Permian brachiopods from the Tupe Formation,San Juan Province,Precordillera, Argentina

The brachiopod fauna from the Tupe Formation at La Herradura Creek, located on the west flank of Perico Hill, San Juan Province, Argentina, palaeogeographically belongs to the western sector of the Paganzo basin (‘Guandacol embayment’). The stratigraphical section of the Tupe Formation at La Herradura Creek is the stratotype of the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone, was previously regarded as being of Late Carboniferous age but here is assigned to the earliest Permian (Asselian). We describe and review the biozone assemblage, which consists of Streptorhynchus inaequiornatus, Tivertonia jachalensis, Kochiproductus sp., Costatumulus sp., Coronalosia argentinensis, Tupelosia paganzoensis, Trigonotreta pericoensis, Septosyringothyris sp. aff. Septosyringothyris jaguelensis and Crurithyris? sp. This brachiopod assemblage is related to Indian and Australian Early Permian faunas and its presence in the La Herradura Creek section provides new evidence in support of an Asselian (Early Permian) age for the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone. This assemblage is also important for intra- and inter-basinal correlation because several of its characteristic species have been identified from other sections of the Paganzo basin and the Río Blanco basin. The proposed age for this biozone is consistent with the age of palynological data from slightly above the marine faunas from the stratotype locality.     

A new Late Permian brachiopod fauna from Tanggula,Qinghai-Tibet Plateau and its palaeogeographical implications

Weihong He, Jianjun Bu, Zhijun Niu & Yang Zhang, June, 2009. A new Late Permian brachiopod fauna from Tanggula, Qinghai-Tibet Plateau and its palaeogeographical implications. Alcheringa 33, 113–132. ISSN 0311-5518.

A brachiopod fauna described from the Late Permian of the Gongri and Lizai villages, northwest of the Qoima Co Lake, Tanggula area, southern Qinghai, northwest China, includes ten species in nine genera. This fauna from the (Qiangtang Block) has a strong affinity to coeval faunas of South China, sharing 74% of its species. In addition, the Late Permian Tanggula brachiopod assemblages also demonstrate a clear link with the Middle Permian brachiopod faunas of neighbouring microcontinents including the Indochina block (Malaysia and Cambodia) and the Shan-Thai block (Thailand), as suggested by the presence of Caricula sp. cf. C. salebrosa, Transennatia termierorum and Strophalosiina. This phenomenon indicates that there were relatively narrow seaways between these microcontinents to enable ready interchange of brachiopods during the Permian, and that brachiopods tended to retreat towards the palaeoequatorial region throughout the period.     

Carboniferous and Permian Rugosochonetidae (Brachiopoda) from West Spitsbergen

The rugosochonetid brachiopod species Lissochonetes geinitzianus from the Kazimovian of the Nordenskioldbreen Formation, and Dyoros (Dyoros) mucronata sp. nov., Dyoros (Dyoros) spitzbergianus and Lissochonetes superba from the Artinskian to latest Permian Kapp Starostin Formation in West Spitsbergen are described and figured. Dyoros is generally restricted to the Boreal Realm, whereas Lissochonetes is mostly distributed in the Boreal Realm, but occasionally present in the Palaeoequatorial and Gondwanan Realms.     

Early Carboniferous brachiopod faunas from the Baoshan block,west Yunnan,southwest China

38 brachiopod species in 27 genera and subgenera are described from the Yudong Formation in the Shidian-Baoshan area, west Yunnan, southwest China. New taxa include two new subgenera: Unispirifer (Septimispirifer) and Brachythyrina (Longathyrina), and seven new species: Eomarginifera yunnanensis, Marginatia cylindrica, Unispirifer (Unispirifer) xiangshanensis, Unispirifer (Septimispirifer) wafangjieensis, Brachythyrina (Brachythyrina) transversa, Brachythyrina (Longathyrina) baoshanensis, and Girtyella wafangjieensis. Based on the described material and constraints from associated coral and conodont faunas, the age of the brachiopod fauna from the Yudon Formation is considered late Tournaisian (Early Carboniferous), with a possibility extending into earlyViséan.     

