Reassessment of the Styxosaurus snowii (Williston, 1890) holotype specimen and its implications for elasmosaurid plesiosaurian interrelationships

The holotype (KUVP 1301) of Styxosaurus snowii—one of the earliest described elasmosaurid plesiosaurians—consists of a well-preserved cranium, mandible and articulated sequence of anterior–mid-series cervical vertebrae found in the lowermost Campanian strata of the Smoky Hill Chalk Member in the Niobrara Formation of Kansas, USA. This particular specimen has proven important for recent phylogenies of Elasmosauridae, and is integral to resulting definitions of the subfamily-level clade, Styxosaurinae. Despite this, KUVP 1301 has not been redescribed or figured in detail since its original taxonomic establishment. We, therefore, re-evaluated KUVP 1301 and assessed its phylogenetic implications. Several notable character states are pertinent for diagnosing S. snowii at genus and species level: (1) an anisodont functional dentition comprising enlarged premaxillary and dentary teeth with a pair of maxillary ‘fangs’, and elongate posterior-most dentary teeth that overlap the upper tooth row; (2) a prominent dorsomedian crest extending from the tip of the premaxillary rostrum, and expanding into a low ‘mound-like’ boss between the external bony nasal openings and orbits; (3) a pronounced convex projection on the posterolateral edge of the squamosals; and (4) platycoelous post-axial cervical vertebral centra that are substantially longer than high, and bear both lateral longitudinal ridges and ventral notches. Character state comparisons with the congeneric subfamily specifier Styxosaurus browni suggest that taxonomic distinction is possible, but equivocal. We, therefore, restrict our definition of Styxosaurus to morphologies observable in KUVP 1301. Furthermore, phylogenetic analysis of our first-hand data returns inconsistent elasmosaurid intra-clade relationships, especially with regard to Styxosaurinae. Consequently, we posit that a more targeted reassessment of Elasmosauridae is necessary to resolve both species-level topologies and higher taxonomy within the group.

Sven Sachs* [sachs.pal@gmail.com] Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany; Johan Lindgren [johan.lindgren@geol.lu.se], Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden. *Also affiliated with: Im Hof 9, 51766 Engelskirchen, Germany. Received 26.5.2016; revised 13.7.2018; accepted 24.7.2018.     

Editorial

Kear, Benjamin P., 11 November 2019. Editorial. Alcheringa 43, XX–XX. ISSN 0311-5518.

Benjamin P. Kear [benjamin.kear@em.uu.se], Museum of Evolution, Uppsala University, 752 36 Uppsala, Sweden.     

A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods

Kear, B.P., Fordyce, R.E., Hiller, N. & Siversson, M., December 2017. A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods. Alcheringa 42, 461-486. ISSN 0311-5518.

THE LAST 15 years has witnessed a blossoming of research on Australasian Mesozoic marine tetrapod fossils. Much of this work has focused on amniotes, particularly those from the prolific Lower Cretaceous (Aptian–Albian) Lagerstätten of the Eromanga Basin in central and eastern Australia, and Upper Cretaceous (Campanian–Maastrichtian) sequences of the North and South islands of New Zealand. However, rare and less popularized remains have also been found in Lower Triassic–mid-Cretaceous rocks from Australia, New Zealand and the Chatham Islands, and on the tectonically proximal landmasses of New Caledonia and Timor. Currently identified taxa include estuarine–paralic rhytidostean, brachyopid, capitosaurian and trematosaurian temnospondyls from the earliest Triassic (Induan–Olenekian), Middle–Late Triassic (Anisian–Norian) eosauropterygians, and mixosaurian, shastasaurian and euichthyosaurian ichthyosaurians, Early–Middle Jurassic (Sinemurian–Bajocian) ichthyosaurians, together with plesiosauroid and rhomaleosaurid-like plesiosaurians, and diverse Early (Aptian–Albian) through to Late Cretaceous (Campanian–Maastrichtian) elasmosaurid, leptocleidid, polycotylid, probable cryptoclidid and pliosaurid plesiosaurians, as well as ophthalmosaurid ichthyosaurians, sea turtles incorporating protostegids, and mosasaurid squamates. This faunal succession evidences almost continuous occupation of southern high-palaeolatitude seas, and repeated endemic diversifications (including nascent members of some key lineages) amongst emigrant cosmopolitan clades. The primary dispersal routes are likely to have been peri-Gondwanan, with coastal migrations along the western Tethys and polar margins of the Panthalassan Ocean. However, augmentation by increasing continental fragmentation and seaway corridor connectivity probably occurred from the Middle Jurassic to Late Cretaceous. Latest Cretaceous mosasaurid and elasmosaurid taxa also reveal regional affinities with the emergent western Pacific and Weddellian austral bioprovinces. The extreme rarity, or complete absence, of many major groups prevalent elsewhere in Gondwana (e.g., tanystropheids, Triassic sauropterygians, bothremydid marine turtles, thalattosuchians and dyrosaurid crocodylomorphs) is conspicuous, and might be related to stratigraphical/collecting biases, or the predominantly higher-palaeolatitude, cooler-water Mesozoic palaeogeography of the Australasian region. Although the burgeoning record is substantial, much still awaits discovery and adequate documentation; thus Australasia is still one of the most exciting prospects for future insights into the global history of Mesozoic marine tetrapods.

