Fauna and biostratigraphy of the Cambrian (Series 2, Stage 4; Ordian) Tempe Formation (Pertaoorrta Group), Amadeus Basin,Northern Territory

A new faunal assemblage is reported from the Tempe Formation (Cambrian Series 2, Stage 4; Ordian) retrieved from the Hermannsburg 41 drillcore, Amadeus Basin, central Australia. Two trilobite taxa, including one new species Gunnia fava sp. nov., four brachiopod taxa, including the age-diagnostic Karathele napuru (Kruse), Kostjubella djagoran (Kruse) and Micromitra nerranubawu Kruse, together with a bradoriid, helcionellids, hyoliths, echinoderms, chancelloriids, sponges and problematic tubes are described. The fauna has close links to those of the neighbouring Daly, Georgina and Wiso basins and suggests that the Tempe Formation correlates with the Australian Ordian stage (either the Redlichia forresti or Xystridura negrina assemblage zones). The Giles Creek Dolostone in the eastern Amadeus Basin, previously regarded as coeval with the Tempe Formation, has recently been reported to be of early Templetonian age in its type section. The described taxa from the Tempe Formation confirm that these two sedimentary units are not contemporaneous and that regional stratigraphic schemes should be amended.     

First records from Laurentia of some middle Cambrian (Series 3) sponge spicules

Peel, J.S., February 2017. First records from Laurentia of some middle Cambrian (Series 3) sponge spicules. Alcheringa 41, 000–000. ISSN 0311-5518.

Spicules of the sponges Silicunculus Bengtson, Australispongia Dong & Knoll and Thoracospongia Mehl are described from the middle Cambrian (Cambrian Series 3) of North Greenland. The occurrences document the first records from the Cambrian of Laurentia of spicules initially reported from Australia. Obese spicules of Thoracospongia are now known to occur in strata of Cambrian Series 2 (Stage 4, Botoman) to Cambrian Series 3 (Stage 5 and Drumian) age in Australia, northern and southeastern Siberia and in the uppermost Henson Gletscher Formation (Cambrian Series 3, Stage 5) of western Peary Land. Morphologically similar obese spicules from the USA, Jordan, Iran and Sweden suggest an evolutionary trend towards armouring of the outer sponge surface during the middle and late Cambrian. New species described are Silicunculus saaqqutit sp. nov., Thoracospongia lacrimiformis sp. nov.

John S. Peel [john.peel@pal.uu.se], Department of Earth Sciences (Palaeobiology), Uppsala University, Villavägen 16, SE-752 36 Uppsala, Sweden. Received 3.7.2016; revised 13.9.2016; accepted 15.9.2016.     

Biostratigraphic potential of Middle Cambrian hyoliths from the eastern Georgina Basin

Fourteen hyolith taxa are documented from the Middle Cambrian (Templetonian to Floran) of the eastern (Queensland) portion of the Georgina Basin, Australia, as a contribution toward a prospective Australian Cambrian hyolith biozonation. The described fauna is from the Beetle Creek Formation (including Monastery Creek Phosphorite Member) and Gowers Formation. Additionally, the enigmatic Cupittheca and some indeterminate hyoliths are figured to illustrate aspects of hyolith morphology. Guduguwan hardmani, widespread in Ordian-early Templetonian strata of northern Australia, is here recorded from the early Templetonian of the eastern Georgina Basin. A new family Gakidae is established for sulcavitide hyolithomorphs with a conch of pentagonally tabernacular transverse section, to include Gaka, Kalkatungu gen. nov. and possibly Dorsolinevitus. New genera are the hyolithid Yalarrnga mara gen. et sp. nov., sulcavitid Kulangarra kutjurru gen. et sp. nov., gakid Kalkatungu murlu gen. et sp. nov. and angusticornid Yuku tjurtu gen. et sp. nov.; new species are Loculitheca kunka sp. nov., Carinolithes tjikilirri sp. nov., ?Sololites kankari sp. nov., ?Shandongolithes thakal sp. nov., ?Gerkella thuka sp. nov. and ?Yacutolituus rakatju sp. nov. Taxa in open nomenclature are Foersteotheca cf. dubecensis, ?Holmitheca sp. and ?Dorsojugatus sp. On present knowledge, the potential for an Australian Cambrian hyolith biozonation is limited in the Early Cambrian, but for the Middle Cambrian, G. hardmani is a widespread Ordian-early Templetonian indicator, while hyolith distribution in the Monastery Creek Phosphorite Member suggests a faunal turnover at or about the incoming of Acidusus atavus which may provide a basis for biozonation in the Floran stage.     

