinities with more primitive capitosaurs and with pre-capitosaurid groups. A third skull from the Blina Shale of Western Australia is of indeterminate species. These are the first recorded capitosaurs from Queensland and Western Australia and the first Parotosuchus from a Lystrosaurus Zone fauna.     

Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted,early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia

Chamberlain, P.M., Travouillon, K.J., Archer, M. & Hand, S.J., November 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa 40, XX–XX. ISSN 0311-5518.

A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.

Philippa M. Chamberlain [philippa.chamberlain@uq.net.au], School of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia; Kenny J. Travouillon [kenny.travouillon@museum.wa.gov.au; kennytravouillon@hotmail.com], Western Australian Museum, Locked Bag 49, Welshpool DC, WA, 6986, and School of Earth Sciences, University of Queensland, St Lucia, Queensland, 4072, Australia; Michael Archer [m.archer@unsw.edu.au] and Suzanne J. Hand [s.hand@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, New South Wales, 2052, Australia.     

Chlorozyga,a new caenogastropod from the Early Carboniferous of Australia

Chlorozyga, a new genus of Australian Tournaisian (Early Carboniferous) caenogastropod, is proposed and assigned to the Imoglobidae. Chlorozyga has a larval shell like Imogloba from the Early Carboniferous of North America but differs in teleoconch morphology. As in Imogloba, the initial whorl of Chlorozyga is openly coiled, a feature unknown in Mesozoic and Recent caenogastropods. However, Chlorozyga and Imogloba belong to the Caenogastropoda, given their multi-whorled orthostrophic larval shells.     

Two middle Cambrian trilobite genera,Cyclolorenzella Kobayashi, 1960 and Jiulongshania gen. nov., from Korea and China

As is widely recognised, fossil florules are difficult to correlate because they appear to have been more controlled by environmental and ecological factors than uniform changes which reflect the passage of geological time. Here I present a worked example of a generally applicable approach to this problem in dating and mapping fossil plant associations.

Floral associations are best described and named by the classical methods of phytosociology. The three dimensional shape of an association in rocks and its palaeoecology can then be critically assessed. For example, during Middle Triassic time the following plant associations would have been encountered on a traverse from the coast to several hundred kilometres inland from the Pacific margin of Gondwanaland: Pachydermophylletum (mangrove scrub), Linguifolietum (coastal swamp woodland), Dicroidietum odontopteroidium (floodplain forest), Phoenicopsetum (levee bank scrub), Dicroidietum odontopteroidium xylopterosum (xerophytic woodland) and Johnstonietum (mallee-like woodland).

The fourth dimension of fossil plant associations (time) is best assessed from the evolution of a prominent group of plants. In conjunction with the ranges of other plant megafossils, evolutionary changes can be used to define Oppel-zones. For the Late Permian to Early Jurassic of eastern Australasia, I have used the evolution of Dicroidium and leaves of related pteridosperm plants together with other species of restricted stratigraphic range or prominent in the definition of the fossil plant associations. From this evidence four Oppel-zones can be recognised. ‘Thinnfeldia’ callipteroides Oppel-zone (Chhidruan to Smithian, 250–230 × 10⁶ years), Dicroidium zuberi Oppel-zone (Smithian to Middle Anisian, 230–220 × 10⁶ years), D. odontopteroides Oppel-zone (Late Anisian to Ladinian, 220–210 × 10⁶ years) and Yabeiella Oppel-zone (Carnian to Rhaetian, 210–200 × 10⁶ years.

In a complete revision of the pteridosperm form-genera Dicroidium, Johnstonia and Xylopteris, a large number of new combinations and the following new taxa are proposed; Dicroidium gouldii sp. nov., D. odontopteroides var. argenteum var. nov., D. odontopteroides var. moltenense var. nov., D. radiatum sp. nov., D. townrovii sp. nov. and Johnstonia coriacea var. obesa var. nov.     

Millerocaulis tekelili sp. nov., a new species of osmundalean fern from the Aptian Cerro Negro Formation (Antarctica)

Vera, E.I. iFirstarticle. Millerocaulis tekelili sp. nov., a new species of osmundalean fern from the Aptian Cerro Negro Formation (Antarctica). Alcheringa, 1–10. ISSN 0311-5518.

Ezequiel Ignacio Vera [evera@macn.gov.ar] División Paleobotánica, Museo Argentino de Ciencias Naturales 'Bernardino Rivadavia', Av. Angel Gallardo 470, C1524DJR, Buenos Aires, Argentina. Also affiliated with Área de Paleontología, Departamento de Ciencias Geológicas, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Pabellón II, Ciudad Universitaria, C1428EGA, Buenos Aires, Argentina. Received 29.10.2010, revised 14.3.2011; accepted 24.3.2011.