Benjamin P. Kear* [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, SE-752 36 Uppsala, Sweden; R. Ewan Fordyce [ewan.fordyce@otago.ac.nz] Department of Geology, University of Otago, Post Box 56, Dunedin 9054, New Zealand; Norton Hiller [norton.hiller@gmail.com] Canterbury Museum, Rolleston Avenue, Christchurch 8013, New Zealand; Mikael Siversson [mikael.siversson@museum.wa.gov.au] Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106.     

Early Cretaceous polar biotas of Victoria,southeastern Australia—an overview of research to date

Poropat, S.F., Martin, S.K., Tosolini, A.-M.P., Wagstaff, B.E, Bean, L.B., Kear, B.P., Vickers-Rich, P. &; Rich, T.H., May 2018. Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date. Alcheringa 42, 158–230. ISSN 0311-5518.

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in Victoria since the 1960s, including the Koonwarra Fossil Bed (Strzelecki Group) near Leongatha, the Dinosaur Cove and Eric the Red West sites (Otway Group) at Cape Otway, and the Flat Rocks site (Strzelecki Group) near Cape Paterson. Systematic exploration over the past five decades has resulted in the collection of thousands of fossils representing various plants, invertebrates and vertebrates. Some of the best-preserved and most diverse Hauterivian–Barremian floral assemblages in Australia derive from outcrops of the lower Strzelecki Group in the Gippsland Basin. The slightly younger Koonwarra Fossil Bed (Aptian) is a Konservat-Lagerstätte that also preserves abundant plants, including one of the oldest known flowers. In addition, insects, crustaceans (including the only syncaridans known from Australia between the Triassic and the present), arachnids (including Australia’s only known opilione), the stratigraphically youngest xiphosurans from Australia, bryozoans, unionoid molluscs and a rich assemblage of actinopterygian fish are known from the Koonwarra Fossil Bed. The oldest known—and only Mesozoic—fossil feathers from the Australian continent constitute the only evidence for tetrapods at Koonwarra. By contrast, the Barremian–Aptian-aged deposits at the Flat Rocks site, and the Aptian–Albian-aged strata at the Dinosaur Cove and Eric the Red West sites, are all dominated by tetrapod fossils, with actinopterygians and dipnoans relatively rare. Small ornithopod (=basal neornithischian) dinosaurs are numerically common, known from four partial skeletons and a multitude of isolated bones. Aquatic meiolaniform turtles constitute another prominent faunal element, represented by numerous isolated bones and articulated carapaces and plastrons. More than 50 specimens—mostly lower jaws—evince a high diversity of mammals, including monotremes, a multituberculate and several enigmatic ausktribosphenids. Relatively minor components of these fossil assemblages are diverse theropods (including birds), rare ankylosaurs and ceratopsians, pterosaurs, non-marine plesiosaurs and a lepidosaur. In the older strata of the upper Strzelecki Group, temnospondyl amphibians—the youngest known worldwide—are a conspicuous component of the fauna, whereas crocodylomorphs appear to be present only in up-sequence deposits of the Otway Group. Invertebrates are uncommon, although decapod crustaceans and unionoid bivalves have been described. Collectively, the Early Cretaceous biota of Victoria provides insights into a unique Mesozoic high-latitude palaeoenvironment and elucidates both palaeoclimatic and palaeobiogeographic changes throughout more than 25 million years of geological time.

Stephen F. Poropat*? [sporopat@swin.edu.au; stephenfporopat@gmail.com], Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Sarah K. Martin*? [sarah.martin@dmirs.wa.gov.au; martin.sarahk@gmail.com] Geological Survey of Western Australia, 100 Plain St, East Perth, Western Australia 6004, Australia; Anne-Marie P. Tosolini [a.tosolini@unimelb.edu.au] and Barbara E. Wagstaff [wagstaff@unimelb.edu.au] School of Earth Sciences, The University of Melbourne, Melbourne, Victoria 3010, Australia; Lynne B. Bean [lynne.bean@anu.edu.au] Research School of Earth Sciences, Australian National University, Acton, Canberra, Australian Capital Territory 2001, Australia; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden; Patricia Vickers-Rich§ [prich@swin.edu.au; pat.rich@monash.edu] Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Thomas H. Rich [trich@museum.vic.gov.au] Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia. *These authors contributed equally to this work. ?Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia. ?Also affiliated with: Earth and Planetary Sciences, Western Australian Museum, Welshpool, Western Australia 6101, Australia. §Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne, Victoria 3800, Australia.     