An Early Permian brachiopod fauna of Gondwanan affinity from the Baoshan block,western Yunnan,China

The present paper describes and illustrates an Early Permian brachiopod fauna collected from two localities from the upper part of the type Dingjiazhai Formation near Youwang, 30 km south of Baoshan in the Baoshan block, western Yunnan, China. The brachiopod fauna is dominated by Stenoscisma sp. and Elivina yunnanensis sp. nov. and exhibits strong generic and some specific links with faunas from the Bisnain assemblage of Timor and the Callytharra Formation of Western Australia and, to a lesser extent, faunas from the Jilong Formation of southern Tibet, the Tashkazyk Formation of southeastern Pamir, the lower Toinlungkongba Formation of northwestern Tibet, the upper Pondo Group of central Tibet, and the Jimba Jimba Calcarenite of the Carnarvon Basin, Western Australia. Based on these correlations, a Late Sakmarian (Sterlitamakian) age is preferred for the Dingjiazhai brachiopod fauna. Two new species are proposed: Globiella youwangensis sp. nov. and Elivina yunnanensis sp. nov.     

Late Palaeozoic South American pectinids revised: biostratigraphical and palaeogeographical implications

Neves, J.P., Anelli, L.E., Pagani, M.A. & Simões, M.G., 2014. Late Palaeozoic South American pectinids revised: biostratigraphical and palaeogeographical implications. Alcheringa 38 ISSN 0311-5518.

A revision of the late Palaeozoic South American pectinid Heteropecten multiscalptus (Thomas) and the establishment of Heteropecten paranaensis sp. nov. have important implications for the relationship between faunal realms within South America. Late Palaeozoic bivalve faunas occur in three distinct realms in South America: a Central Gondwanic Realm with endemic taxa showing affinities to Australia, New Zealand, South Africa and Namibia, a cold Perigondwanic Realm, and a warm Extragondwanic Realm with tethyan-like affinities similar to faunas of the American Midcontinent. In South America, faunas east of the southern Andes belong to the first two realms and previous interpretations of bivalve faunas suggested biocorrelations with those of the Extragondwanic Realm because they shared the taxon Heteropecten multiscalptus (Thomas). A revision of the Peruvian and Brazilian material does not confirm this. Instead, a re-analysis suggests that two species are present, rather than one: Heteropecten multiscalptus in the Cerro Prieto Formation, Amotape Mountains (Peru; Extragondwanic Realm), and Heteropecten paranaensis sp. nov. in the upper part of the Itararé Group, Paraná Basin (Brazil; Central Gondwanic Realm). Thus, the correlation between the late Palaeozoic faunas of the Central Gondwanic and Extragondwanic Realms in South America can no longer be supported. Heteropecten paranaensis sp. nov. lived in a siliciclastic-dominated, cold, epeiric sea of Brazil and Argentina, and is morphologically similar to some Australian species, whereas the Peruvian H. multiscalptus thrived in the warm seas of the Extragondwanic Realm.

Jacqueline P. Neves [nevesjp.unesp@gmail.com], Instituto de Geociências e Ciências Exatas, Universidade Estadual Paulista, Rio Claro campus, SP, Brazil, 13506-900; Luiz E. Anelli [anelli@usp.br], Instituto de Geociências, Universidade de São Paulo, 05508–080, Brazil; M. Alejandra Pagani, [apagani@mef.org.ar], CONICET-Museo Paleontológico Egidio Feruglio (MEF), U9100GYO, Chubut, Argentina; Marcello, G. Simões [profmgsimoes@gmail.com], Instituto de Biociências, Universidade Estadual Paulista, Botucatu campus, SP, 18618-970, Brazil. Received 19.7.2013; revised 12.11.2013; accepted 19.11.2013     

Early Permian Jakutoproductus (Productida: Brachiopoda) from Patagonia,Argentina

The productid brachiopod genus Jakutoproductus, dominant in the Early Permian marine faunas of northeastern Siberia, is described for the first time from the southern hemisphere. Jakutoproductus australis sp. nov. is described from the Rio Genoa Formation, Chubut Province (Patagonia), Argentina. The age of the Patagonian species is considered to be Sakmarian (Early Permian), possibly Sterlitamakian.     

