A Tremadocian asterozoan from Tasmania and a late Llandovery edrioasteroid from Victoria

Jell, P.A., 2014. A Tremadocian asterozoan from Tasmania and a late Llandovery edrioasteroid from Victoria. Alcheringa 38, 528–540. ISSN 0311-5518.

An asterozoan, Maydena roadsidensis gen. et sp. nov., is described from the mid-Tremadocian (La1b Zone) Florentine Valley Formation in southwestern Tasmania and is the oldest known asterozoan in the world. Although only a single, largely dissociated, specimen is available, enough is preserved to recognize distinctive ambulacral plates similar to those of Archegonaster Jaekel from the Llanvirn of the Czech Republic. Reciprocodiscus transambus n. gen., n. sp. is an isorophid edrioasteroid from the uppermost late Llandovery Springfield Formation exposed in the bed of Deep Creek, near Springfield, 65?km north northwest of Melbourne. It occurs with a low-diversity trilobite fauna indicating a deepwater, subphotic environment. This edrioasteroid has in each ambulacrum a single series of floor plates that are not visible on the oral surface, indicating its isorophid affinity and retains certain apparently primitive features that are not seen in post-Cambrian edrioasteroids, such as the very large plates of the marginal circlet, plated aboral surface and ambulacral tips extending onto the marginal circlet plates.

Peter A. Jell [p.jell@uq.edu.au] School of Earth Sciences, The University of Queensland, St Lucia Queensland 4072, Australia.     

Ordovician stromatoporoids from Tasmania

The Hunter Siltstone near Grenfell, New South Wales, contains a rich Upper Devonian fish fauna including the sinolepid Grenfellaspis and the new antiarchs Bothriolepis grenfellensis sp. nov. and Remigolepis redcliffensis sp. nov. Bothriolepis grenfellensis sp. nov. is the first bothriolepid species described from N.S.W., and R. redcliffensis sp. nov. is the first species of Remigolepis described from Australia. Traditionally, the Hunter Siltstone was considered to be uppermost Famennian or earliest Carboniferous in age based on the presence of Grenfellaspis, and the related taxon Sinolepis, which is known from the Wutung and Sanmentan formations of southeastern China. However, available data indicates the Hunter Siltstone may be early Famennian in age. Ongoing work suggests that all Famennian Bothriolepis from N.S.W., including B. grenfellensis, possess a trifid preorbital recess, but differ in other aspects of headshield morphology. In North China, the Famennian Zhongning Formation contains six species of Remigolepis and a species of Sinolepis. However, R. redcliffensis does not show any similarity to these species beyond those of Remigolepis as a whole.     

Nothofagus macrofossils from the Tertiary of Tasmania

Leaves assignable to Nothofagus from two fossil deposits in Tasmania represent the first macrofossils of this genus from the Tertiary in southeastern Australia. One fossil species, N. johnstonii, is closely related to the extant Australian species N. cunninghamii while the other fossil species, N. tasmanica, has very close affinites with the extant Australian species N. moorei. All four of these species are closely interrelated. The pollen type produced by the fossil species is unknown, since all three types are present in the microfloras. However, both N. cunninghamii and N. moorei produce N. menziesii-type pollen. The macrofossils confirm the conclusion from pollen studies that evolution in Nothofagus has occurred very slowly.     

Tertiary gymnosperms from Tasmania: Araucariaceae

Three new species of the Araucariaceae are described from leaf remains, Araucaria readiae from the Early Eocene Regatta Point flora, A. hastiensis from the Middle-Late Eocene Hasties flora, and Agathis tasmanica from the Early Oligocene-Early Miocene Little Rapid River flora. Additionally, emended diagnoses are presented for Araucarioides linearis and A. annulata. A. readiae is the first organically preserved species in the section Eutacta described from Tasmania, A. hastiensis is the first record of a species not in section Eutacta in Tasmania, and A. tasmanica is the first record of Agathis in Tasmania. These species, along with other records from south-eastern Australia indicate the presence of a high diversity of araucarian species in the region in the Early-Middle Tertiary, although no species survive there today. Climatic change and competition from angiosperms may have led to their demise in that region.     

Tertiary Nothofagus macrofossils from Cethana,Tasmania

Two new Tertiary species of Nothofagus from the Early Eocene-Oligocene deposit at Cethana represent the first reports of fossil species which are not closely related to the extant Australian species N. moorei and N. cunninghamii. N. cethanica sp. nov. is most closely related to the extant New Zealand species N. fusca and N. truncata and gives further evidence of the relatively slow evolution within this genus. The other specimen is indistinguishable from extant adult N. gunnii leaves, and has been assigned to that species. This fossil shows that the deciduous habit was probably already present in N. gunnii by the Oligocene, and this may have helped N. gunnii to survive the Late Tertiary/Quaternary glaciations. Juvenile N. gunnii foliage gives some insight into the possible origins of this species, which may have been from the same ancestral stock as N. fusca, N. truncata, and N. cethanica.     

