Editorial

Kear, Benjamin P., 11 November 2019. Editorial. Alcheringa 43, XX–XX. ISSN 0311-5518.

Benjamin P. Kear [benjamin.kear@em.uu.se], Museum of Evolution, Uppsala University, 752 36 Uppsala, Sweden.     

Two new skeletons of the enigmatic,rail-like avian taxon Songzia Hou, 1990 (Songziidae) from the early Eocene of China

Sachs, S. &; Kear, B.P. November 2018. A rare new Pliensbachian plesiosaurian from the Amaltheenton Formation of Bielefeld in northwestern Germany. Alcheringa 42, 487-500. ISSN 0311-5518.

We describe a new plesiosaurian from the upper Pliensbachian Amaltheenton Formation of Bielefeld in northwestern Germany. The taxon is based upon an incomplete associated skeleton comprising part of the right mandibular ramus, several teeth, a series of cervical, pectoral, dorsal and caudal vertebrae, as well as ribs, limb girdle elements including a nearly complete right scapula, and various distal limb bones. A unique character state combination serves to distinguish the Amaltheenton Formation remains from other previously documented Early Jurassic plesiosaurians. The most important features are the presence of a longitudinal notch incising the posterior rim of the glenoid fossa and retroarticular process, and a pronounced ventrolateral shelf on the scapula, both of which constitute derived states otherwise shared with Early Cretaceous leptocleidians. However, phylogenetic analysis using a ‘total group’ Plesiosauria data-set that specifically accommodates for Pliensbachian taxa unanimously placed the Amaltheenton Formation plesiosaurian among Early–Middle Jurassic pliosaurids. This discovery is significant because it reveals unexpected homoplasy, but also because it establishes what is only the third formally named plesiosaurian taxon thus far documented from Pliensbachian strata worldwide.

Sven Sachs^* [sachs.pal@gmail.com], Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany and Im Hof 9, 51766 Engelskirchen, Germany; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden.     

A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods

Kear, B.P., Fordyce, R.E., Hiller, N. & Siversson, M., December 2017. A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods. Alcheringa 42, 461-486. ISSN 0311-5518.

THE LAST 15 years has witnessed a blossoming of research on Australasian Mesozoic marine tetrapod fossils. Much of this work has focused on amniotes, particularly those from the prolific Lower Cretaceous (Aptian–Albian) Lagerstätten of the Eromanga Basin in central and eastern Australia, and Upper Cretaceous (Campanian–Maastrichtian) sequences of the North and South islands of New Zealand. However, rare and less popularized remains have also been found in Lower Triassic–mid-Cretaceous rocks from Australia, New Zealand and the Chatham Islands, and on the tectonically proximal landmasses of New Caledonia and Timor. Currently identified taxa include estuarine–paralic rhytidostean, brachyopid, capitosaurian and trematosaurian temnospondyls from the earliest Triassic (Induan–Olenekian), Middle–Late Triassic (Anisian–Norian) eosauropterygians, and mixosaurian, shastasaurian and euichthyosaurian ichthyosaurians, Early–Middle Jurassic (Sinemurian–Bajocian) ichthyosaurians, together with plesiosauroid and rhomaleosaurid-like plesiosaurians, and diverse Early (Aptian–Albian) through to Late Cretaceous (Campanian–Maastrichtian) elasmosaurid, leptocleidid, polycotylid, probable cryptoclidid and pliosaurid plesiosaurians, as well as ophthalmosaurid ichthyosaurians, sea turtles incorporating protostegids, and mosasaurid squamates. This faunal succession evidences almost continuous occupation of southern high-palaeolatitude seas, and repeated endemic diversifications (including nascent members of some key lineages) amongst emigrant cosmopolitan clades. The primary dispersal routes are likely to have been peri-Gondwanan, with coastal migrations along the western Tethys and polar margins of the Panthalassan Ocean. However, augmentation by increasing continental fragmentation and seaway corridor connectivity probably occurred from the Middle Jurassic to Late Cretaceous. Latest Cretaceous mosasaurid and elasmosaurid taxa also reveal regional affinities with the emergent western Pacific and Weddellian austral bioprovinces. The extreme rarity, or complete absence, of many major groups prevalent elsewhere in Gondwana (e.g., tanystropheids, Triassic sauropterygians, bothremydid marine turtles, thalattosuchians and dyrosaurid crocodylomorphs) is conspicuous, and might be related to stratigraphical/collecting biases, or the predominantly higher-palaeolatitude, cooler-water Mesozoic palaeogeography of the Australasian region. Although the burgeoning record is substantial, much still awaits discovery and adequate documentation; thus Australasia is still one of the most exciting prospects for future insights into the global history of Mesozoic marine tetrapods.