Ordovician trilobite faunas and depositional history of the Taebaeksan Basin,Korea: implications for palaeogeography

The Taebaeksan Basin occupies the central-eastern part of the Korean peninsula and was a low-relief shallow marine carbonate shelf on which the Cambro-Ordovician Choson Supergroup was deposited. In the Taebaeksan Basin trilobites are among the most dominant fossil groups in the Lower Ordovician, but they become less important in Middle Ordovician faunal assemblages. The Early Ordovician trilobite faunas of the Taebaeksan Basin are characterised by the common occurrence of pandemic genera such as Jujuyaspis, Leiostegium, Asaphellus, Protopliomerops, Hystricurus, Apatokephalus, Shumardia, Asaphopsoides, and Kayseraspis. Biogeographically significant trilobite taxa include Yosimuraspis, Dikelokephalina, Koraipsis, and Chosenia. These Ordovician trilobite faunas, which thrived in shallow marine environments, show a remarkable similarity with faunas from North China, implying that the Taebaeksan Basin was connected through contiguous shallow waters to North China. These Sino-Korean faunas exhibit a close biogeographic connection with Australian faunas, with which they share some endemic genera, whereas they are more distantly related to the faunas of South China, South America, and North America. Based on these palaeobiogeographical features, it is suggested that in the early Palaeozoic much of the present Korean peninsula including the Taebaeksan Basin belonged to the Sino-Korean block, while part of the peninsula was derived from the Yangtze block.     

Late Mississippian (Early Carboniferous) brachiopods from the western Daba Mountains,central China

Qiao L. & Shen S.Z., September 2012. Late Mississippian (Early Carboniferous) brachiopods from the western Daba Mountains, central China. Alcheringa 36, 287–309. ISSN 0311–5518.

Fifteen brachiopod species in 12 genera are described for the first time from four intervals in the middle and upper parts of the Zhanpo Formation at the Huoyanxi section near Zhenba in the western Daba Mountains, southern Shaanxi, central China. The Zhenba brachiopod fauna is dominated by diverse and abundant species of Productida, together with some species of Athyridida, Orthida, Orthotetida and Spiriferida. It ranges from late Viséan to Serpukhovian in age based on the presence of Gigantoproductus species in association with diagnostic foraminifera and conodonts. This fauna generally shows palaeobiogeographical links with the palaeoequatorial realm, including Western Europe, the Moscow Basin, the Ural Mountains, Japan, eastern Tibet and South and North China. Its closest palaeobiogeographical affinity is with South China assemblages rather than those of North and Northwest China, therefore, indicating that the Zhenba area was palaeogeographically close to the South China Block and relatively far from the blocks in Northwest China (e.g., the Qilian Mountains and Qaidam Basin, Kunlun Mountains, Tarim Basin and Tianshan Mountains) during the late Viséan–Serpukhovian.     

Significance of Claraia from the Late Permian of South Guizhou,China

Recently collectée material of two Claraia taxa, Claraia zhiyunica Yang et al, 2001 and Claraia sp. nov. from the Late Permian of South China, are described. Late Permian Claraia species are compared with those from the Early Triassic, and the survival of Claraia across the mass extinction period across the Permian- Triassic boundary (PTB) is discussed.     

Early Carboniferous Bivalvia from western Argentina

González, C. R., Early carboniferous Bivalvia from western Argentina. Alcheringa 18, 169–185. ISSN 0311-5518.