Metriorhynchid crocodylomorphs from the lower Kimmeridgian of Southern Germany: evidence for a new large-bodied geosaurin lineage in Europe

Abstract

Over the last two centuries, numerous exquisitely preserved thalattosuchian crocodylomorph skeletons have been found in the Jurassic strata of Southern Germany. While the majority of these specimens occur in Toarcian and upper Kimmeridgian–lower Tithonian deposits, thalattosuchian remains are otherwise rare in strata representing different stages of the Jurassic. Here, we describe skeletal elements from two large-bodied thalattosuchians attributable to the family Metriorhynchidae – these were recovered from lower Kimmeridgian sediments in Bavaria and Baden-Württemberg, respectively. These new metriorhynchid fossils are closely comparable in both stratigraphic age and dental morphology, and thus may be congeneric. Furthermore, our phylogenetic analysis suggests affinity with metriorhynchid remains from France, Switzerland, and the UK. We interpret these taxa as members of an as-yet unnamed geosaurine metriorhynchid lineage (herein termed the ‘E-clade’) from the Kimmeridgian and Tithonian of Europe, which appears to be related to species of Torvoneustes from England and Mexico, and Purranisaurus potens from Argentina, collectively contributing to ‘Subclade T’ of the tribe Geosaurini. Finally, the metriorhynchid material described herein suggests preservation as a ‘bloat and float’ carcass that underwent diagenetic dispersal within a limestone-marl-alternation deposited in an off-shore epicontinental marine environment.

Pascal Abel [pascal.abel@senckenberg.de], Senckenberg Center for Human Evolution and Palaeoenvironment, Eberhard-Karls-University Tübingen, Sigwartstraße 10, 72076 Tübingen, Germany; Sven Sachs [sachs.pal@gmail.com], Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany; Mark T. Young [marktyoung1984@gmail.com], School of GeoSciences, Grant Institute, University of Edinburgh, James Hutton Road, Edinburgh, EH9 3FE, UK.     

A new antiarch placoderm from the Emsian (Early Devonian) of Wuding,Yunnan, China

Pan, Z., Zhu, M., Zhu, Y. &; Jia, L., August 2017. A new antiarch placoderm from the Emsian (Early Devonian) of Wuding, Yunnan, China. Alcheringa 42, 10–21. ISSN 0311-5518.

Wufengshania magniforaminis, a new genus and species of the Euantiarcha (Placodermi: Antiarcha), is described from the late Emsian (Early Devonian) of Wuding, Yunnan, southwestern China. The referred specimens were three-dimensionally preserved in black shales, allowing a high-resolution computed tomography reconstruction of anatomical details. The new euantiarch is characterized by a large orbital fenestra, an arched exoskeletal band around the orbital fenestra and a developed obtected nuchal area of the skull roof. Maximum parsimony analysis, using a revised data-set of antiarchs with 44 taxa and 66 characters, resolves Wufengshania gen. nov. as a member of the Bothriolepididae, which is characterized by the presence of the infraorbital sensory canal diverging on the lateral plate, and the nuchal plate with orbital facets. New analysis supports a sister group relationship between Dianolepis and the Bothriolepididae. Luquanolepis, a coeval euantiarch from the neighboring site of the new form, is referred to the Asterolepidoidei and represents the basalmost and earliest member of the Asterolepidoidei.

Zhaohui Pan* [panzhaohui@ivpp.ac.cn], Min Zhu* [zhumin@ivpp.ac.cn], You’an Zhu^? [zhuyouan@ivpp.ac.cn] and Liantao Jia [jialiantao@ivpp.ac.cn] Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, PO Box 643, Beijing 100044, PR China. *Also affiliated with University of Chinese Academy of Sciences, Beijing, 100049, PR China. ^?Also affiliated with Uppsala University, PO Box 256, 751 05 Uppsala, Sweden.     

Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae,Caimaninae) in the Yecua Formation (late Miocene) of Bolivia

Tineo, D.E., Bona, P., Pérez, L.M., Vergani, G.D., González, G., Poiré, D.G., Gasparini, Z.N. & Legarreta, P., 1.10.2014. Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae, Caimaninae) in the Yecua Formation (late Miocene) of Bolivia. Alcheringa 39, xxx–xxx. ISSN 0311-5518

Outcrops of the Yecua Formation (late Miocene) are exposed for approximately 230 m along the La Angostura section of the Piraí River (50 km southwest of Santa Cruz de la Sierra). These reveal massive (argillic palaeosols) and laminated (quiet-water lacustrine and marsh settings) mudstones interbedded with thin sandstones containing microfossils, molluscs and vertebrate remains. Significantly, the succession hosts a giant crocodylian, Mourasuchus (Alligatoridae, Caimaninae), which is represented by both skull and postcranial fragments found in association with freshwater turtles and fishes. Mourasuchus was distributed widely from the middle Miocene of Colombia to upper Miocene of Venezuela, Brazil and Argentina, suggesting connections between major fluvial systems and an active mechanism for dispersal of South American freshwater vertebrates during the Miocene.