Benjamin P. Kear* [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, SE-752 36 Uppsala, Sweden; R. Ewan Fordyce [ewan.fordyce@otago.ac.nz] Department of Geology, University of Otago, Post Box 56, Dunedin 9054, New Zealand; Norton Hiller [norton.hiller@gmail.com] Canterbury Museum, Rolleston Avenue, Christchurch 8013, New Zealand; Mikael Siversson [mikael.siversson@museum.wa.gov.au] Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106.     

First records from Laurentia of some middle Cambrian (Series 3) sponge spicules

Peel, J.S., February 2017. First records from Laurentia of some middle Cambrian (Series 3) sponge spicules. Alcheringa 41, 000–000. ISSN 0311-5518.

Spicules of the sponges Silicunculus Bengtson, Australispongia Dong & Knoll and Thoracospongia Mehl are described from the middle Cambrian (Cambrian Series 3) of North Greenland. The occurrences document the first records from the Cambrian of Laurentia of spicules initially reported from Australia. Obese spicules of Thoracospongia are now known to occur in strata of Cambrian Series 2 (Stage 4, Botoman) to Cambrian Series 3 (Stage 5 and Drumian) age in Australia, northern and southeastern Siberia and in the uppermost Henson Gletscher Formation (Cambrian Series 3, Stage 5) of western Peary Land. Morphologically similar obese spicules from the USA, Jordan, Iran and Sweden suggest an evolutionary trend towards armouring of the outer sponge surface during the middle and late Cambrian. New species described are Silicunculus saaqqutit sp. nov., Thoracospongia lacrimiformis sp. nov.

John S. Peel [john.peel@pal.uu.se], Department of Earth Sciences (Palaeobiology), Uppsala University, Villavägen 16, SE-752 36 Uppsala, Sweden. Received 3.7.2016; revised 13.9.2016; accepted 15.9.2016.     

Biostratigraphical and palaeogeographical implications of Early Cambrian hyoliths from the North China Platform

Pan, B., Skovsted, C.B., Sun, H.J. & Li, G.X., 18 June 2019. Biostratigraphical and palaeogeographical implications of Early Cambrian hyoliths from the North China Platform. Alcheringa 43, 351–380. ISSN 0311-5518.

A succession of diverse hyolith assemblages comprising 10 genera and 14 species are reported from the lower Cambrian Shangwan and Sanjianfang sections of the Xinji Formation, and Xiaomeiyao section of the Houjiashan Formation, which crop out along the southern margin of the North China Platform. Most of the specimens are represented by both conchs and opercula. The identified orthothecids include Conotheca australiensis, Cupitheca holocyclata, C. costellata, Neogloborilus applanatus, N. spinatus, Tegminites hymenodes, Triplicatella disdoma, T. xinjia sp. nov. and Paratriplicatella shangwanensis gen. et sp. nov. The hyolithids comprise Protomicrocornus triplicensis gen. et sp. nov., Microcornus eximius, M. petilus, Parkula bounites and Parakorilithes mammillatus. Some anomalous taxa possess characteristics of both Hyolithida and Orthothecida, such as C. australiensis, Neogloborilus and P. triplicensis. Protomicrocornus may constitute a sister group of other hyolithids. The teeth of Parkula bounites and clavicles of Parakorilithes mammillatus are documented for the first time. The hyolith assemblages from North China are probably coeval, and can be correlated with the Cambrian upper Stage 3–lower Stage 4. Many taxa are also globally distributed and have significant potential for biostratigraphical correlations. In accordance, the hyoliths from North China reveal closest compositional similarities to faunas from eastern Gondwana, and especially South Australia. However, some taxa are shared with Laurentian assemblages suggesting cosmopolitanism, and possibly planktonic larval dispersal.

Bing Pan* [bpan@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, PR China; Christian B. Skovsted [christian.skovsted@nrm.se], Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden; Haijing Sun [hjsun1987@163.com], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, PR China; Guoxiang Li [gxli@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, PR China. *Also affiliated with: University of Science and Technology of China, Hefei 230026, PR China and Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden.     