Lower Carboniferous deposits of western Argentina yield invertebrates and plant remains. In the Tournaisian epoch, a transgression from the ‘Pacific’ flooded the Rio Blanco Basin, forming a semi-restricted inland sea. Marine invertebrates of this age comprise the Malimanian fauna, which is based on the Protocanites-Rossirhynchus Assemblage. Eleven species of Bivalvia are here described Palaeoneilo subquadratum sp. nov., Malimania triangularis gen. et sp. nov., Phestia sp., Volsellina? sp. Posidoniella malimanensis sp. nov., Leptodesma? sp., Schizodus sp., Cypricardinia? sp., Edmondia? sp., Sanguinolites punillanus sp. nov., and Vacunella? sp. nov. They are accompanied by gastropods, trilobites, conularids and corals. The Malimanian fauna is regarded as a poorly to moderately varied assemblage that was influenced by some basinal restrictions. It was probably connected to faunas of Chile and Peru, and lived during a stage of mild climate before the beginning of the Late Palaeozoic ice age.     

Late Paleocene foraminiferal faunas with Chapmanina and Reticulophragmium from offshore southeastern Australia: a Tethyan influence

Samples from off southern New South Wales in approximately 1750 m water depth have yielded late Paleocene faunas of benthic and planktonic foraminifera and other invertebrates. Two faunal associations with a strong Tethyan influence, dominated respectively by Chapmanina conjuncta sp. nov. and Reticulophragmium naroomaensis sp. nov., are described; this is the first formal record of these genera from Australasia. The age of the host sediment is based on both benthic species and few planktonic foraminifera.

The environment at the time was warm temperate to subtropical at the southwestern extremity of a counter-clockwise Pacific Ocean gyre. The new occurrence extends the record of Palaeogene marine sediments and palaeoenvironments along the eastern margin of Australia and confirms earlier observations that conditions in the marine environment at that time were considerably warmer along the east coast of Australia than those along the southern margin; the biogeographic association was Tethyan. The circulation model may go some way to explaining coeval warmth along the east Antarctic coast.     

Tivertonia yarrolensis (maxwell) (chonetidina,brachiopoda) from the Permian of the Sydney Basin,Australia

Tivertonia yarrolensis from the Farley Formation of the Sydney Basin is illustrated for the first time. The age of the formation is discussed. The palaeogeographical distribution of Tivertonia provides an additional generic link within the Austrazean Province of the Gondwanan Realm.     

The Late Palaeozoic brachiopod genus Jakutoproductus Kashirtsev 1959 and the Jakutoproductus verchoyanicus Zone,northern Yukon Territory,Canada

Shi, Guang R., 1994:03:28. The Late Palaeozoic brachiopod genus Jakutoproductus Kashirtsev 1959 and the Jakutoproductus verchoyanicus Zone, northern Yukon Territory, Canada. Alcheringa 18, 103–120. ISBN 0311-5518.

The familial and subfamilial position, species composition, and geographic distribution of the Late Palaeozoic productid genus Jakutoproductus Kashirtsev 1959 are reviewed. Jakutoproductus is placed in the subfamily Plicatiferinae Muir-Wood & Cooper 1960 of the family Plicatiferidae. Eighteen described species from the Russian Arctic. Mongolia, northeast China, and northern Yukon Territory, Canada are assigned to Jakutoproductus. The Jakutoproductus verchoyanicus Zone of late Sakmarian to Artinskian age, most likely early Artinskian (Aktastinian), here established is based on material from the Jungle Creek Formation, northern Yukon Territory, Canada, and is correlated with the following horizons in Russia: the Osennin Horizon in the Verchoyan Mountains, the lower Munugudjak Horizon of the Kolyma-Omolon Massif, the Hipkhoshin Suite of east Zabaikal, the lower Bhang Horizon of Taimyr, and an unnamed sandstone-shale unit on the north island of Novaya Zemlya.     