David Eric Tineo [tineo.d.e@gmail.com] and Daniel Gustavo Poiré [dgpoire@yahoo.com.ar], CONICET—Centro de Investigaciones Geológicas, Universidad Nacional de La Plata. Calle 1 (644), B1900FWA, La Plata, Argentina; Paula Bona [paulabona26@gmail.com] and Zulma Gasparini [zgaspari@fcnym.unlp.edu.ar], CONICET—División Paleontología Vertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Leandro Martín Pérez [pilosaperez@gmail.com] CONICET—División Paleozoología Invertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Gustavo Dardo Vergani [gvergani@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina; Gloria González Rigas [ggonzalez@pluspetrol.net]—Pluspetrol Bolivia Corporation SA, Av. Grigotá esq. Las Palmas, Santa Cruz de la Sierra, Bolivia; Pablo Legarreta [plegarreta@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina.     

A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin,northwestern China

Zheng, D., Wang, H., Nel, A., Dou, L., Dai, Z., Wang, B. & Zhang, H. 27 June 2019. A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin, northwestern China. Alcheringa XX, X–X. ISSN 0311-5518.

A new genus and species of campterophlebiid damsel-dragonfly, Jurassophlebia xinjiangensis gen. et sp. nov., is described from the Lower Jurassic Badaowan Formation in the Junggar Basin, northwestern China. Jurassophlebia differs from all other campterophlebiid genera in having PsA in the same orientation as the distal branch of AA, and in its uniquely open subdiscoidal cell with very acute apical angle in the hind wing. The new discovery adds to the Asian diversity of damsel-dragonflies in the earliest Jurassic.

Daran Zheng* [dranzheng@gmail.com], He Wang [hwang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Longhui Dou [345317122@qq.com], Comprehensive Geology Exploration Team, Xinjiang Coalfield Geology Bureau, West Mountain Road, Ürümqi 830000, PR China; Zhenlong Dai [286321388@qq.com], No.9 Geological Team, Xinjiang Bureau of Geology and Mineral Resources, Ürümqi 830011, PR China; Daran Zheng also affiliated with Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China.     

The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri

Rich, T.H., Hopson, J.A., Gill, P.G., Trusler, P., Rogers-Davidson, S., Morton, S., Cifelli, R.L., Pickering, D., Kool, L., Siu, K., Burgmann, F.A., Senden, T., Evans, A.R., Wagstaff, B.E., Seegets-Villiers, D., Corfe, I.J., Flannery, T.F., Walker, K., Musser, A.M., Archer, M., Pian, R. & Vickers-Rich, P., June 2016. The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri. Alcheringa 40, xx–xx. ISSN 0311-5518.

The monotreme Teinolophos trusleri Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren, 1999 Rich, T.H., Vickers-Rich, P., Constantine, A., Flannery, T.F., Kool, L. & van Klaveren, N., 1999. Early Cretaceous mammals from Flat Rocks, Victoria, Australia. Records of the Queen Victoria Museum and Art Gallery 106, 1–34. [Google Scholar] from the Early Cretaceous of Australia is redescribed and reinterpreted here in light of additional specimens of that species and compared with the exquisitely preserved Early Cretaceous mammals from Liaoning Province, China. Together, this material indicates that although T. trusleri lacked a rod of postdentary bones contacting the dentary, as occurs in non-mammalian cynodonts and basal mammaliaforms, it did not share the condition present in all living mammals, including monotremes, of having the three auditory ossicles, which directly connect the tympanic membrane to the fenestra ovalis, being freely suspended within the middle ear cavity. Rather, T. trusleri appears to have had an intermediate condition, present in some Early Cretaceous mammals from Liaoning, in which the postdentary bones cum ear ossicles retained a connection to a persisting Meckel’s cartilage although not to the dentary. Teinolophos thus indicates that the condition of freely suspended auditory ossicles was acquired independently in monotremes and therian mammals. Much of the anterior region of the lower jaw of Teinolophos is now known, along with an isolated upper ultimate premolar. The previously unknown anterior region of the jaw is elongated and delicate as in extant monotremes, but differs in having at least seven antemolar teeth, which are separated by distinct diastemata. The dental formula of the lower jaw of Teinolophos trusleri as now known is i2 c1 p4 m5. Both the deep lower jaw and the long-rooted upper premolar indicate that Teinolophos, unlike undoubted ornithorhynchids (including the extinct Obdurodon), lacked a bill.