New Early Triassic Lingulidae (Brachiopoda) genera and species from South China

Peng, Y. & Shi, G.R., June, 2008. New Early Triassic Lingulidae (Brachiopoda) genera and species from South China. Alcheringa 32, 149–170. ISSN 0311-5518.

Two new genera, Sinolingularia gen. nov. and Sinoglottidia gen. nov., together with three new species, Sinolingularia huananensis gen. et sp. nov., Sinolingularia yini gen. et sp. nov. and Sinoglottidia archboldi gen. et sp. nov., are described on the basis of a large collection of well-preserved specimens from several sections straddling the Permian – Triassic boundary in South China.     

Exceptional fossils and biotas of Gondwana: the fortieth anniversary issue of Alcheringa

McLoughlin, S., September 2016. Exceptional fossils and biotas of Gondwana: the fortieth anniversary issue of Alcheringa. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

Stephen McLoughlin [steve.mcloughlin@nrm.se], Department of Palaeobiology, Swedish Museum of Natural History, Stockholm, Box 50007, S-104 05, Sweden. Submitted 22.8.2016.     

Early Cretaceous polar biotas of Victoria,southeastern Australia—an overview of research to date

Poropat, S.F., Martin, S.K., Tosolini, A.-M.P., Wagstaff, B.E, Bean, L.B., Kear, B.P., Vickers-Rich, P. &; Rich, T.H., May 2018. Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date. Alcheringa 42, 158–230. ISSN 0311-5518.

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in Victoria since the 1960s, including the Koonwarra Fossil Bed (Strzelecki Group) near Leongatha, the Dinosaur Cove and Eric the Red West sites (Otway Group) at Cape Otway, and the Flat Rocks site (Strzelecki Group) near Cape Paterson. Systematic exploration over the past five decades has resulted in the collection of thousands of fossils representing various plants, invertebrates and vertebrates. Some of the best-preserved and most diverse Hauterivian–Barremian floral assemblages in Australia derive from outcrops of the lower Strzelecki Group in the Gippsland Basin. The slightly younger Koonwarra Fossil Bed (Aptian) is a Konservat-Lagerstätte that also preserves abundant plants, including one of the oldest known flowers. In addition, insects, crustaceans (including the only syncaridans known from Australia between the Triassic and the present), arachnids (including Australia’s only known opilione), the stratigraphically youngest xiphosurans from Australia, bryozoans, unionoid molluscs and a rich assemblage of actinopterygian fish are known from the Koonwarra Fossil Bed. The oldest known—and only Mesozoic—fossil feathers from the Australian continent constitute the only evidence for tetrapods at Koonwarra. By contrast, the Barremian–Aptian-aged deposits at the Flat Rocks site, and the Aptian–Albian-aged strata at the Dinosaur Cove and Eric the Red West sites, are all dominated by tetrapod fossils, with actinopterygians and dipnoans relatively rare. Small ornithopod (=basal neornithischian) dinosaurs are numerically common, known from four partial skeletons and a multitude of isolated bones. Aquatic meiolaniform turtles constitute another prominent faunal element, represented by numerous isolated bones and articulated carapaces and plastrons. More than 50 specimens—mostly lower jaws—evince a high diversity of mammals, including monotremes, a multituberculate and several enigmatic ausktribosphenids. Relatively minor components of these fossil assemblages are diverse theropods (including birds), rare ankylosaurs and ceratopsians, pterosaurs, non-marine plesiosaurs and a lepidosaur. In the older strata of the upper Strzelecki Group, temnospondyl amphibians—the youngest known worldwide—are a conspicuous component of the fauna, whereas crocodylomorphs appear to be present only in up-sequence deposits of the Otway Group. Invertebrates are uncommon, although decapod crustaceans and unionoid bivalves have been described. Collectively, the Early Cretaceous biota of Victoria provides insights into a unique Mesozoic high-latitude palaeoenvironment and elucidates both palaeoclimatic and palaeobiogeographic changes throughout more than 25 million years of geological time.