Editorial

Similarities in late Middle Cambrian trilobite, inarticulate brachiopod and molluscan faunas are apparent between Bornholm (Denmark) and New Zealand. Tuarangia gravgaerdensis sp. nov. is described from the late Middle Cambrian Andrarum Limestone of Bornholm, Denmark. The Andrarum Limestone is stratigraphically correlated with the Tasman Formation, New Zealand, from where the genus Tuarangia was first described. A single specimen, referred to Tuarangia, has been found in an erratic boulder from NW Poland. The boulder is estimated to be of early Late Cambrian age. The initial taxonomic assignment of Tuarangia to the Bivalvia, Subclass Pteriomorphia, is upheld. The distance between Bornholm and New Zealand, the position at opposite hemispheres, and the palaeogeography in general clearly indicate isocommunities with restricted possibility of exchange of the gene pool at species level.     

Early Middle Cambrian corals from western New South Wales

Three forms, Cothonion sympomatum gen. et sp. nov. with a bi-radially septate operculum, and the new species lissa and daseia of the genus Lipopora gen. nov., are described from the early Middle Cambrian Coonigan Formation in the Mootwingee district of western New South Wales, Australia. They may be representatives of hitherto unknown groups of organisms, but they are tentatively interpreted as coelenterates that reached a level of development comparable with the Anthozoa. Cothonion is placed in the new family Cothoniidae and questionably referred to the Rugosa. Lipopora is considered to be similar to Coelenteratella Korde; both are grouped in the new family Lipoporidae and tentatively referred to the Tabulata.     

The worm Palaeoscolex from the Cambrian of NW Argentina: extending the biogeography of Cambrian priapulids to South America

Over one hundred years of palaeontological research in northwestern Argentina has provided extensive knowledge of Andean lower Palaeozoic fossil assemblages, with trilobites, graptolites, brachiopods and echinoderms being among the most prominent groups. This record is enriched by the recent discovery of soft-bodied worms in Cambrian outcrops of northwestern Argentina. Palaeoscolex sp. cf. P. ratcliffei from the Furongian Lampazar Formation in Jujuy is described, considerably expanding the biogeographical range of this genus and filling the distributional gap, between the well-known early–middle Cambrian occurrences of Palaeoscolex and those of its Ordovician species.     

Archaeocyaths from Yorke Peninsula,South Australia and archaeocyathan Early Cambrian zonation

Zhuravlev, A. Yu., & Gravestock, D. I., 1994:03:28. Archaeayaths from Yorke Peninsula, South Australia and archaeocyathan Early Cambrian zonation. Alcheringa 18, 1–54. ISSN 0311-5518.

Two assemblages of archaeocyaths are documented from Lower Cambrian outcrops and drillholes on Yorke Peninsula. South Australia. The older assemblage (11 species) occurs in the uppermost Kulpara Formation and conformably overlying basal Parara Limestone, and is equivalent to Lower Faunal Assemblage II in the Flinders Ranges. The younger assemblage (28 species plus Acanthhcyathus and Rodiocyathus) occurs in the Koolywurtie Member near the top of the Parara Limestone. Equivalent taxa are widespread in the Flinders Ranges, western New South Wales and Antarctica. Archaeocyathan distribution is now sufficiently well known to propose three assemblage zones and two informal assemblages for regional correlation. No new taxa have been added, but Erugatocyathus scutatus (Hill) and Pycnoidocyathus latiloculatus (Hill), hitherto known only from Antarctica, are found in the upper assemblage on Yorke Peninsula. Irregular archaeocyathan systematics are discussed, the ontogeny of Archaeopharetra irregularis (Tylor) clarified, and Kruseicnema Debrenne. Gravestock & Zhuravlev, represented by K. gracilis (Gordon), is fully described.     

Review of the Cambrian acrotretid brachiopod Neotreta

The Cambrian acrotretid brachiopod Neotreta Sobolev 1976 is reviewed and re-illustrated, based on new material of the type species, N. tumida Sobolev 1976, from the lower Upper Cambrian of Siberia; Neotreta orbiculata Koneva 1990, originally described from the Middle Cambrian of Kazakhstan, has been obtained from roughly coeval beds in Shropshire, England; Neotreta pusilla Koneva 1986 was unavailable for study. Two new species, N. davidi and N. karagailensis, are described from Queensland, Australia, and Kirgizia, Central Asia respectively.     