Thomas H. Rich [trich@museum.vic.gov.au], Sally Rogers-Davidson [srogers@museum.vic.gov.au], David Pickering [dpick@museum.vic.gov.au], Timothy F. Flannery [tim.flannery@textpublishing.com.au], Ken Walker [kwalker@museum.vic.gov.au], Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; James A. Hopson [jhopson@uchicago.edu], Department of Organismal Biology & Anatomy, University of Chicago,1025 East 57th Street, Chicago, IL 60637, USA; Pamela G. Gill [pam.gill@bristol.ac.uk], School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, U.K. and Earth Science Department, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Peter Trusler [peter@petertrusler.com.au], Lesley Kool [koollesley@gmail.com], Doris Seegets-Villiers [doris.seegets-villiers@monash.edu], Patricia Vickers-Rich [pat.rich@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Victoria 3800, Australia; Steve Morton [steve.morton@monash.edu], Karen Siu [karen.siu@monash.edu], School of Physics and Astronomy, Monash University, Victoria 3800, Australia; Richard L. Cifelli [rlc@ou.edu] Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, OK 73072, USA; Flame A. Burgmann [flame.burgmann@monash.edu], Monash Centre for Electron Microscopy, 10 Innovation Walk, Monash University, Clayton, Victoria 3800, Australia; Tim Senden [Tim.Senden@anu.edu.au], Department of Applied Mathematics, Research School of Physical Sciences and Engineering, The Australian National University, Canberra, Australian Capital Territory 0200, Australia; Alistair R. Evans [alistair.evans@monash.edu], School of Biological Sciences, Monash University, Victoria 3800, Australia; Barbara E. Wagstaff [wagstaff@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Ian J. Corfe [ian.corfe@helsinki.fi], Institute of Biotechnology, Viikinkaari 9, 00014, University of Helsinki, Finland; Anne M. Musser [anne.musser@austmus.gov.au], Australian Museum, 1 College Street, Sydney NSW 2010 Australia; Michael Archer [m.archer@unsw.edu.au], School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia; Rebecca Pian [rpian@amnh.org], Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA. Received 7.4.2016; accepted 14.4.2016.     

A new saucrosmylid lacewing (Insecta,Neuroptera) from the Middle Jurassic of Daohugou,Inner Mongolia,China

Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A., 2014. A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa 38. ISSN 0311-5518.

A new genus and new species of Saucrosmylidae (Insecta, Neuroptera) are described (Daohugosmylus castus) based on a well-preserved hindwing from the Middle Jurassic of Daohugou, Inner Mongolia, China. Daohugosmylus gen. nov. is distinguished by a large and nearly semi-circular hindwing, relatively wide R1 space possessing several rows of cells, anteriorly bent Rs, dense crossveins over the entire wing, and smooth outer margin.

Qing Liu (corresponding author) [qingliu1029@gmail.com], Haichun Zhang [hczhang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], Yan Fang [yanfang@nigpas.ac.cn], Daran Zheng [dranzheng@gmail.com], Qi Zhang [zhqi1105@126.com] and Edmund A Jarzembowski [jarzembowski2@live.co.uk], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China; secondary address of Daran Zheng & Qi Zhang, University of Chinese Academy of Sciences, Beijing, 100049, PR China; and Ed Jarzembowski, Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. Received 13.11.2013; revised 20.1.2014; accepted 21.1.2014.     

First records of Wuchiapingian (Late Permian) conodonts in the Xainza area,Lhasa Block,Tibet, and their palaeobiogeographic implications

Yuan, D.X., Zhang, Y.C., Zhang, Y.J., Zhu, T.X. & Shen, S.Z., 2014. First records of Wuchiapingian (Late Permian) conodonts in the Xainza area, Lhasa Block, Tibet, and their palaeobiogeographic implications. Alcheringa 38, 546–556. ISSN 0311-5518.

Conodonts are among the best fossil groups to provide high-resolution biostratigraphic correlation and resolve the palaeobiogeographic evolution of the Permian. However, they have been rarely reported from the Lhasa Block in Tibet. Here we report the first discovery of Wuchiapingian (early Lopingian) conodonts from the Xiala Formation in the Lhasa Block, Tibet. This conodont fauna includes two genera and three species (Clarkina liangshanensis, C. orientalis, Iranognathus sp.). The conodont fauna indicates that the Xiala Formation previously assigned to the Guadalupian actually ranges from late Kungurian to late Wuchiapingian. The existence of the late Wuchiapingian conodont species Clarkina orientalis and C. liangshanensis in the Lhasa Block provides additional data to support the viewpoint that this block probably had been in a warm-water regime during the Wuchiapingian (Lopingian).