Stephen F. Poropat*? [sporopat@swin.edu.au; stephenfporopat@gmail.com], Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Sarah K. Martin*? [sarah.martin@dmirs.wa.gov.au; martin.sarahk@gmail.com] Geological Survey of Western Australia, 100 Plain St, East Perth, Western Australia 6004, Australia; Anne-Marie P. Tosolini [a.tosolini@unimelb.edu.au] and Barbara E. Wagstaff [wagstaff@unimelb.edu.au] School of Earth Sciences, The University of Melbourne, Melbourne, Victoria 3010, Australia; Lynne B. Bean [lynne.bean@anu.edu.au] Research School of Earth Sciences, Australian National University, Acton, Canberra, Australian Capital Territory 2001, Australia; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden; Patricia Vickers-Rich§ [prich@swin.edu.au; pat.rich@monash.edu] Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Thomas H. Rich [trich@museum.vic.gov.au] Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia. *These authors contributed equally to this work. ?Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia. ?Also affiliated with: Earth and Planetary Sciences, Western Australian Museum, Welshpool, Western Australia 6101, Australia. §Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne, Victoria 3800, Australia.     

First record of Carinodens (Squamata,Mosasauridae) from the uppermost Maastrichtian of Stevns Klint,Denmark

Milàn, J., Jagt, J.W.M., Lindgren, J. & Schulp, A.S., November 2017. First record of Carinodens (Squamata, Mosasauridae) from the uppermost Maastrichtian of Stevns Klint, Denmark. Alcheringa 42, 597-602. ISSN 0311-5518.

Here we report on an important addition to the Late Cretaceous fossil record of marine reptiles from Denmark: a tooth crown of the rare durophagous mosasaur Carinodens minalmamar found in the uppermost Maastrichtian strata at the UNESCO World Heritage Site Stevns Klint. The tooth was found within the uppermost few metres of the Maastrichtian chalk placing it within the latest 50.000 years prior to the K/Pg boundary. The new find is a shed crown probably representing a tooth from the 11th to 13th position in the jaw. The tooth represents the northernmost occurrence of the genus Carinodens. Previous mosasaur finds from Denmark have all been from the hypercarnivorous mosasaurids Mosasaurus hoffmannii and Plioplatecarpus sp., thus our specimen adds a new trophic niche exploited by marine tetrapods in the food web of the latest Maastrichtian of Denmark.

Jesper Milàn* [jesperm@oesm.dk], Geomuseum Faxe/Østsjællands Museum, Østervej 2, 4640 Faxe, Denmark; John W.M. Jagt [john.jagt@maastricht.nl], Natuurhistorisch Museum Maastricht, De Bosquetplein 6–7, 6211 kJ Maastricht, the Netherlands; Johan Lindgren [johan.lindgren@geol.lu.se], Department of Geology, Lund University, Sölvegatan 12, 223 62 Lund, Sweden. Anne S. Schulp? [anne.schulp@naturalis.nl], Natuurhistorisch Museum Maastricht, De Bosquetplein 6–7, 6211 kJ Maastricht, the Netherlands. *Also affiliated with: Natural History Museum of Denmark, Øster Voldgade 5–7, 1465 Copenhagen K, Denmark. ?Also affiliated with: Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands, and Faculteit Bètawetenschappen, Vrije Universiteit Amsterdam, De Boelelaan 1085, 1081 HV Amsterdam, the Netherlands.     

The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri

Rich, T.H., Hopson, J.A., Gill, P.G., Trusler, P., Rogers-Davidson, S., Morton, S., Cifelli, R.L., Pickering, D., Kool, L., Siu, K., Burgmann, F.A., Senden, T., Evans, A.R., Wagstaff, B.E., Seegets-Villiers, D., Corfe, I.J., Flannery, T.F., Walker, K., Musser, A.M., Archer, M., Pian, R. & Vickers-Rich, P., June 2016. The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri. Alcheringa 40, xx–xx. ISSN 0311-5518.

The monotreme Teinolophos trusleri Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren, 1999 Rich, T.H., Vickers-Rich, P., Constantine, A., Flannery, T.F., Kool, L. & van Klaveren, N., 1999. Early Cretaceous mammals from Flat Rocks, Victoria, Australia. Records of the Queen Victoria Museum and Art Gallery 106, 1–34. [Google Scholar] from the Early Cretaceous of Australia is redescribed and reinterpreted here in light of additional specimens of that species and compared with the exquisitely preserved Early Cretaceous mammals from Liaoning Province, China. Together, this material indicates that although T. trusleri lacked a rod of postdentary bones contacting the dentary, as occurs in non-mammalian cynodonts and basal mammaliaforms, it did not share the condition present in all living mammals, including monotremes, of having the three auditory ossicles, which directly connect the tympanic membrane to the fenestra ovalis, being freely suspended within the middle ear cavity. Rather, T. trusleri appears to have had an intermediate condition, present in some Early Cretaceous mammals from Liaoning, in which the postdentary bones cum ear ossicles retained a connection to a persisting Meckel’s cartilage although not to the dentary. Teinolophos thus indicates that the condition of freely suspended auditory ossicles was acquired independently in monotremes and therian mammals. Much of the anterior region of the lower jaw of Teinolophos is now known, along with an isolated upper ultimate premolar. The previously unknown anterior region of the jaw is elongated and delicate as in extant monotremes, but differs in having at least seven antemolar teeth, which are separated by distinct diastemata. The dental formula of the lower jaw of Teinolophos trusleri as now known is i2 c1 p4 m5. Both the deep lower jaw and the long-rooted upper premolar indicate that Teinolophos, unlike undoubted ornithorhynchids (including the extinct Obdurodon), lacked a bill.