Upper Ordovician Phacopida (Trilobita) from Tasmania

Eastonian trilobite faunas of the Gordon Group in Tasmania include the new species Ceraurinella oepiki, Erratencrinurus trippi, and Pliomerina trisulcata, as well as a reedocalymeninid probably allied to Sarrabesia Hammann & Leone, 1997. Ceraurinella and Erratencrinurus have not previously been reported from Australia, the former being predominantly Laurentian but also known from NE China, the Himalaya, and Vietnam, and the latter mostly Baltic/Laurentian. Peri-Gondwanan species of Ceraurinella appear to form a clade, within which Tasmanian and Indian (central Himalayan) taxa are closest relatives.     

Acaste (Trilobita: Phacopina) from the Early Devonian of Tasmania

Edgecombe, G.D. March 2007. Acaste (Trilobita: Phacopina) from the Early Devonian of Tasmania. Alcheringa 31, 59-66. ISSN 0311-5518.

The widespread Siluro-Devonian acastid trilobites Acaste and Acastella are known from few occurrences in Australasia, these being confined to Lochkovian strata in Victoria and New Zealand. A new species, Acaste andersoni, is described from a correlative of the Bell Shale near St Valentines Peak in north-western Tasmania. Acaste supplements the biogeographic links between the Early Devonian faunas of New Zealand, Victoria and Tasmania.

Gregory D. Edgecombe [greged@austmus.gov.au], Australian Museum, 6 College Street, Sydney, NSW 2010, Australia; received 2.3.2005, revised 6.4.2005.     

Late Silurian vertebrate microfossils from the Carnarvon Basin,Western Australia

Burrow, C.J., Turner, S., Trinajstic, K. &; Young, G.C., 27 February 2019. Late Silurian vertebrate microfossils from the Carnarvon Basin, Western Australia. Alcheringa 43, 204–219. ISSN 0311-5518.

A core sample from the offshore Pendock 1A well, Carnarvon Basin, Western Australia yielded microvertebrate residues at an horizon in the lower part of the Hamelin Formation, dated as late Silurian, ? Ludlow, based on associated conodonts. The fish fauna comprises loganelliiform thelodont scales, the ? stem gnathostome Aberrosquama occidens nov. gen. et sp., the acanthodian Nostolepis sp. aff. N. alta, and the ? stem osteichthyan Andreolepis sp. aff. A. petri. Because of the paucity of the material, and some differences between the Pendock scales and those of established species, a precise age can not be confirmed; however, the composition of the fauna at generic level most closely resembles that of late Silurian (Ludlow) assemblages from northern Eurasia.

Carole J. Burrow* [carole.burrow@gmail.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Susan Turner [palaeoich@yahoo.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Kate Trinajstic [k.trinajstic@curtin.edu.au], School of Molecular and Life Sciences, Curtin University, Perth, WA 6102, Australia; Gavin C. Young [gavin.young@anu.edu.au], Research School of Physics and Engineering, Australian National University, Canberra, ACT 2000, Australia.     

Quaternary Organic Deposit from Newton Creek Valley,Western Tasmania

Pollen and macrofossils from an organic deposit probably 44 ka BP in age record subalpine heath with abundant local Athrotaxis cupressoides. This was succeeded by subalpine heath that included Casuarina, Phyllocladus and Eucalyptus. The pollen/vegetation assemblages are comparable with present subalpine vegetation located between 850–1050 m, though the site is at 550 m and potentially below the limit of temperate rainforest (620 m). It is inferred that the vegetation altered from upper to lower subalpine heath and that the climate became warmer and wetter. The mean temperature is estimated to have been at least 3.2°C to 1.7°C colder than today. The trend of vegetation and climate change compares closely with Late Glacial changes but does not compare with changes recorded from sites of Middle Last Glacial age in the West Coast Ranges. A radiocarbon date of 34 ka BP and a trend towards warming suggests that the deposit may represent the onset of an interstadial during the Early Last Glacial Stage.     

The first record of palaeopathology from the zooarchaeological record of late Pleistocene,southwest Tasmania

Several bones from the Late Pleistocene archaeological site, Kutikina Cave, in southwest Tasmania, show pathological conditions. The majority of these specimens represent Bennett's or red‐necked wallaby (Macropus rufogriseus), with a single specimen of the Common wombat (Vombatus ursinus) recorded. While there has been extensive work in this region of Australia since the early 1980s, this is the first record of palaeopathology from the late Pleistocene of Tasmania, and the first from a human accumulation of predominately macropod material. The palaeopathology raises several questions concerning the mobility of these animals which might have increased their susceptibly to predation, while some of the effected elements are fragmented, suggesting that people utilised these bones despite their deformities. Copyright © 2008 John Wiley & Sons, Ltd.     

Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough,Tasmania

Jago, J.B., Bentley, C.J., Laurie, J.R. &; Corbett, K.B., 26 June 2018. Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough, Tasmania. Alcheringa 43, 1-17. ISSN 0311-5518.