A new species of the osmundalean fossil morphogenus Millerocaulis Tidwell emend. Vera, Millerocaulis tekelili sp. nov. is defined, based on several permineralized stems recovered from exposures of the Lower Cretaceous Cerro Negro Formation on Livingston Island, South Shetland Islands, Antarctica. This new species is characterized by the presence of an ectophloic–dictyoxylic siphonostele, inner parenchymatic and outer sclerotic cortices, heterogeneous sclerotic ring in the petiole bases, absence of sclerenchyma associated with the petiolar xylem trace, petiolar inner cortex with sclerenchyma strands and stipular wings having a large sclerenchyma bundle and several smaller ones. The presence of non-homogeneous sclerotic rings in the petiole bases allows this new species to be clearly distinguished from other Antarctic Millerocaulis, and suggests that it may represent an intermediate form in the evolutionary lineage leading from Millerocaulis to subgenus Claytosmunda of Osmunda.     

A short-snouted trematosauroid (Tetrapoda,Temnospondyli) from the Early Triassic of Australia: the oldest known trematosaurine

The trematosauroid temnospondyl Tirraturhinus smisseni gen. et sp. nov. from the Arcadia Formation of central Queensland, Australia, is described on the basis of its rostrum. This is the first trematosaurine (short-snouted) trematosauroid from Australia, and is considered to be most closely related to Tertrema acuta from Spitzbergen. Tirraturhinus smisseni occurs alongside lonchorhynchine (long-snouted) trematosauroids in the Arcadia Formation; the co-occurrence of both trematosauroid morphotypes in that fauna is repeated in a number of non-marine Early Triassic faunas elsewhere in Pangaea. The Arcadia Formation is probably Griesbachian (earliest Triassic), so that T. smisseni is the oldest known trematosaurine.     

A lapillopsid temnospondyl from the Early Triassic of India

A temnospondyl mandible from the Panchet Formation, described as Manubrantlia khaki, new genus and species, has several apomorphic characters that indicate it belongs to the Lapillopsidae, a taxon previously known only from the Early Triassic of Australia. The postglenoid area of the lower jaw of lapillopsids is quite similar to that of lydekkerinids. However, when other characters are considered, the two families do not form a clade exclusive of all other temnospondyls, indicating that the similarity in the morphology of the postglenoid area is probably the result of convergence.     

New corystosperm ovulate fructifications from the Middle Triassic of eastern Australia

The genus Umkomasia Thomas is emended to include the corystosperm ovulate inflorescent genera Pilophorosperma Thomas and Karibacarpon Lacey. Three new species of large inflorescences are described and illustrated: Umkomasia polycarpa sp. nov. from the Esk Formation of Queensland and U. distans sp. nov. and U. sessilis sp. nov. from the Basin Creek Formation of New South Wales.     

Early Triassic mastodonsaurids (Temnospondyli,Stereospondyli) from Western Australia,with remarks on mastodonsauroid palaeobiogeography

A newly discovered partial skull of a temnospondyl from the Early Triassic Blina Shale of the Erskine Range, Western Australia, is referred to the mastodonsaurid species Watsonisuchus aliciae. A partial skull described as ‘Parotosuchus’ sp. by Warren (1980) is also referred to W. aliciae. Both taxonomic assignments should be treated with caution because of the relatively poor state of preservation of the specimens. W. aliciae is the fourth temnospondyl species to be described from the Blina Shale. The worldwide distribution of mastodonsauroids at the base of the Triassic suggests a rapid phase of radiation and dispersal following the Permo-Triassic extinction event.     

Australian Cenozoic Bryozoa, 2: Free-living Cheilostomata of the Eocene St. Vincent Basin,S.A., including Bonellina gen. nov

Schmidt, R., March 2007. Australian Cenozoic Bryozoa, 2: Free-living Cheilostomata of the Eocene St. Vincent Basin, S.A., including Bonellina gen. nov. Alcheringa 31, 67-84. ISSN 0311-5518.

Free-living bryozoans are diverse in the Eocene sediments of the St. Vincent Basin, South Australia. They include Bonellina pentagonalis gen. et sp. nov., Otionellina sp. cf. O. exigua (Tenison Woods), Otionellina sp. cf. O. cupola (Tenison Woods), Tubiporella magna (Tenison Woods), Celleporaria nummularia (Tenison Woods), and an indeterminate species only found as moulds. This diversity and abundance is highest in the sediments representing the initial transgressive marine facies, where they occur in ‘sand fauna’ bryozoan assemblages (e.g. with Melicerita and Siphonicytara). Free-living bryozoans decrease up-section and are absent from latest Eocene sediments, indicating a significant environmental shift.

Rolf Schmidt [rschmid@museum.vic.gov.au], Museum Victoria, Melbourne, Vic 3001, Australia; received 18.3.2005, revised 14.12.2005.     

Nogoa nom. nov., a replacement name for Cometia McLoughlin

MCLOUGHLIN, S. June 2012. Nogoa nom. nov., a replacement namefor Cometia McLoughlin. Alcheringa, 279–281. ISSN 0311-5518.

A junior homonym was detected among the Rigbyaceae (Glossopteridales). The replacement name Nogoa is here proposed for Cometia McLoughlin, 1990 McLoughlin, S. 1990c. “Palaeobotany and palaeoenvironments of Permian strata, Bowen Basin”. 312Queensland: PhD thesis, University of Queensland. (unpublished) [Google Scholar] with a single included species, Nogoa biloba (McLoughlin) comb. nov.