Thomas H. Rich [trich@museum.vic.gov.au], Sally Rogers-Davidson [srogers@museum.vic.gov.au], David Pickering [dpick@museum.vic.gov.au], Timothy F. Flannery [tim.flannery@textpublishing.com.au], Ken Walker [kwalker@museum.vic.gov.au], Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; James A. Hopson [jhopson@uchicago.edu], Department of Organismal Biology & Anatomy, University of Chicago,1025 East 57th Street, Chicago, IL 60637, USA; Pamela G. Gill [pam.gill@bristol.ac.uk], School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, U.K. and Earth Science Department, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Peter Trusler [peter@petertrusler.com.au], Lesley Kool [koollesley@gmail.com], Doris Seegets-Villiers [doris.seegets-villiers@monash.edu], Patricia Vickers-Rich [pat.rich@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Victoria 3800, Australia; Steve Morton [steve.morton@monash.edu], Karen Siu [karen.siu@monash.edu], School of Physics and Astronomy, Monash University, Victoria 3800, Australia; Richard L. Cifelli [rlc@ou.edu] Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, OK 73072, USA; Flame A. Burgmann [flame.burgmann@monash.edu], Monash Centre for Electron Microscopy, 10 Innovation Walk, Monash University, Clayton, Victoria 3800, Australia; Tim Senden [Tim.Senden@anu.edu.au], Department of Applied Mathematics, Research School of Physical Sciences and Engineering, The Australian National University, Canberra, Australian Capital Territory 0200, Australia; Alistair R. Evans [alistair.evans@monash.edu], School of Biological Sciences, Monash University, Victoria 3800, Australia; Barbara E. Wagstaff [wagstaff@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Ian J. Corfe [ian.corfe@helsinki.fi], Institute of Biotechnology, Viikinkaari 9, 00014, University of Helsinki, Finland; Anne M. Musser [anne.musser@austmus.gov.au], Australian Museum, 1 College Street, Sydney NSW 2010 Australia; Michael Archer [m.archer@unsw.edu.au], School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia; Rebecca Pian [rpian@amnh.org], Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA. Received 7.4.2016; accepted 14.4.2016.     

A new Late Silurian plant with complex branching from Xinjiang,China

Wang Yi, Fu Qiang, Xu Honghe, & Hao Shougang, June, 2007. A new Late Silurian plant with complex branching from Xinjiang, China. Alcheringa 31, 111-120. ISSN 0311-5518.

A new fossil plant is described from the middle part of the Wutubulake Formation (late Pridoli) of Xinjiang, China. This plant demonstrates at least two orders of branching. The first-order axis has pseudomonopodial branching with alternately attached second-order axes. Fertile units are alternately inserted along the second-order axis, and consist of a branching system and two sporangia at each tip. Sporangia are narrowly obovate with rounded apex and tapering base. This plant is characterized by more complex branching than other Silurian and Early Devonian plants, and is named Wutubulaka multidichotoma gen. et sp. nov., and placed under open higher-order nomenclature.     

A new antiarch placoderm from the Emsian (Early Devonian) of Wuding,Yunnan, China

Pan, Z., Zhu, M., Zhu, Y. &; Jia, L., August 2017. A new antiarch placoderm from the Emsian (Early Devonian) of Wuding, Yunnan, China. Alcheringa 42, 10–21. ISSN 0311-5518.