A short-snouted trematosauroid (Tetrapoda,Temnospondyli) from the Early Triassic of Australia: the oldest known trematosaurine

The trematosauroid temnospondyl Tirraturhinus smisseni gen. et sp. nov. from the Arcadia Formation of central Queensland, Australia, is described on the basis of its rostrum. This is the first trematosaurine (short-snouted) trematosauroid from Australia, and is considered to be most closely related to Tertrema acuta from Spitzbergen. Tirraturhinus smisseni occurs alongside lonchorhynchine (long-snouted) trematosauroids in the Arcadia Formation; the co-occurrence of both trematosauroid morphotypes in that fauna is repeated in a number of non-marine Early Triassic faunas elsewhere in Pangaea. The Arcadia Formation is probably Griesbachian (earliest Triassic), so that T. smisseni is the oldest known trematosaurine.     

Microdictyon plates from the lower Cambrian Ajax Limestone of South Australia: Implications for species taxonomy and diversity

A small assemblage of isolated Microdictyon plates is described from the lower Cambrian Ajax Limestone, Mt Scott Range (Flinders Ranges), South Australia. Microdictyon plates are primarily known from single, isolated, perforated phosphatic sclerites; only one species (M. sinicum) from the Chengjiang Lagerstätte is known from soft-bodied preservation of the complete organism. The isolated plates from South Australia display a wide range of morphologies potentially reflecting: 1, considerable diversification within the group at this time; 2, extensive intraspecific morphological variation; 3, different plate morphotypes along the trunk of the animal; or 4, significant ontogenetic variation in successive growth stages. The South Australian specimens are similar to several individual sclerites described from other Cambrian palaeocontinents, but appear closest to faunas described from South China. Problems associated with the taxonomy of isolated Microdictyon plates are discussed, including the lack of knowledge associated with intraspecific and/or ontogenetic variability and how individual plate morphology may relate to size or relative position along the length of the complete organism.     

A phosphatic stromatolite (Ilicta cf. composita Sidorov) from the Middle Cambrian,northern Australia

Phosphatic stromatolites from the early Middle Cambrian (Ordian) of the Georgina Basin are described and identified as Ilicta cf. composita Sidorov. Based on Öpik's interpretation of the early Middle Cambrian, the age of the Georgina Basin specimens is similar to that of the type specimens described from the late Early Cambrian of Eastern Siberia. Phosphatic stromatolites occur at the base of dolomitized and partially silicified bioherms up to 4 m thick. The phosphatic forms overlie impermeable, cemented pavement surfaces which were covered by stratiform stromatolites from which rose columnar forms up to 5 cm in height. The stromatolites were phosphatized by penecontemporaneous diagenetic reactions that took place just below the sediment water interface and above the impermeable substrates.     

Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough,Tasmania

Jago, J.B., Bentley, C.J., Laurie, J.R. &; Corbett, K.B., 26 June 2018. Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough, Tasmania. Alcheringa 43, 1-17. ISSN 0311-5518.

Cambrian Series 3 and Furongian trilobites and brachiopods are described from the Adamsfield Trough in southwestern Tasmania. The oldest fossils are very poorly preserved trilobites, assigned to Asaphiscidae gen. et sp. indet. from within the Island Road Formation a short distance above the unconformity with the underlying Proterozoic Wedge River Beds. A trilobite species from within the isolated Boyd River Formation is referred to Lioparia sp. The Island Road Formation and the Boyd River Formation are stratigraphically equivalent to the Trial Ridge Beds which have previously been dated as belonging to the Lejopyge laevigata Zone. The Trial Ridge Beds are overlain unconformably by the Singing Creek Formation. In the Adamsfield, Clear Hill and Stepped Hills areas, stratigraphic equivalents of the Singing Creek Formation collectively contain the trilobites Pseudaphelaspis sp., Pseudaphelaspis? sp., Prochuangia sp., Mindycrusta sp., Nepeidae gen. et sp. indet., and Olenidae gen. et sp. indet. plus the brachiopods described herein as Billingsella sp. aff. costata, Billingsella sp. A, Billingsella sp. B and a possible member of the Billingselloidea. The Singing Creek Formation has been previously correlated with the Stigmatoa diloma Zone. The genus Lotosoides Shergold 1975 is placed in synonymy with Prochuangia Kobayashi 1935.

