Lower Devonian ischnacanthid fish (Gnathostomata: Acanthodii) from the Taemas Limestone,Lake Burrinjuck,New South Wales

Dentigerous jawbones of ischnacanthid acanthodians from the Lower Devonian Currajong Limestone Member and the lower part of the Bloomfield Limestone Member of the Taemas Limestone (Murrumbidgee Group: early Emsian) in the Murrumbidgee valley at Good Hope, Taemas district, and the Goodradigbee valley at Wee Jasper are described. Three taxa of Taemasacanthus are recognized: T. sp. cf. erroli Long, 1986; T. narrengullenensis sp. nov. and T. cooradigbeensis sp. nov. Fragmentary fin spine remains and scales are also described.     

Acanthodian fish remains from the Lower Devonian Cavan Bluff Limestone (Murrumbidgee Group), Taemas district,New South Wales

A well-preserved acanthodian fish fauna from the Lower Devonian (early Emsian) Cavan Bluff Limestone (Murrumbidgee Group), Taemas, Yass district, New South Wales, consists of dentigerous jawbones, fin spines and scales. Four taxa belonging to the Order Ischnacanthida are recognized including Taemasacanthus erroli Long, 1986 and newly described genera and species Cavanacanthus warrooensis gen. et sp. nov., Cambaracanthus goodhopensis gen. et sp. nov. and Taemasacanthus porca sp. nov. An amended diagnosis is provided for T. erroli. The jawbone of C. warrooensis gen. et sp. nov. is of moderate size and bears a single row of teeth with a circular parabasal section. The jawbone extends in an anterior direction beyond the foremost tooth. C. goodhopensis gen. et sp. nov. is a small to moderate sized jawbone bearing two teeth rows separated by a longitudinal ridge. The teeth of the mesial tooth row, the main tooth row, are circular in parabasal section. The lateral tooth row is weakly developed, bearing one or two incipient teeth. T. porca sp. nov. is represented by a small curved jawbone (mesially concave) bearing two teeth rows separated by a prominent longitudinal ridge. The teeth of both tooth rows have a circular parabasal section. The anterior extension of this ridge beyond the foremost tooth represents approximately one quarter the length of the jawbone. These fishes inhabited a Lower Devonian carbonate platform consisting of patch reefs built upon a muddy substrate on a low energy shallow marine shelf which was subjected to frequent storm surges.     

A new Late Devonian acanthodian fish from the Hunter Formation near Grenfell,New South Wales

The ischnacanthid acanthodian Grenfellacanthus zerinae gen. et sp. nov. is described on the basis of two large jaw bones from the Late Devonian (late Famennian) Hunter Formation, near Grenfell, N.S.W. The new species is the youngest known ischnacanthid, and the largest ischnacanthid from Gondwana. As for many ischnacanthids, the structure of the jaws and teeth indicate that Grenfellacanthus was probably an ambush predator.     

Late Silurian pentameride brachiopods from Yass and Molong,New South Wales

The five known species of pentameride brachiopods from the Yass Syncline Ludlow (LateSilurian) succession, belonging to the superfamilies Pentameroidea, Gypiduloidea and Clorindoidea, are fully revised; no new species are recognised. The pentameroids Conchidium sp. cf. hospes and Aliconchidium yassi are confined to the Bowspring Limestone Member (Silverdale Formation). The gypiduloid Ascanigypa glabra and externally homeomorphic clorindoids Barrandina wilkinsoni and Clorinda minor replace them in the overlying Barrandella Shale Member, the last two extending into the Yarwood Siltstone Member (Black Bog Shale). Clorinda minor is also possibly present in the Rainbow Hill Member (Rosebank Shale). All except C. minor are uncommon to rare components of the Yass brachiopod fauna. Clorinda molongensis, a species of uncertain mid- to late Silurian age from the Molong Limestone, is also revised. Aliconchidium and Barrandina are known only from Yass, whereas Clorinda is cosmopolitan. Conchidium alsois widespread, but C. hospes is a species from the Prague Basin probably also known from the Urals and the Tien Shan. Ascanigypa is another Prague Basin taxon, recently recognised in Arctic Canada.     

New data on the structure of the Early Devonian lungfish,Dipnorhynchus

A skull of a young specimen of Dipnorhynchus sussmilchi (Etheridge Jr), from the base of the Bloomfield Limestone Member (mid-Emsian, probably perbonus Zone) at Taemas, has both sides of the skull roof preserved, but the cheeks are missing. The posterior end of the endocranium, up to its dorsal extremity, provides new data on the endolymphatic sacs, and the attachment of the epaxial muscles. The endolymphatic region is compared with that of Chirodipterus australis. The palate is not fully formed and the marginal ridges are not fully developed. The fine structures around the dermopalatines and the pterygoids are preserved, indicating the manner in which the plates grew. The position of the parasphenoid is clearly outlined against the pterygoids, and confirms the view that this primitive genus of dipnoans did not have the pterygoids divided by the parasphenoid. All these features provide new information and comparison with Dipterus valenciennesi, the next oldest genus on which the same regions are preserved. In addition, a new mandible from a horizon low in the Cavan Bluff Formation is well preserved and provides information on the external bones of Dipnorhynchus.     