Carboniferous postglacial faunas in the late Serpukhovian–Bashkirian interval of central-western Argentina

Cisterna, G.A., Sterren, A.F., López Gamundí, O. & Vergel, M.M., March 2017. Carboniferous postglacial faunas in the late Serpukhovian–Bashkirian interval of central-western Argentina. Alcheringa, ISSN 0311-5518.

Typical glacial–postglacial sequences associated with the Late Palaeozoic Ice Age (LPIA) are recognized in the Calingasta-Uspallata Basin, central-western Argentina, particularly in the Hoyada Verde and El Paso formations (late Serpukhovian–Bashkirian) at Barreal Hill (San Juan province). Brachiopods and bivalves accompanied by gastropods, conulariids, nautiloids, corals and ostracods constitute the marine assemblages of the El Paso Formation. They are assigned to the Aseptella–Tuberculatella/Rhipidomella–Micraphelia (AT/RM) fauna, characterized by two fossil assemblages: Aseptella–Tuberculatella, identified in the lower fossiliferous interval, and Rhipidomella–Micraphelia in the upper. The development of the different invertebrate assemblages within the El Paso Formation, and their relationship with coeval suite in the Hoyada Verde Formation, can be explained by a complex array of abiotic factors (substrate stability, turbidity, nutrient availability, variation in oxygen levels, poor circulation and salinity variations in the water column) that were directly related to glacial retreat dynamics and coastal configuration. A restricted palaeofjord setting is proposed for the depositional environment of the El Paso Formation in contrast to an exposed open marine coast with a gently sloping shelf for the Hoyada Verde Formation. The study of the postglacial fauna of the El Paso Formation and its relationship with the Levipustula fauna in the Calingasta-Uspallata Basin, help determine the main controls on the distribution of the postglacial faunas in other late Palaeozoic South American basins, such as the Tepuel Genoa Basin in Patagonia and the Tarija Basin in Bolivia.

Gabriela A. Cisterna [gabrielacisterna@conicet.gov.ar], CONICET-UNLAR, Av. Dr. Luis M. de la Fuente s/n, La Rioja, 5300, Argentina; Andrea F. Sterren [asterren@unc.edu.ar], CICTERRA (CONICET – Universidad Nacional de Córdoba), Av. Vélez Sarsfield 1611, X5016GCA, Córdoba, Argentina; Oscar López Gamundí [orlg2003@yahoo.com], P1C Consultants 1121 Banks Street, Houston, TX 77006, USA; María del Milagro Vergel [vergelmar@tucbbs.com.ar], CONICET – INSUGEO – Facultad de Ciencias Naturales e Instituo Miguel Lillo (UNT), Miguel Lillo 205, 4000, San Miguel de Tucumán, Argentina.     

Ngapakaldia bonythoni (Marsupialia,Diprotodontidae): new material from Riversleigh,northwestern Queensland,and a reassessment of the genus Bematherium

A latest Permian (late Changhsingian) radiolarian fauna is recorded from the upper Talung Formation, Hushan, Nanjing, Jiangsu Province, South China. This fauna includes 24 species belonging to 16 genera; new species are Albaillella hushanensis, Copicyntroides stellatus and Trilonche crassus. The presence of the radiolarian fauna and its taxonomic composition reveal that the Eastern Qinling-Dabie deep sea, which was located along the northern margin of the northeastern Yangtze Basin, persisted at least until the end of the Palaeozoic and that the collision between the North China and South China plates had not occurred by the end of the Permian.     

A new crustacean clam shrimp (Spinicaudata: Eosestheriidae) from the Upper Triassic of Argentina and its importance for ‘conchostracan’ taxonomy

A new species of a fossil crustacean clam shrimp (Spinicaudata: Eosestheriidae) Menucoestheria wichmanni is described from the lower Upper Triassic Vera Formation (Los Menucos Complex) in Río Negro Province, southern Argentina. This discovery represents the first record of this family in the Triassic of Argentina and the southernmost record of South American Triassic ‘conchostracans’ (Spinicaudata). The new species shows close affinities with Middle Jurassic faunas from Antarctica and offers important data on the taxonomy (notably the use of ornamentation characters), palaeobiogeography (as South America hosts the oldest-known fossils of this family) and evolution of the Gondwanan faunas. Other South American eosestheriid species are tentatively recognized. Menucoestheria is hypothesized to be the ancestral form of the Triassic–Jurassic Gondwanan eosestheriids. Relationships between European and Gondwanan eosestheriids remain unresolved.     