Wufengshania magniforaminis, a new genus and species of the Euantiarcha (Placodermi: Antiarcha), is described from the late Emsian (Early Devonian) of Wuding, Yunnan, southwestern China. The referred specimens were three-dimensionally preserved in black shales, allowing a high-resolution computed tomography reconstruction of anatomical details. The new euantiarch is characterized by a large orbital fenestra, an arched exoskeletal band around the orbital fenestra and a developed obtected nuchal area of the skull roof. Maximum parsimony analysis, using a revised data-set of antiarchs with 44 taxa and 66 characters, resolves Wufengshania gen. nov. as a member of the Bothriolepididae, which is characterized by the presence of the infraorbital sensory canal diverging on the lateral plate, and the nuchal plate with orbital facets. New analysis supports a sister group relationship between Dianolepis and the Bothriolepididae. Luquanolepis, a coeval euantiarch from the neighboring site of the new form, is referred to the Asterolepidoidei and represents the basalmost and earliest member of the Asterolepidoidei.

Zhaohui Pan* [panzhaohui@ivpp.ac.cn], Min Zhu* [zhumin@ivpp.ac.cn], You’an Zhu^? [zhuyouan@ivpp.ac.cn] and Liantao Jia [jialiantao@ivpp.ac.cn] Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, PO Box 643, Beijing 100044, PR China. *Also affiliated with University of Chinese Academy of Sciences, Beijing, 100049, PR China. ^?Also affiliated with Uppsala University, PO Box 256, 751 05 Uppsala, Sweden.     

Reassessment of the Styxosaurus snowii (Williston, 1890) holotype specimen and its implications for elasmosaurid plesiosaurian interrelationships

The holotype (KUVP 1301) of Styxosaurus snowii—one of the earliest described elasmosaurid plesiosaurians—consists of a well-preserved cranium, mandible and articulated sequence of anterior–mid-series cervical vertebrae found in the lowermost Campanian strata of the Smoky Hill Chalk Member in the Niobrara Formation of Kansas, USA. This particular specimen has proven important for recent phylogenies of Elasmosauridae, and is integral to resulting definitions of the subfamily-level clade, Styxosaurinae. Despite this, KUVP 1301 has not been redescribed or figured in detail since its original taxonomic establishment. We, therefore, re-evaluated KUVP 1301 and assessed its phylogenetic implications. Several notable character states are pertinent for diagnosing S. snowii at genus and species level: (1) an anisodont functional dentition comprising enlarged premaxillary and dentary teeth with a pair of maxillary ‘fangs’, and elongate posterior-most dentary teeth that overlap the upper tooth row; (2) a prominent dorsomedian crest extending from the tip of the premaxillary rostrum, and expanding into a low ‘mound-like’ boss between the external bony nasal openings and orbits; (3) a pronounced convex projection on the posterolateral edge of the squamosals; and (4) platycoelous post-axial cervical vertebral centra that are substantially longer than high, and bear both lateral longitudinal ridges and ventral notches. Character state comparisons with the congeneric subfamily specifier Styxosaurus browni suggest that taxonomic distinction is possible, but equivocal. We, therefore, restrict our definition of Styxosaurus to morphologies observable in KUVP 1301. Furthermore, phylogenetic analysis of our first-hand data returns inconsistent elasmosaurid intra-clade relationships, especially with regard to Styxosaurinae. Consequently, we posit that a more targeted reassessment of Elasmosauridae is necessary to resolve both species-level topologies and higher taxonomy within the group.

Sven Sachs* [sachs.pal@gmail.com] Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany; Johan Lindgren [johan.lindgren@geol.lu.se], Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden. *Also affiliated with: Im Hof 9, 51766 Engelskirchen, Germany. Received 26.5.2016; revised 13.7.2018; accepted 24.7.2018.     

Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia

Benson, R.B.J., Fitzgerald, E.M.G., Rich, T.H. & Vickers-Rich, P., 2013. Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia. Alcheringa 37, 1–6. ISSN 0311-5518

We report a large plesiosaurian tooth from the freshwater early–middle Aptian (Early Cretaceous) Eumeralla Formation of Victoria, Australia. This, combined with records of smaller plesiosaurian teeth with an alternative morphology, provides evidence for a multitaxic freshwater plesiosaurian assemblage. Dental and body size differences suggest ecological partitioning of sympatric freshwater plesiosaurians analogous to that in modern freshwater odontocete cetaceans. The evolutionarily plastic body plan of Plesiosauria may have facilitated niche differentiation and helped them to exclude ichthyosaurs from freshwater environments during the Mesozoic. However, confirmation of this hypothesis requires the discovery of more complete remains.