Macropalaeontology of the Trochocyathus-Trematotrochus band (Paleocene/Eocene boundary), Dilwyn Formation,Otway Basin,Victoria

Two new brachythoracids are described from the Taemas Limestone. Arenipiscis westolli gen. et sp. nov. possesses a skull-roof with deeply embayed posterior and elongate anterolateral margins, a large fused rostropineal plate, and orbits anteriorly placed and deeply notching the skull. The visceral skull-roof surface is developed with strong relief. The trunk-shield is poorly known, but the median dorsal plate probably had a prominent carinal process. Errolosteus goodradigbeensis gen. et sp. nov. is known only by an incomplete skull-roof, and anterior lateral and posterior ventrolateral plates. It has a highly distinctive ridged ornament. The affinities of these forms are considered in a discussion of brachythoracid interrelationships, and it is concluded that they are interchangeable on a cladogram with Buchanosteus. Other brachythoracids from the fauna, recently described by White, are considered, and it is suggested that Goodradigbeeon is phyletically primitive and Taemasosteus phyletically advanced relative to Buchanosteus. Schemes of brachythoracid interrelationships recently proposed by Denison and by Miles &; Dennis are compared, and a new scheme is proposed.     

Ordovician (Bendigonian,Darriwilian to Gisbornian) faunas from the northern Molong Volcanic Belt of central New South Wales

Diverse shallow water assemblages dominated by brachiopods, molluscs, sponges and stromatoporoids, and a tabulate coral, in the Wahringa Limestone Member (Darriwilian-Gisbornian), and Yuranigh Limestone Member (Gisbornian, or early Late Ordovician) of the Fairbridge Volcanics, are documented from the northern Molong Volcanic Belt in central N.S.W. New species described include Billingsaria spissa, Shlyginia printhiensis and Sowerbyites? wahringaensis. Elements of the Wahringa Limestone Member assemblage such as Labechia banksi, Labechiella regularis, and Maclurites cf. M. florentinensis are biogeographically significant in displaying strong similarities with contemporaneous Tasmanian faunas. The brachiopods Ishimia and Shlyginia from the Yuranigh Limestone Member are recognised for the first time outside Kazakhstan and Sibumasu. The presence of the brachiopod Anoptambonites in allochthonous limestone breccia within the lower Fairbridge Volcanics provides evidence of a regionally significant hiatus of 10–15 Ma duration separating this unit from the underlying Hensleigh Siltstone, of Early Ordovician (Bendigonian) age. The sponge Archaeoscyphia?, from allochthonous limestones in the latter formation, is the oldest macrofossil yet described from the Lachlan Fold Belt in central N.S.W.     

Campanile (Mollusca: Gastropoda): a new record from the early Miocene of Victoria,Australia

Campanile rupicolum sp. nov. is described from the early Miocene Upper Maude Limestone Member of Maude, Victoria. The genus has not hitherto been recorded from Victoria, a part of the Southeast Australian biogeographic Province, and was thought to be characteristic of the Austral-Indopacific Province. This is the oldest occurrence of the genus so far recorded from Australia.     

Archaeocyaths from Yorke Peninsula,South Australia and archaeocyathan Early Cambrian zonation

Zhuravlev, A. Yu., & Gravestock, D. I., 1994:03:28. Archaeayaths from Yorke Peninsula, South Australia and archaeocyathan Early Cambrian zonation. Alcheringa 18, 1–54. ISSN 0311-5518.

Two assemblages of archaeocyaths are documented from Lower Cambrian outcrops and drillholes on Yorke Peninsula. South Australia. The older assemblage (11 species) occurs in the uppermost Kulpara Formation and conformably overlying basal Parara Limestone, and is equivalent to Lower Faunal Assemblage II in the Flinders Ranges. The younger assemblage (28 species plus Acanthhcyathus and Rodiocyathus) occurs in the Koolywurtie Member near the top of the Parara Limestone. Equivalent taxa are widespread in the Flinders Ranges, western New South Wales and Antarctica. Archaeocyathan distribution is now sufficiently well known to propose three assemblage zones and two informal assemblages for regional correlation. No new taxa have been added, but Erugatocyathus scutatus (Hill) and Pycnoidocyathus latiloculatus (Hill), hitherto known only from Antarctica, are found in the upper assemblage on Yorke Peninsula. Irregular archaeocyathan systematics are discussed, the ontogeny of Archaeopharetra irregularis (Tylor) clarified, and Kruseicnema Debrenne. Gravestock & Zhuravlev, represented by K. gracilis (Gordon), is fully described.     