Ordovician (Bendigonian,Darriwilian to Gisbornian) faunas from the northern Molong Volcanic Belt of central New South Wales

Diverse shallow water assemblages dominated by brachiopods, molluscs, sponges and stromatoporoids, and a tabulate coral, in the Wahringa Limestone Member (Darriwilian-Gisbornian), and Yuranigh Limestone Member (Gisbornian, or early Late Ordovician) of the Fairbridge Volcanics, are documented from the northern Molong Volcanic Belt in central N.S.W. New species described include Billingsaria spissa, Shlyginia printhiensis and Sowerbyites? wahringaensis. Elements of the Wahringa Limestone Member assemblage such as Labechia banksi, Labechiella regularis, and Maclurites cf. M. florentinensis are biogeographically significant in displaying strong similarities with contemporaneous Tasmanian faunas. The brachiopods Ishimia and Shlyginia from the Yuranigh Limestone Member are recognised for the first time outside Kazakhstan and Sibumasu. The presence of the brachiopod Anoptambonites in allochthonous limestone breccia within the lower Fairbridge Volcanics provides evidence of a regionally significant hiatus of 10–15 Ma duration separating this unit from the underlying Hensleigh Siltstone, of Early Ordovician (Bendigonian) age. The sponge Archaeoscyphia?, from allochthonous limestones in the latter formation, is the oldest macrofossil yet described from the Lachlan Fold Belt in central N.S.W.     

Chonetoidea (Brachiopoda) from the Lopingian (Late Permian) of South China

Ten species of the superfamily Chonetoidea from the Lopingian (Late Permian) of South China are described or revised. A review of all recorded Chonetoidea species from the Lopingian (Late Permian) of South China indicates that some 22 species of five genera can be recognised. Species of Tethyochonetes and Neochonetes are characteristic in the lithofacies dominated by mudstone, siltstone or siliceous rocks in the Lopingian and some argillaceous limestone and clay rock facies near the Permian-Triassic boundary. New taxa are Neochoneles (Zhongyingia) subgen. nov., Neochonetes (Huangichonetes) subgen. nov. and Tethyochonetes flatus sp. nov.     

Early Ordovician conodonts from Zhejiang Province,southeast China and their biostratigraphic and palaeobiogeographic implications

Forty-two conodont species are documented from the Liuxia, Shijiatou and Jingshan formations in Zhejiang Province of southeast China, located palaeogeographically on the Jiangnan Slope offshore to the Yangtze Platform. From these faunas, eight successive conodont biozones of Tremadocian to middle Floian (Early Ordovician) age are recognized, including the Cordylodus lindstromi Biozone, Cordylodus angulatus Biozone, Chosonodina herfurthi Biozone, Paltodus deltifer Biozone, Paroistodus proteus Biozone, Triangulodus bifidus Biozone, Serratognathus diversus Biozone and Prioniodus elegans Biozone. Several zonal index species of the Baltoscandian succession—Paltodus deltifer, Paroistodus proteus and Prioniodus elegans—are described and illustrated in detail for the first time from South China. Co-occurrence of P. proteus and Serratognathus bilobatus in several samples below the appearance of P. elegans also confirms correlation of the S. diversus Biozone (basal Floian) with the upper P. proteus Zone of the Baltoscandian succession. These Zhejiang faunas are dominated by pandemic forms, and are similar to those of the Open-Sea Realm elsewhere, inhabiting deep, offshore environments.