Cambrian Series 3 and Furongian trilobites and brachiopods are described from the Adamsfield Trough in southwestern Tasmania. The oldest fossils are very poorly preserved trilobites, assigned to Asaphiscidae gen. et sp. indet. from within the Island Road Formation a short distance above the unconformity with the underlying Proterozoic Wedge River Beds. A trilobite species from within the isolated Boyd River Formation is referred to Lioparia sp. The Island Road Formation and the Boyd River Formation are stratigraphically equivalent to the Trial Ridge Beds which have previously been dated as belonging to the Lejopyge laevigata Zone. The Trial Ridge Beds are overlain unconformably by the Singing Creek Formation. In the Adamsfield, Clear Hill and Stepped Hills areas, stratigraphic equivalents of the Singing Creek Formation collectively contain the trilobites Pseudaphelaspis sp., Pseudaphelaspis? sp., Prochuangia sp., Mindycrusta sp., Nepeidae gen. et sp. indet., and Olenidae gen. et sp. indet. plus the brachiopods described herein as Billingsella sp. aff. costata, Billingsella sp. A, Billingsella sp. B and a possible member of the Billingselloidea. The Singing Creek Formation has been previously correlated with the Stigmatoa diloma Zone. The genus Lotosoides Shergold 1975 is placed in synonymy with Prochuangia Kobayashi 1935.

James B. Jago* [jim.jago@unisa.edu.au] University of South Australia, School of Natural and Built Environment, Mawson Lakes, SA 5095, Australia; Christopher J. Bentley [bigfossil@bigpond.com] 30 Albert Street, Clare, SA 5453, Australia; John R. Laurie [john.r.laurie@gmail.com] Department of Biological Sciences, Macquarie University, NSW 2109, Australia; Keith D. Corbett [keith.corbett@bigpond.com] 35 Pillinger Drive, Fern Tree, Tas 7054, Australia.     

Late quaternary Palorchestes azael (Mammalia,Diprotodontidae) from northwestern Tasmania

A fossil mandible and incisor of the diprotodontid marsupial Palorchestes azeal Owen is reported from a new locality at Pulbeena, near Smithon, in northwestern Tasmania. The fossils occurred with a piece of wood which has a ¹⁴C age of 54,200_-4,500 ^+11,000 B.P. Both fossils and wood were deposited contemporaneously in shallow-lake shell marls and swamp peat deposits of late Quaternary age. Pollen analysis indicates that this P. azael inhabited a Eucalyptus woodland.     

Glaciation of the Upper Forth River Catchment,Tasmania

The Upper Forth Valley was glaciated on at least four occasions during the late Cainozoic. The two most extensive glaciations were no younger than early Pleistocene in age. During the last glaciation the limited ice catchment at the head of the valley nourished only a short valley glacier with separate glaciers forming in the tributary valleys to the west During earlier glaciations the Forth catchment was invaded by ice from the Mersey Valley to the east.     

Glacial Stratigraphy of the Upper Mersey Valley,Tasmania

The upper Mersey Valley was glaciated on three occasions during the Pleistocene. The youngest, Rowallan Claciation, probably commenced after 28000 years B.P. The maximum ice limit was attained before 13500 years B.P., and retreat occurred before 10000years B.P. Deposits associated with Rowallan Glaciation are weakly weathered chemically. They overlie moderately weathered deposits that were formed during the Arm Glaciation, which is inferred from relative dating data to have occurred before the Last Interglacial Stage. North of the deposits and ice limits of the Arm Glaciation extremely weathered tills and rhythmites occur. They were formed by ice of the Croesus Glaciation which is inferred to be of Early Pleistocene age or older.     

The radiation of early Silurian spiriferide brachiopods,with new taxa from the Llandovery of Iran

Popov, L.E. & Cocks, L.R.M., 2013. The radiation of early Silurian spiriferide brachiopods, with new taxa from the Llandovery of Iran. Alcheringa 38, 560–566. ISSN 0311–5518.

Although there were Late Ordovician spiriferides in the superfamily Cyrtioidea, namely Eospirifer and Odakella, only the former genus survived the terminal Ordovician extinction, and only Eospirifer is known from the earliest Llandovery (Rhuddanian). However, in the succeeding mid-Llandovery (Aeronian), the spiriferides radiated to include not only more species within Eospirifer and Striispirifer in the Eospiriferidae, but also the new genus Iranospirifer described here, which is the earliest representative of the other family within the superfamily, the Hedeinopsidae. The Ordovician species were confined to the South China continent and the Boshchekul volcanic island arc in Kazakhstan, but by the Aeronian the superfamily had spread westwards to various other continents, including the main Gondwanan superterrane (which included Iran) in the early Aeronian, and Avalonia-Baltica and Laurentia in the late Aeronian. The new species Eospirifer ghobadiae and Iranospirifer qarabilensis are both described from the lower Aeronian of Iran, and there is a note on the Aeronian rhynchonellide Stegocornu, which is endemic to Iran and nearby areas.

Leonid E. Popov [lepbarry@yahoo.co.uk; leonid.popov@museumwales.ac.uk] Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK. L. Robin M. Cocks [r.cocks@nhm.ac.uk], Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK. Received 29.5.2013, revised 12.7.2013, accepted 21.7.2013.