Late Carboniferous marine invertebrate zones of eastern Australia

Brachiopods of the Marginirugus barringtonensis and Levispustula levis Zones comprise the youngest of three major Carboniferous biostratigraphic units in eastern Australia. The remarkable change in the composition of marine invertebrates between the second and third of these units, a transition that only two brachiopod species are known to survive, has been attributed to the isolation of eastern Australia and to a drop in temperature. The Marginirugus barringtonensis and Levipustula levis Zones are redefined from reference sections in the Myall region, N.S.W.; the L. levis Zone now includes the Syringothyris bifida Zone of Campbell (1961). Faunal sequences show that these two zones are closely related and that there is no hiatus between them, as formerly proposed by McKellar (1965). Conodonts identified by Mr. D. Crane indicate a late Viséan to early Namurian age for the Marginirugus barringtonensis Zone, and an age of early Namurian for the base of the Levipustula levis Zone; brachiopod evidence indicates that the L. levis Zone extends into the Westphalian. The younger Auriculispina levis Zone from the Yarrol Basin may be coeval with the Trigonotreta campbelli Zone from N.S.W. Faunas from both the L. levis and A. levis Zones are present in Argentina and many species from the L. levis Zone have affinities with forms from the Baikal region, U.S.S.R. Correlations based on the zones have eliminated the need for a hiatus between the Branch Creek and Baywulla Formations in the Yarrol Basin.

Taxa described from the Marginirugus barringtonensis and Levipustula levis Zones in the Myall region include Yagonia gibberensis Roberts gen. et sp. nov., Bulahdelia myallensis Roberts gen. et sp. nov., Alispirifer yagonensis Thompson sp. nov., A. alatus Thompson sp. nov., Licharewia bootiensis Thompson sp. nov., Spirifer pristinus (Maxwell), Spiriferi sp., Neospirifer campbelli Maxwell and N. senilis Maxwell.     

Microflora of the middle Proterozoic Balbirini Dolomite (McArthur Group) of Australia

New genera of the Spiriferidae are described from the Permian sequences of the Carnarvon, Canning and Bonaparte Gulf Basins of Western and northwestern Australia. New taxa described are Latispirifer gen. nov. with type species Latispirifer callytharrensis sp. nov. and Latispirifer amplissimus sp. nov.; Costatispirifer gen. nov. with type species Costatispirifer gracilis sp. nov.; Cratispirifer gen. nov. with type species Cratispirifer nuraensis sp. nov. and Crassispirifer gen. nov. with type species Spirifer rostalinus Hosking (1931) and Crassispirifer pinguis sp. nov. The holotype and only extant specimen of Spirifer kimberleyensis Foord (1890) is re-examined and is assigned to Crassispirifer gen. nov. with a query.     

Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia,China

SHI, C.F., WANG, Y.J., YANG, Q. & REN, D., September 2012. Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China. Alcheringa 36, 311–320. ISSN 0311-5518.

A new grammolingiid genus, Chorilingia containing four new species (C. euryptera, C. parvica, C. translucida and C. peregrina) is described and illustrated from the Jiulongshan Formation at Daohugou, Inner Mongolia, China. The new genus is differentiated mainly on the first branch of vein Rs (Rs1) separating distal to the forks of both CuA and CuP. A key to species of the genus is provided.     

Paracordyline aureonemoralis (Lomandraceae): an Eocene monocotyledon from South Australia

A monocotyledonous leaf macrofossil taxon from Golden Grove in Adelaide, South Australia is recognised as being close to several extant Australasian species of Cordyline, especially those in the C. stricta (Sims) Endl. / C. fruticosa (L.) A. Chev. complex. The fossil is assigned to the form genus Paracordyline, known previously from the Oligocene Kerguélen Islands. However, as the Golden Grove taxon differs markedly from the Kerguélen species, it is considered to be a new species, P. aureonemoralis Conran & Christophel.     