James B. Jago* [jim.jago@unisa.edu.au] University of South Australia, School of Natural and Built Environment, Mawson Lakes, SA 5095, Australia; Christopher J. Bentley [bigfossil@bigpond.com] 30 Albert Street, Clare, SA 5453, Australia; John R. Laurie [john.r.laurie@gmail.com] Department of Biological Sciences, Macquarie University, NSW 2109, Australia; Keith D. Corbett [keith.corbett@bigpond.com] 35 Pillinger Drive, Fern Tree, Tas 7054, Australia.     

Late Devonian thelodonts (Agnatha) from the Gneudna Formation,Carnarvon Basin,Western Australia

Dermal denticles of two thelodont agnathans, Australolepis seddoni gen. et sp. nov., and a possible nikoliviid gen. et sp. indet. are described from the Gneudna Formation, a marine sequence of interbedded limestone and shale exposed on the eastern edge of the Carnarvon Basin, Western Australia. The age and significance of these new forms are discussed. The Gneudna Formation is Late Devonian, probably early Frasnian, which, if confirmed, makes these the youngest thelodonts known to date.     

Palaeobiogeography of the Ordovician trilobite Prosopiscus,with a new species from western New South Wales

Prosopiscus is particularly important in Ordovician palaeobiogeography because of its wide geographic distribution in Gondwana and peri-Gondwanan regions. It appears to have been confined to low palaeolatitudes, representing a characteristic member of the warm water eastern Gondwanan shelf faunas. Trends in the distribution of the Ordovician genus can be observed due to its long stratigraphic range. Prosopiscus was restricted to, and may have originated in, Australia during the late Early Ordovician (Bendigonian-Chewtonian). By the Middle Ordovician (Darriwilian), Prosopiscus had dispersed to other parts of Gondwana and peri-Gondwana, including the North and South China blocks, Tarim, central Himalayas, and the Argentine Precordillera (South America). Possible explanations for the distribution of Prosopiscus are that: (1) there were no oceanic barriers preventing dispersal of trilobites between different regions of Gondwana, thus permitting uninhibited migration over vast distances; (2) Prosopiscus was not restricted to a specific biofacies; (3) a major eustatic transgression during the early Darriwilian may have facilitated the dispersal of Prosopiscus in allowing further development and expansion of marine environments; and (4) a prolonged planktonic larval stage may have permitted wide dispersal.

Prosopiscus lauriei sp. nov. is described from the late Early Ordovician (Bendigonian-Chewtonian) Tabita Formation at Mount Arrowsmith, northwestern New South Wales, Australia. The new species is closely related to P. praecox, from the Nora Formation, Georgina Basin, central Australia, and to P. magicus from northwest China.     

Stromatolites from Adelaidean (Late Proterozoic) sequences in central and South Australia

New forms of late Precambrian stromatolites are described from the Amadeus and Georgian Basins and additional information is given on Minjaria pontifera Walter from the Bitter Springs Formation. Tesca stewartii gr. et f. nov. is described from the Boord Formation, Elleria minuta gr. et f. nov. from the Pioneer Sandstone, and Tungussia julia f. nov. from the Julie Formation. The latter is compared with Tungussia cf. T. julia from the Wonoka Formation, South Australia. Acaciella australica Walter has been recognised in the Yackah Beds of the Georgina Basin, supporting a correlation with the Bitter Springs Formation. The stromatolites described were selected because of their potential regional and global stratigraphic significance.     

Shelly fauna from the Cambrian (Miaolingian,Guzhangian) Shannon Formation and the SPICE event in the Amadeus Basin,Northern Territory

Smith, P.M., Brock, G.A. & Paterson J.R. 20 September 2019. Shelly fauna from the Cambrian (Miaolingian, Guzhangian) Shannon Formation and the SPICE event in the Amadeus Basin, Northern Territory. Alcheringa 44, 1–24. ISSN 0311-5518