Dong-Xun Yuan [yuanzi55@163.com], School of Earth Sciences and Engineering, Nanjing University, 22 Hankou Road, Nanjing, 210093, PR China and State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, PR China; Yi-Chun Zhang [yczhang@nigpas.ac.cn] and Shu-Zhong Shen [szshen@nigpas.ac.cn] (corresponding author), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, PR China; Yu-Jie Zhang [zebiac@163.com] and Tong-Xing Zhu [ztongxing88@yahoo.com.cn], Chengdu Center, China Geological Survey, 2 Renming Road North, Chengdu, 610081, PR China. Received 9.1.2014; revised 1.4.2014; accepted 28.4.2014.     

A new species of Aboilus (Insecta,Orthoptera) from the Jurassic Daohugou beds of China,and discussion of forewing coloration in Aboilus

Wang, H., Li, S., Zhang, Q., Fang, Y., Wang, B. & Zhang, H., 13.02.2015. A new species of Aboilus (Insecta, Orthoptera) from the Jurassic Daohugou beds of China, and discussion of forewing coloration in Aboilus. Alcheringa 39, xxx–xxx. ISSN 0311-5518

He Wang* [wanghe0701@163.com], Sha Li* [lisha.paleo@gmail.com], Qi Zhang* [zhqi1105@126.com], Yan Fang [yanfang@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.*Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China. ?Also affiliated with Steinmann Institute, University of Bonn, Bonn 53115, Germany.

A new species of Aboilinae (Orthoptera: Prophalangopsidae), Aboilus perbellus, is described and illustrated based on three well-preserved forewings recovered from the Middle–Upper Jurassic Daohugou beds of Inner Mongolia, China. The new species differs from all congeneric forms in its special forewing coloration and features of its wing venation. To date, three types of forewing coloration have been found among different species of Aboilus at Daohugou, suggesting that these taxa inhabited different ecotopes.     

Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan,southern Guangxi,South China

Yang, T.L., He, W.H., Zhang, K.X., Wu, S.B., Zhang, Y., Yue, M.L., Wu, H.T. & Xiao, Y.F., November 2015. Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan, southern Guangxi, South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

The Talung Formation (latest Permian) and basal part of Luolou Formation (earliest Triassic) of the Dongpan section have yielded 30 bivalve species in 17 genera. Eight genera incorporating 11 species are systematically described herein, including three new species: Nuculopsis guangxiensis, Parallelodon changhsingensis and Palaeolima fangi. Two assemblages are recognized, i.e., the Hunanopecten exilis–Euchondria fusuiensis assemblage from the Talung Formation and the Claraia dieneri–Claraia griesbachi assemblage from the Luolou Formation. The former is characterized by abundant Euchondria fusuiensis, an endemic species, associated with other common genera, such as Hunanopecten, which make it unique from coeval assemblages of South China. A palaeoecological analysis indicates that the Changhsingian bivalve assemblage at Dongpan is diverse and represented by various life habits characteristic of a complex ecosystem. This also suggests that redox conditions were oxic to suboxic in deep marine environments of the southernmost Yangtze Basin during the late Changhsingian, although several episodes of anoxic perturbations and declines in palaeoproductivity saw deterioratation of local habitats and altered the taxonomic composition or population size of the bivalve fauna.

Tinglu Yang [yang@geology.so], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Weihong He* [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Shunbao Wu [shbwu@cug.edu.cn], Yang Zhang [zhangy05@163.com], Mingliang Yue [812182779@qq.com], Huiting Wu [ht_wu415@163.com] and Yifan Xiao [shadowyi@sohu.com], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

The age of the Takatika Grit,Chatham Islands,New Zealand

Hollis, C.J, Stickley, C.E., Bijl, P.K., Schiøler, P., Clowes, C.D., Li, X, Campbell, H. March 2017. The age of the Takatika Grit, Chatham Islands, New Zealand. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

The oldest Paleogene strata on Chatham Islands, east of New Zealand, are the phosphatized conglomerates and sandstones of the Takatika Grit that crops out on the northeastern coast at Tioriori and unconformably overlies the Chatham Schist. An intact Cretaceous–Paleogene boundary transition is not preserved at this locality. New biostratigraphic analysis of dinoflagellate, diatom and radiolarian microfossil assemblages confirms that the Takatika Grit is of late early–middle Paleocene (New Zealand Teurian stage) age but contains reworked microfossils of early Campanian (Early Haumurian) age. Vertebrate fossils found in this unit are inferred to be a mixture of reworked Cretaceous and in situ Paleocene bones and teeth. The overlying Tutuiri Greensand is of middle–late Paleocene age in its lower part and also contains reworked Cretaceous microfossils.