Umkomasia (megasporophyll): part 1 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed

The genus Umkomasia, a megasporophyll, belonging to the pteridosperms (seed ferns) in the family Umkomasiaceae (Corystospermaceae), is reassessed comprehensively worldwide. All previous records are analysed. Certain fertile structures previously attributed are reclassified. Umkomasia is shown to be restricted to the Triassic of Gondwana where it is associated with the genus Pteruchus, a microsporophyll, and the genus Dicroidium, a vegetative leaf. It is well represented from Argentina, Australia and southern Africa where the Molteno Formation is by far the most comprehensively sampled with eight species described. Two specimens from the upper Permian of India attributed to Umkomasia are reclassified as cf. Arberiopsis sp. A whorled fertile structure from Antarctica, previously assigned to Umkomasia, is reclassified in a new genus as Axsmithia uniramia. Another compression fossil and the permineralized Umkomasia resinosa remain as valid records from Antarctica. The material described as Umkomasia from the Triassic of China is reclassified as Stenorachis asiatica. The Lower Jurassic record from Germany is placed in a new genus as Kirchmuellia franconica. The records of Umkomasia sp. from the Rhaetic of Germany are reclassified as cf. Kirchmuellia sp. and the single specimen from the Jurassic of Libya as genus et sp. indet. The Lower Cretaceous record from Mongolia has been reclassified by other researchers as Doylea mongolica. A pictorial key to Umkomasia species is provided, geographic and stratigraphic distributions are tabulated.     

An Eocene moncotyledon from Nelly Creek,Central Australia,with affinities to Hemerocallidaceae (Lilianae: Asparagales)

A monocotyledonous partial leaf macrofossil from Eocene beds at Nelly Creek near Lake Eyre in Central Australia is described. It is recognised as being close to several extant Australasian species of Dianella Lam. ex Juss. (Hemerocallidaceae), especially the D. odorata Blume / D. bambusifolia Hallier f. complex. However, because it has unique cuticular features relative to extant Hemerocallidaceae, the fossil is assigned to the new genus and species Dianellophyllum eocenicum Conran, Christophel & Cunningham.     

A new fossil genus in Pamphiliidae (Hymenoptera) from China

A new genus with two new species, Scabolyda orientalis gen. et sp. nov. and Scabolyda incompleta sp. nov., assigned to the subfamily Juralydinae in the family Pamphiliidae are described and illustrated. They were collected from the Middle Jurassic Jiulongshan Formation and the Lower Cretaceous Yixian Formation in northeastern China. They represent the first fossil pamphiliids described from China.     

The Late Palaeozoic brachiopod genus Jakutoproductus Kashirtsev 1959 and the Jakutoproductus verchoyanicus Zone,northern Yukon Territory,Canada

Shi, Guang R., 1994:03:28. The Late Palaeozoic brachiopod genus Jakutoproductus Kashirtsev 1959 and the Jakutoproductus verchoyanicus Zone, northern Yukon Territory, Canada. Alcheringa 18, 103–120. ISBN 0311-5518.

The familial and subfamilial position, species composition, and geographic distribution of the Late Palaeozoic productid genus Jakutoproductus Kashirtsev 1959 are reviewed. Jakutoproductus is placed in the subfamily Plicatiferinae Muir-Wood & Cooper 1960 of the family Plicatiferidae. Eighteen described species from the Russian Arctic. Mongolia, northeast China, and northern Yukon Territory, Canada are assigned to Jakutoproductus. The Jakutoproductus verchoyanicus Zone of late Sakmarian to Artinskian age, most likely early Artinskian (Aktastinian), here established is based on material from the Jungle Creek Formation, northern Yukon Territory, Canada, and is correlated with the following horizons in Russia: the Osennin Horizon in the Verchoyan Mountains, the lower Munugudjak Horizon of the Kolyma-Omolon Massif, the Hipkhoshin Suite of east Zabaikal, the lower Bhang Horizon of Taimyr, and an unnamed sandstone-shale unit on the north island of Novaya Zemlya.     

Namilamadeta snideri,a new diprotodontan (Marsupialia,Vombatoidea) from the medial Miocene of South Australia

Namilamadeta snideri new genus and species, from the medial Miocene Namba Formation, was found on the western side of Lake Tarkarooloo, South Australia. It is tentatively assigned to the Wynyardiidae and may be the plesiomorphous sister group of the Vombatidae.     