A low-diversity assemblage of linguliformean brachiopods, hyoliths, molluscs, bradoriid arthropods and echinoderms (ten taxa in total) is documented from the upper Shannon Formation (Cambrian, Miaolingian, Guzhangian; Mindyallan) in the Ross River Syncline and Todd River Anticline area, NE Amadeus Basin, central Australia. Taxa described include: the linguliformean brachiopods Treptotreta jucunda Henderson & MacKinnon, 1981, Opisthotreta sp., Stittia? sp., an undetermined acrotretid and an undetermined linguloid; the molluscs Kobayashiella? heritagensis Webers, Pojeta & Yochelson, 1992 (a paragastropod), and Apoptopegma joyceae sp. nov. (a rostroconch); the bradoriid Mongolitubulus sp.; Hyolithida gen. et sp. indet.; and isolated echinoderm plates. All fauna (including previously documented trilobites) from the upper Shannon Formation disappear at the beginning of a ～5‰ positive shift in δ¹³C, identified as the Steptoean Positive Isotopic Carbon Excursion (SPICE). Despite previous links between SPICE and a global extinction, as well as the disappearance of all faunal elements in the uppermost Shannon Formation, certain taxa reported herein range through the event elsewhere in the world. The occurrence of Treptotreta jucunda and Kobayashiella? heritagensis in younger (Furongian) deposits indicates these species survived the SPICE extinction event. This study advocates for more systematic and comprehensive sampling through stratigraphic intervals where the SPICE event occurs, especially for range data of complete faunas (rather than individual taxonomic groups), to facilitate a more complete picture of diversity trends through this important phase.

P.M. Smith* [patrick.smith@austmus.gov.au], Palaeontology Department, Australian Museum Research Institute, Sydney, New South Wales, 2010, Australia; G.A. Brock [glenn.brock@mq.edu.au], Department of Biological Sciences, Macquarie University, Sydney, New South Wales, 2109, Australia. J.R. Paterson [jpater20@une.edu.au], Palaeoscience Research Centre, School of Environmental and Rural Science, University of New England, Armidale, New South Wales, 2351, Australia. *Also affiliated with: Department of Biological Sciences, Macquarie University, Sydney, New South Wales, 2109, Australia.     

Parotosuchus from the Early Triassic of Queensland and Western Australia

Three members of the labyrinthodont Family Capitosauridae are described. Two new species from the Arcadia Formation of the Rewan Group, Queensland, show affinities with more primitive capitosaurs and with pre-capitosaurid groups. A third skull from the Blina Shale of Western Australia is of indeterminate species. These are the first recorded capitosaurs from Queensland and Western Australia and the first Parotosuchus from a Lystrosaurus Zone fauna.     

Gracilitheca and Nephrotheca (Hyolitha,Orthothecida) in the Cambrian of the Barrandian area,Czech Republic

VALENT, M., FATKA, O. & MAREK, L., iFirst article. Gracilitheca and Nephrotheca (Hyolitha, Orthothecida) in the Cambrian of the Barrandian area, Czech Republic. Alcheringa, 1–10. ISSN 0311-5518.

Five orthothecid hyoliths, Gracilitheca mirabilis sp. nov., Gracilitheca triangularis sp. nov., Gracilitheca sp., Nephrotheca betula sp. nov. and Nephrotheca sp. are described from the ‘middle’ Cambrian Buchava Formation of the Skryje–Tý?ovice Basin in the Czech Republic. The new forms are based on about forty well-preserved external and internal moulds of conchs; opercula of all species remain unknown. Stratigraphic ranges and geographic distributions within the Skryje Tý?ovice Basin are established for all taxa.     

A second incisoscutid arthrodire (Pisces,Placodermi) from the Late Devonian Gogo Formation,Western Australia

Long, J. A., 1994:03:28. A second incisoscutid arthrodire (Pisces, Placodermi) from the Late Devonian Gogo Formation. Western Australia. Alcheringa 18. 59–69. ISSN 0311-5518.

A new incisoscutid arthrodire is described from the Upper Devonian Gogo Formation of Western Australia as Gogosteus sarahae gen. et sp. nov. It is characterised by its narrow headshield with cheek unit firmly attached to skull roof, crushing gnathal plates, deep postnasal plates, elongate anterior lateral plate with deeply embayed postbranchial notch and squarely cut posterior margin, and fine dermal ornamentation. The family Incisoscutidae Denison 1984 is redefined to include Incisoscutum ritchiei Dennis & Miles 1981 and Gogosteus gen. nov., and the superfamily Incisoscutoidea nov. defined to include Incisoscutidae and Camuropiscidae.     

Eoshumardia (Trilobita,Cambrian), a junior synonym of Koldinioidia

Well-preserved internal and external moulds of a shumardiid species are described from the late Furongian (Late Cambrian) “Fengshan Formation” at Poben, Napo, southwestern Guangxi, which is close to the type locality of Shumardia orientalis, the type species of Eoshumardia. All features observed on the internal moulds indicate that the new material is definitely S. orientalis, whereas the morphology of the external moulds indicates that S. orientalis should be referred to Koldinioidia. The change of generic assignment of the type species means that Eoshumardia is a junior synonym of Koldinioidia.