Christopher J. Hollis [c.hollis@gns.cri.nz], Chris Clowes [c.clowes@gns.cri.nz], Xun Li [x.li@gns.cri.nz], Hamish Campbell [h.campbell@gns.cri.nz], GNS Science, PO Box 30-368, Lower Hutt 5040, New Zealand; Catherine Stickley, Evolution Applied Limited, 50 Mitchell Way, Upper Rissington, Cheltenham GL54 2PL, UK [catherine.stickley@gmail.com]; Peter Bijl [p.k.bijl@uu.nl], Marine Palynology and Paleoceanography, Department of Earth Sciences, Utrecht University, 3584 CS Utrecht, the Netherlands; Poul Schiøler [poul.schioler@mgpalaeo.com.au], Morgan Goodall Palaeo, Unit 1/5 Arvida St, Malaga, WA 6090, Australia.     

A new Miocene carnivorous marsupial,Barinya kutjamarpensis (Dasyuromorphia), from central Australia

Binfield, P., Archer, M., Hand, S.J., Black, K.H., Myers, T.J., Gillespie, A.K. & Arena, D.A., June 2016. A new Miocene carnivorous marsupial, Barinya kutjamarpensis (Dasyuromorphia), from central Australia. Alcheringa 41, xx–xx. ISSN 0311-5518.

A new dasyuromorphian, Barinya kutjamarpensis sp. nov., is described on the basis of a partial dentary recovered from the Miocene Wipajiri Formation of northern South Australia. Although about the same size as the only other species of this genus, B. wangala from the Miocene faunal assemblages of the Riversleigh World Heritage Area, northwestern Queensland, it has significant differences in morphology including a very reduced talonid on M₄ and proportionately wider molars. Based on the structural differences and the more extensive wear on its teeth, the central Australian species might have consumed harder or more abrasive prey in a more silt-rich environment than its congener, which hunted in the wet early to middle Miocene forests of Riversleigh.

Pippa Binfield [pippa.binfield@outlook.com], Michael Archer [m.archer@unsw.edu.au], Suzanne J. Hand [s.hand@unsw.edu.au], Karen H. Black [k.black@unsw.edu.au], Troy J. Myers [t.myers@unsw.edu.au] Anna K. Gillespie [a.gillespie@unsw.edu.au] and Derrick A. Arena [rick@sitc.net.au], PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales 2052, Sydney, Australia.     

The first palaeontinid from the Late Jurassic of Australia (Hemiptera,Cicadomorpha, Palaeontinidae)

Chen, J., Beattie, R., Wang, B., Jiang, H., Zheng, Y. & Zhang, H., 12 April 2019. The first palaeontinid from the Late Jurassic of Australia (Hemiptera, Cicadomorpha, Palaeontinidae). Alcheringa 43, 449–454. ISSN 0311-5518.

Palaeontinidae, an extinct group of large arboreal insects, has the most diverse record among the Mesozoic Hemiptera, but only a few taxa have been reported from the Southern Hemisphere. Herein, Talbragarocossus jurassicus Chen, Beattie & Wang gen. et sp. nov., one of the earliest representatives of ‘late’ Palaeontinidae, is described and illustrated from the Upper Jurassic Talbragar Fossil Fish Bed in New South Wales, Australia. This new taxon constitutes the first representative of Palaeontinidae in Australia and the first Jurassic example in Gondwanaland, providing significant distributional and stratigraphic extensions to the family.

Jun Chen*? [rubiscada@sina.com] and Yan Zheng? [zhengyan536@163.com], Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi 276000, China. Bo Wang? [bowang@nigpas.ac.cn], Hui Jiang [huijiang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. Robert Beattie [rgbeattie@bigpond.com], Australian Museum, 1 William St., Sydney, NSW 2010, Australia. ?Also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. ?Also affiliated with: Shandong Provincial Key Laboratory of Depositional Mineralization & Sedimentary Minerals, Shandong University of Science and Technology, Qingdao, Shandong 266590, China.     

New species of Tanyderidae (Diptera) from the Jiulongshan Formation of China

Dong, F., Shih, C.K., Skibińska, K., Krzemiński, W. & Ren, D., 10.4.2015. New species of Tanyderidae (Diptera) from the Jiulongshan Formation of China. Alcheringa 39, xxx–xxx. ISSN 0311-5518

Two new tanyderid species of Praemacrochile Kalugina, 1985 (P. dryasis, P. ovalum) and one new tanyderid species of Protanyderus Handlirsch, 1909 (P. astictum) are described and illustrated from the late Middle Jurassic Jiulongshan Formation of Daohugou in eastern Inner Mongolia, China. These species are circumscribed using well-preserved fossil specimens with bodies and complete wings. We also collected and identified new material of two species of Praemacrochile (P. ansorgei Lukashevich & Krzemiński and P. chinensis, Krzemiński & Ren) and one species of Protanyderus (P. vulcanium Zhang) from the same locality.