Roger B.J. Benson [roger.benson@earth.ox.ac.uk], Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK; Erich M.G. Fitzgerald [efitzgerald@museum.vic.gov.au], Thomas H. Rich [trich@museum.vic.gov.au], Museum Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia; Thomas H. Rich and Patricia Vickers-Rich [pat.rich@monash.edu], School of Geosciences, Monash University, Clayton, Victoria 3800, Australia. Received 30.10.2012; revised 27.1.2013; accepted 31.1.2013.     

A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin,northwestern China

Zheng, D., Wang, H., Nel, A., Dou, L., Dai, Z., Wang, B. & Zhang, H. 27 June 2019. A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin, northwestern China. Alcheringa XX, X–X. ISSN 0311-5518.

A new genus and species of campterophlebiid damsel-dragonfly, Jurassophlebia xinjiangensis gen. et sp. nov., is described from the Lower Jurassic Badaowan Formation in the Junggar Basin, northwestern China. Jurassophlebia differs from all other campterophlebiid genera in having PsA in the same orientation as the distal branch of AA, and in its uniquely open subdiscoidal cell with very acute apical angle in the hind wing. The new discovery adds to the Asian diversity of damsel-dragonflies in the earliest Jurassic.

Daran Zheng* [dranzheng@gmail.com], He Wang [hwang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Longhui Dou [345317122@qq.com], Comprehensive Geology Exploration Team, Xinjiang Coalfield Geology Bureau, West Mountain Road, Ürümqi 830000, PR China; Zhenlong Dai [286321388@qq.com], No.9 Geological Team, Xinjiang Bureau of Geology and Mineral Resources, Ürümqi 830011, PR China; Daran Zheng also affiliated with Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China.     

A new saucrosmylid lacewing (Insecta,Neuroptera) from the Middle Jurassic of Daohugou,Inner Mongolia,China

Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A., 2014. A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa 38. ISSN 0311-5518.

A new genus and new species of Saucrosmylidae (Insecta, Neuroptera) are described (Daohugosmylus castus) based on a well-preserved hindwing from the Middle Jurassic of Daohugou, Inner Mongolia, China. Daohugosmylus gen. nov. is distinguished by a large and nearly semi-circular hindwing, relatively wide R1 space possessing several rows of cells, anteriorly bent Rs, dense crossveins over the entire wing, and smooth outer margin.

Qing Liu (corresponding author) [qingliu1029@gmail.com], Haichun Zhang [hczhang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], Yan Fang [yanfang@nigpas.ac.cn], Daran Zheng [dranzheng@gmail.com], Qi Zhang [zhqi1105@126.com] and Edmund A Jarzembowski [jarzembowski2@live.co.uk], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China; secondary address of Daran Zheng & Qi Zhang, University of Chinese Academy of Sciences, Beijing, 100049, PR China; and Ed Jarzembowski, Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. Received 13.11.2013; revised 20.1.2014; accepted 21.1.2014.     

The oldest silken fungus beetle from the Early Cretaceous of southern China (Coleoptera: Cryptophagidae: Atomariinae)

Cai, C.-y. & Wang, B., 2013. The oldest silken fungus beetle from the Early Cretaceous of southern China (Coleoptera: Cryptophagidae: Atomariinae). Alcheringa 37, 1–4. ISSN 0311-5518

Atomaria cretacea sp. nov., a new silken fungus beetle, is described and figured based on an impression fossil from the Lower Cretaceous Shixi Formation at a locality near Qingxi Town, Jiangxi Province, southern China. The new species can be referred to the extant family Cryptophagidae as supported by the tiny body size, the clubbed antenna with dilated antennomere 1, closely spaced antennal insertions, and abdominal ventrite 1 being longer than the remaining ventrites. It is placed in the extant subfamily Atomariinae based on the presence of a frontoclypeal suture and the absence of gular sutures; and tentatively in Atomaria based on its body size, sub-parallel body shape, and the presence of a frontoclypeal suture.

Chen-Yang Cai [caichenyang1988@163.com], Bo Wang [savantwang@gmail.com], State Key Laboratory of Palaeobiology and Stratigraphy; Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China. Received 11.11.2012; revised 26.1.2013; accepted 31.1.2013.     