Stephen McLoughlin [steve.mcloughlin@nrm.se], Department of Paleobotany, Swedish Natural History Museum, Box 50007, S-104 05 Stockholm, Sweden. Received 29.9.2011, revised 10.10.2011.     

Belowicysta nom. nov., a new name for the Jurassic dinoflagellate cyst Belowia Riding & Helby, 2001

The Jurassic dinoflagellate cyst genus Belowia Riding & Helby, 2001 is a junior homonym of Belowia Moquin-Tandon, 1849, a genus of the Chenopodiaceae. The new generic name Belowicysta is proposed here to replace Belowia Riding & Helby, 2001.     

Caledogonia,a new smooth trigoniacean bivalve from the Triassic of New Caledonia

Caledogonia globosa gen. et sp. nov. is considered to be a member of the Myophoriidae following the classification of Newell & Boyd (1975). It is a more evolved genus than Agonisca Fleming from the Ladinian of New Zealand. Both are probably the descendants of Neoschizodus of the Andean-West Antarctic region, a dispersal route on the southern Gondwanaland coastline. Caledogonia was probably a rapid burrower in a high energy paleoenvironment. It occurs in a rhythmic sequence of greywackes and conglomeratic shell beds that lie on pre-Triassic basement. Like the associated bivalves, Maoritrigonia and Manticula, Caledogonia is an endemic genus of the Late Triassic Maorian Province.

Caledogonia globosa gen. et sp. nov. est a rapporter aux Myophoriidae, d'après la classification de Newell & Boyd (1975). Plus évolué que le genre Agonisca Fleming du Ladinien de Nouvelle Zélande, il descend probablement, comme celuici de Neoschizodus représenté dans la region Andine et West Antarctique, zone de dispersion faunique de la cote Sud des Terres de Gondwana. Caledogonia, était un fouisseur actif dans un paléoenvironnement de haute énergie et de sédimentation rapide. On le trouve aujourdhui dans une séquence rythmique de grauwackes alternant avec des lumachelles et conglomérats, transgressive sur le substratum antétriasique. Comme les bivalves associés Maoritrigonia et Manticula, Caledogonia est un genre endémique de la province máorie du Trias supérieur.     

Organic-walled microfossils from the Early Cambrian of South Australia

Microfossils which are hollow, possess a two-layered vesicle wall, and occur as single elements or, more rarely, as compound forms, have been recovered from the Early Cambrian Heatherdale Shale, on the Fleurieu Peninsula, South Australia. The microfossils, which range in size from 4 to 14.5 μm, are informally and tentatively assigned to the genus Sphaerocongregus Moorman 1974. Superficially they resemble forms assigned to Pyritosphaera Love 1958 and its probable junior synonym, Bavlinella Shepeleva 1962. Topotypes of these genera, however, have yet to be studied using SEM techniques, and their morphologic details remain uncertain. The organic composition of the present microfossils is supported by energy-dispersive X-ray analyses. Samples of the Heatherdale Shale were also analysed using pyrolysis techniques; the organic matter is, however, over-mature with respect to petroleum generation, and no geochemical assessment of original kerogen type is possible.     

First occurrence of Janospira from the Early Ordovician of Australia

Cavanosteus, a new genus of homostiid arthrodire, with C. australis (McCoy) as type species, is erected for new material from the Early Devonian (Emsian) of the Burrinjuck area, NSW. This new material compares closely with the holotype from Buchan, Victoria, identified as the central plates from the skull; it includes lower jawbones (infragnathals) lacking denticulation also closely similar to those of Homostius from the Middle Devonian of the Baltic Province and Scotland. The only bone from the dermal trunk armour referred to the new genus is an anterior dorsolateral plate. Relationships of other large Early Devonian arthrodires assigned to the family Homostiidae are discussed. Cavanosteus gen. nov. is probably the closest relative to the type genus known from the Early Devonian.     

A horned member of the labyrinthodont super-family Brachyopoidea from the Early Triassic of Queensland

A horned, short-skulled labyrinthodont amphibian from the Arcadia Formation of the Rewan Group near Bluff, Queensland, is placed in the family Chigutisauridae of the superfamily Brachyopoidea. This is the first occurrence of this family outside Argentina. The material, consisting of two skulls and three mandibles is described as Keratobrachyops australis gen. et sp. nov., and used as a basis for an examination of the relationship between the families Brachyopidae and Chigutisauridae.     

A new temnospondyl amphibian from the Lower Triassic of South America

A new temnospondyl from the Early Triassic Buena Vista Formation (Paraná Basin) in Uruguay is described. The specimen is represented by a skull table fragment which includes the left orbit, and, although fairly incomplete, shows close affinities with the basal temnospondyl clade (Dvinosaurus + Tupilakosauridae). Previously, members of this clade were only known from Upper Permian and Lower Triassic deposits from the Northern Hemisphere. The new specimen described herein together with a recent record from the Lower Triassic of South Africa (Karoo Basin), support the hypothesis of an invasion of members of the basal (Dvinosaurus + Tupilakosauridae) clade into Gondwana as early as the Late Permian.