A new giant python from the Pliocene Bluff Downs Local Fauna of northeastern Queensland

Liasis dubudingala n. sp., described on the basis of isolated vertebrae from the Early Pliocene Bluff Downs Local Fauna, is the largest snake known from Australia. Dependance of vertebral proportions on intracolumnar position indicates that the fossil taxon can be excluded from the Morelia/Python clade. High neural spines suggest possible affinity with Liasis olivacea, whereas a posterior dentary fragment with small teeth is unlike L. olivacea and more similar to Liasis mackloti or species of Bothrochilus and Leiopython. As these extant species have all recently been treated as members of Liasis, the new species is assigned to that genus.     

Miralinidae (Marsupialia: Phalangeroidea) from northern Australia,including the youngest occurrence of the family

Five isolated molars from two localities in the Northern Territory, the middle Miocene Bullock Creek and late Oligocene Kangaroo Well sites, are assigned to the new miralinid genus Barguru, which includes the three new species: Barguru kayir, Barguru maru and Barguru kula. The Miralinidae was previously thought to be restricted to the late Oligocene and early Miocene, but the occurrence at Bullock Creek extends the time range of this family into the middle Miocene. Analysis of metaloph development in the Miralinidae suggests that loph formation in this family followed a different trajectory to that of phalangerids.     

A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China

LI, Y.J., HAN, G., NEL, A. REN, D., PANG, H. & LIU. X.L., September 2012. A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China. Alcheringa 36, 321–324. ISSN 0311-5518.

The new petalurid species Pseudocymatophlebia boda is described from Lower Cretaceous strata of Inner Mongolia, China. It provides additional morphological characters for this genus, which has been previously recorded from the Lower Cretaceous of England. Together with Aktassia, it is the second aktassiid genus with a very wide distribution, even though this family remains known only from Eurasia. Furthermore, a new name, Brachaktassia gen. nov., is proposed to replace the brachiopod genus Aktassia Popov, 1976.     

Metaceratodus kaopen comb. nov. and M. wichmanni comb. nov., two Late Cretaceous South American species of an austral lungfish genus (Dipnoi)

Cione, A.L. & Gouiric-Cavalli, S., June 2012. Metaceratodus kaopen comb. nov. and M. wichmanni comb. nov., two Late Cretaceous South American species of an austral lungfish genus (Dipnoi). Alcheringa 36, 203–216. ISSN 0311-5518.

Metaceratodus wollastoni, an Australian species, was reported from Upper Cretaceous beds of Patagonia in 1997. Later, three new species (Ceratodus wichmanni, Ptychoceratodus kaopen and Ptychoceratodus cionei), based on scarce material, were described from the same region. Two of these species were later referred to Ferganoceratodus. After examining much more abundant and better-preserved material, we conclude that neither the occurrence of Metaceratodus wollastoni nor those of Ptychoceratodus and Ferganoceratodus in the Cretaceous of South America are supported. We consider that C. wichmanni and P. cionei are synonyms and we reassign the three putative species to Metaceratodus under two new combinations: M. kaopen comb. nov. and M. wichmanni comb. nov. Both differ from the other species of the genus in having pits over most of the occlusal surface and a different occlusal profile of the tooth plate, and most have four ridges in the lower and upper tooth plates. Metaceratodus wichmanni differs from M. kaopen in oclussal profile, inner angle, and symphysis development among other features. Metaceratodus kaopen is known from the upper Santonian–lower Campanian Anacleto Formation of Río Negro province and M. wichmanni from upper Campanian–lower Maastrichtian units of Chubut, Río Negro, Neuquén and Mendoza provinces, Argentina. The occurrence of Metaceratodus in southern South America corroborates a close biogeographical relationship with Australia in the latest Cretaceous.

Alberto Luis Cione [acione@museo.fcnym.unlp.edu.ar] and Soledad Gouiric-Cavalli [sgouiric@museo.fcnym.unlp.edu.ar], División Paleontología Vertebrados, Museo de La Plata, Paseo del Bosque s/n, W1900FWA La Plata, Argentina. Received 23.11.2010, revised 11.7.2011, accepted 7.8.2011.