An unusual theropod frontal from the Upper Cretaceous of north Patagonia

Paulina-Carabajal, A. & Coria, R.A. 15.4.2015. An unusual theropod frontal from the Upper Cretaceous of north Patagonia. Alcheringa 39, 514–518. ISSN 0311-5518

We report an isolated left frontal (MCF-PVPH 320) corresponding to a medium-sized theropod dinosaur from the Portezuelo Formation (Coniacian) of northern Patagonia. It shows a unique combination of traits that are not present in any other known Cretaceous theropod from South America. MCF-PVPH 320 is robust and anteroposteriorly short, with a flat and smooth dorsal surface largely excavated by the supratemporal fossa. Endocranially, the olfactory bulb impression is elongate, and the olfactory tract impression is markedly shortened anteroposteriorly. MCF-PVPH 320 differs greatly from the frontals of Late Cretaceous theropods, such as abelisaurids, megaraptorines and carcharodontosaurids. In contrast, character states including the thickness of the bone, V-shaped frontoparietal suture, reduced participation on the orbital margin and markedly short olfactory tract impression suggest the presence of an unknown mid-sized to large allosauroid for the Portezuelo Formation.

Ariana Paulina-Carabajal [a.paulinacarabajal@conicet.gov.ar], CONICET-Museo Carmen Funes, Av. Córdoba 55 (8318), Plaza Huincul, Neuquén, Argentina; Rodolfo A. Coria [rcoria@unrn.edu.ar], CONICET-Instituto de Paleobiología y Geología, Universidad Nacional de Río Negro. Av. Roca 1242 (8332), Gral. Roca, Río Negro, Argentina.     

New damselflies (Odonata: Zygoptera: Hemiphlebiidae,Dysagrionidae) from mid-Cretaceous Burmese amber

Zheng, D., Zhang, Q., Nel, A., Jarzembowski, E.A., Zhou, Z., Chang, S.-C. &; Wang, B., May 2016. New damselflies (Odonata: Zygoptera: Hemiphlebiidae, Dysagrionidae) from mid-Cretaceous Burmese amber. Alcheringa XX, xxx–xxx. ISSN 0311-5518

Two damselflies, Burmahemiphlebia zhangi gen. et sp. nov. and Palaeodysagrion cretacicus gen. et sp. nov., are described from the mid-Cretaceous Burmese amber. Burmahemiphlebia zhangi is the first record of Hemiphlebiidae from this amber, although the family was cosmopolitan during the Mesozoic. It can be readily distinguished from all other members of Hemiphlebiidae in having very short MP and CuA veins, and in its rectangular discoidal cell. The new fossils support the view that hemiphlebiid damselflies were one of the dominant groups of Zygoptera during the Mesozoic. Palaeodysagrion cretacicus is the first dysagrionid damselfly from Burmese amber and the second Mesozoic representative of this predominantly Paleogene group. It differs from other members of Dysagrionidae in having a unique elongate discoidal cell. These new finds increase the diversity of damselflies in mid-Cretaceous Burmese amber.

Daran Zheng* [dranzheng@gmail.com], Su-Chin Chang [suchin@hku.hk], Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Qingqing Zhang [qqzhang@nigpas.ac.cn], Edmund A. Jarzembowski? [edj@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Zhicheng Zhou [172692308@qq.com], The PLA Information Engineering University, 62 Kexue Ave, Gaoxin District, Zhengzhou 450002, Henan, PR China. *Also affiliated with State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China. ?Also affiliated with Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. ?Also affiliated with Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China.     

A new species of the aphid family Burmitaphididae (Hemiptera: Sternorrhyncha: Aphidomorpha) from Upper Cretaceous Burmese amber

Liu, X., Qiao, G.X., Yao, Y. & Ren, D., 28 March 2019. A new species of the aphid family Burmitaphididae (Hemiptera: Sternorrhyncha: Aphidomorpha) from Upper Cretaceous Burmese amber. Alcheringa 43, 455–460. ISSN 0311-5518

A new species of extinct aphids is reported based on a fossil specimen with a relatively complete body and broken wings from Upper Cretaceous Burmese amber. Vasteantenatus reliquialaus sp. nov. (Hemiptera: Aphidomorpha: Burmitaphididae) differs from other burmitaphidids in having antennae distinctly longer than the body. The diagnosis of Burmitaphididae is emended, and a key to all species of the family is provided

Xue Liu [katharina0521@foxmail.com], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China; Gexia Qiao [qiaogx@ioz.ac.cn], Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, No. 1 Beichen West Road, Chaoyang District, Beijing, 100101, China; Yunzhi Yao* [yaoyz100@126.com], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China; Dong Ren [rendong@mail.cnu.edu.cn], Key Lab of Insect Evolution and Environmental Change, College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China.