Fei Dong [123dongfeifei@163.com], Dong Ren [rendong@mail.cnu.edu.cn] and Chungkun Shih [chungkun.shih@gmail.com], College of Life Sciences, Capital Normal University, Xisanhuanbeilu 105, Haidian District, Beijing, PR China 100048; Kornelia Skibińska [yukisiak@gmail.com] Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, Poland; Wies?aw Krzemiński [wieslawk4@gmail.com] Pedagogical University of Cracow, Faculty of Geography and Biology, Institute of Biology, ul. Podchor??ych 2, 30-084 Kraków, ma?opolskie, Poland.     

First record of Homoctenus (Tentaculitoidea,Homoctenida) from the Late Devonian of northwest Peninsular Malaysia

Meor, H.A.H., Yong, A.M., Zakaria, M.Z.Z. & Ghani, A.A., 2.6.2015. First record of Homoctenus (Tentaculitoidea, Homoctenida) from the Late Devonian of northwest Peninsular Malaysia. Alcheringa 39, 550–558. ISSN 0311-5518.

The homoctenid tentaculitoid genus Homoctenus is reported for the first time from Peninsular Malaysia. The fossils derive from the Upper Devonian Sanai Limestone, exposed in the state of Perlis, northwest Peninsular Malaysia. The fossils are closely related to Homoctenus tenuicinctus tenuicinctus and are described as Homoctenus sp. cf. H. tenuicinctus. The homoctenids were recovered from an interval containing a rich conodont assemblage, indicating a Frasnian age (Palmatolepis linguiformis Zone).

Meor Hakif Amir Hassan [meorhakif@um.edu.my], Yong Adilah Mustafa [yongadilah@gmail.com], Mohamad Z.Z. Zakaria [zulhafiszariq@siswa.um.edu.my], Azman A. Ghani [azmangeo@um.edu.my], Geology Department, University of Malaya, 50603 Kuala Lumpur, Malaysia. Received 12.4.2015; revised 27.5.2015; accepted 2.6.2015.     

A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile

Hampe, O., Witzmann, F. & Asbach, P., 2014. A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile. Alcheringa 38, xxx–xxx. ISSN 0311–5518.

A pathology of the fossil baleen whale ‘Megaptera’ hubachi from the early Pliocene of Chile is described. It is a bony outgrowth on the left side of the supraoccipital, which is interpreted as a benign bone-forming tumour (osteoma). This diagnosis is based on X-ray imaging and CT scans of the abnormal bone, revealing a homogeneously dense internal structure with no evidence for lytic areas. The osteoma described here in ‘Megaptera’ hubachi is the first unequivocal evidence of a bone tumour in a cetacean, fossil or extant.

Oliver Hampe [oliver.hampe@mfn-berlin.de] and Florian Witzmann [florian.witzmann@mfn-berlin.de], Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, D-10115 Berlin, Germany; Patrick Asbach [patrick.asbach@charite.de], Institut für Radiologie, Charité—Universitätsmedizin Berlin, Charitéplatz 1, D-10117 Berlin, Germany. Received 28.8.2013, revised 7.11.2013, accepted 12.11.2013.     

Progiraffa (Artiodactyla: Ruminantia: Giraffidae) from the Lower Siwalik Subgroup (Miocene) of Pakistan

Aftab, K., Khan, M.A., Ahmad, Z. & Akhtar, M., February 2016. Progiraffa (Artiodactyla: Ruminantia: Giraffidae) from the Lower Siwalik Subgroup (Miocene) of Pakistan. Alcheringa 40, xxx–xxx. ISSN 0311-5518

Previously, Progiraffa exigua has been reported only from the Kamlial Formation (ca 18.3–14.2 Ma) of the Siwalik Group. We record Progiraffa exigua from the Lower Siwalik Subgroup at five localities: Jaba, Chinji Rest House, Rakh Wasnal, Dhok Bun Amir Khatoon and Ghungrila, Pakistan, thus extending the range of P. exigua to the Chinji Formation of the Siwalik Group (ca 14.2–11.2 Ma).

Kiran Aftab [kiranaftab2012@gmail.com], Zaheer Ahmad [dr.zaheerahmed11@yahoo.com], Zoology Department, GC University, Lahore, Pakistan; Muhammad Akbar Khan [akbaar111@yahoo.ca], Muhammad Akhtar [drakhtarfdrc@hotmail.com], Dr Abu Bakr Fossil Display & Research Centre, Zoology Department, University of the Punjab, Lahore, Punjab, Pakistan.