     

Repairs to an injured early Middle Cambrian Trilobite,Elkedra area,Northern Territory

Wang, Y., Shih, C., Szwedo, J. &; Ren, D. iFirst article. New fossil palaeontinids (Hemiptera, Cicadomorpha, Palaeontinidae) from the Middle Jurassic of Daohugou, China. Alcheringa, 1–12. ISSN 0311-5518.

A new genus and species assigned to the extinct family Palaeontinidae, Synapocossus sciacchitanoae Wang, Shih &; Ren, is described from the Middle Jurassic of Daohugou in Inner Mongolia, China. This new genus is established based on well-preserved fossil specimens with body and complete forewings and hind wings. It differs from other described genera by the following characters: small body size, RP and M₁ coalescing for an interval on the forewings and M_3 + 4 without bifurcation on the hind wings. The RP coalescence with M₁ in Synapocossus Wang, Shih &; Ren previously reported only in Turgaiella Becker-Migdisova &; Wootton, seems to be associated with strengthening of the anterior wing margin. The intra-specific and individual variations of Synapocossus and numerous other insect fossils of northeastern China probably indicate long-lasting ecological stresses and a competitive environment in the Middle Jurassic to Early Cretaceous ecosystems.     

On the Middle Cambrian agnostoid species Agnostus fallax Linnarsson 1869

The types and other specimens of Agnostus fallax Linnarsson 1869 are examined and reinterpreted. It is shown that the application of this specific name to many specimens from around the world is largely in error. The species is considered sufficiently different from the type species of Peronopsis, P. integra (Beyrich, 1845) to warrant erection of a new genus Axagnostus with A. fallax as the type species.     

Bryozoans from the late Cretaceous Kahuitara Tuff of the Chatham Islands,New Zealand

Taylor, P.D., & Gordon, D.P., December, 2007. Bryozoans from the Late Cretaceous Kahuitara Tuff of the Chatham Islands, New Zealand. Alcheringa 31, 339-363. ISSN 0311-5518.

Fourteen bryozoan species are described from the Campanian – Maastrichtian Kahuitara Tuff of Pitt Island, substantially increasing the known diversity in this deposit from the two species recorded previously and making it the most diverse bryozoan biota yet described from the Cretaceous of Australasia. Nine of the Kahuitara Tuff bryozoans are cyclostomes, four are cheilostomes, and one is a shell-boring ctenostome. Seven new species are described: Ceriocava hakepaensis sp. nov., Tholopora australis sp. nov., Crisidmonea lanauzeorum sp. nov., Cookobryozoon cretacea sp. nov., Chiplonkarina preeceorum sp. nov. Chiplonkarina bifoliata sp. nov. and Aechmella rangiauriensis sp. nov. The remaining species are left in open nomenclature because of preservational deficiencies or lack of taxon-diagnostic gonozooids. The ctenostome family Cookobryozoidae is subsumed in the Terebriporidae. The new family Chiplonkarinidae is proposed for anascan cheilostomes previously assigned to the paraphyletic Electridae and distinguished by having primarily erect colonies with long, tubular zooids reminiscent of stenolaemates. None of the Kahuitara Tuff bryozoan species is known elsewhere, but all apart from one genus occur in roughly coeval deposits. No families regarded as particularly characteristic of the austral post-Cretaceous are evident. The relatively large number (three) of co-occurring species of Chiplonkarina is notable, as is the dominance of cyclostomes and the first record of Tholopora in the Southern Hemisphere.

Paul D. Taylor [p.taylor@nhm.ac.uk], Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, UK; Dennis P. Gordon [d.gordon@niwa.co.nz], National Institute of Water and Atmospheric Research, Private Bag 14-901, Kilbirnie, Wellington, New Zealand; received 7.3.2006, revised 3.9.2006.     

Discovery of Paleocene Prophalangopsidae (Insecta,Orthoptera) in the Jiangtang Basin,Northern Tibet,China

A new genus and species of Prophalangopsidae, Hylophalangopsis chinensis gen. and sp. nov., is described here on the basis of one well-preserved complete fore wing from the Paleocene of Northern Tibet. The differences between this species and other described species, its systematic position, and its evolutionary significance are discussed.