Editorial

Similarities in late Middle Cambrian trilobite, inarticulate brachiopod and molluscan faunas are apparent between Bornholm (Denmark) and New Zealand. Tuarangia gravgaerdensis sp. nov. is described from the late Middle Cambrian Andrarum Limestone of Bornholm, Denmark. The Andrarum Limestone is stratigraphically correlated with the Tasman Formation, New Zealand, from where the genus Tuarangia was first described. A single specimen, referred to Tuarangia, has been found in an erratic boulder from NW Poland. The boulder is estimated to be of early Late Cambrian age. The initial taxonomic assignment of Tuarangia to the Bivalvia, Subclass Pteriomorphia, is upheld. The distance between Bornholm and New Zealand, the position at opposite hemispheres, and the palaeogeography in general clearly indicate isocommunities with restricted possibility of exchange of the gene pool at species level.     

Biostratigraphic potential of Middle Cambrian hyoliths from the eastern Georgina Basin

Fourteen hyolith taxa are documented from the Middle Cambrian (Templetonian to Floran) of the eastern (Queensland) portion of the Georgina Basin, Australia, as a contribution toward a prospective Australian Cambrian hyolith biozonation. The described fauna is from the Beetle Creek Formation (including Monastery Creek Phosphorite Member) and Gowers Formation. Additionally, the enigmatic Cupittheca and some indeterminate hyoliths are figured to illustrate aspects of hyolith morphology. Guduguwan hardmani, widespread in Ordian-early Templetonian strata of northern Australia, is here recorded from the early Templetonian of the eastern Georgina Basin. A new family Gakidae is established for sulcavitide hyolithomorphs with a conch of pentagonally tabernacular transverse section, to include Gaka, Kalkatungu gen. nov. and possibly Dorsolinevitus. New genera are the hyolithid Yalarrnga mara gen. et sp. nov., sulcavitid Kulangarra kutjurru gen. et sp. nov., gakid Kalkatungu murlu gen. et sp. nov. and angusticornid Yuku tjurtu gen. et sp. nov.; new species are Loculitheca kunka sp. nov., Carinolithes tjikilirri sp. nov., ?Sololites kankari sp. nov., ?Shandongolithes thakal sp. nov., ?Gerkella thuka sp. nov. and ?Yacutolituus rakatju sp. nov. Taxa in open nomenclature are Foersteotheca cf. dubecensis, ?Holmitheca sp. and ?Dorsojugatus sp. On present knowledge, the potential for an Australian Cambrian hyolith biozonation is limited in the Early Cambrian, but for the Middle Cambrian, G. hardmani is a widespread Ordian-early Templetonian indicator, while hyolith distribution in the Monastery Creek Phosphorite Member suggests a faunal turnover at or about the incoming of Acidusus atavus which may provide a basis for biozonation in the Floran stage.     

A new late Oligocene to early Miocene species o Apatopygus (Echinoidea: Cassiduloida) from the Maltese Islands

The occurrence of Apatopygus gaudensis sp. nov. in the upper part of the late Oligocene (Chattian) Lower Coralline Limestone division, and the lower part of the early Miocene (probably late Aquitanian) Lower Globigerina Limestone division of the Maltese Islands (Central Mediterranean), is the first confirmed record of Apatopygus outside the Australia-New Zealand region. Apatotygus vincentinus (Tate, 1891) from the middle to late Eocene of southern Australia is the earliest known occurrence of the genus, possibly suggesting that Apatotygus may have evolved earlier in the Australasian region, or merely that it is yet to be discovered in pre-Oligocene strata in the Mediterranean area.     

Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America

Camilleri, T.A., Warne, M.T., Holloway, D.J. & Weldon, E.A., 10 May 2019. Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America. Alcheringa XXX, X–X. ISSN 0311-5518.

Known occurrences of the ostracod genus Velibeyrichia are restricted to a number of Silurian to Lower Devonian geological strata in North America: the McKenzie Member of the Mifflintown Formation of Maryland and West Virginia; the Tonoloway Limestone of Maryland, West Virginia, Virginia and Pennsylvania; the Bloomsburg Formation of Maryland, Virginia and Pennsylvania; the Manlius Limestone of New York; and the Decker Limestone of New Jersey and New York. The genus includes six species: V. moodeyi (type species), V. mesleri, V. paucigranulosa, V. reticulosaccula, V. tonolowayensis and V. tricornia. The diagnostic combination of characters for this genus are: distinct deflection of the velum where it crosses the crumina in heteromorphs (adult female specimens), dorsal nodes on lobes L1 and L3, sexual dimorphism of the velum, and in tecnomorph specimens, either a shallow sulcus on lobe L3 or a zygal ridge (in adult tecnomorph specimens) extending from lobe L2 to lobe L3. The presence of one or the other of the latter two characters defines two distinct species groups.

Tamara T.A. Camilleri* [tamara.camilleri@deakin.edu.au], Mark T. Warne* [mark.warne@deakin.edu.au] and Elizabeth A. Weldon [l.weldon@deakin.edu.au], Deakin University, Geelong, School of Life and Environmental Sciences & Centre for Integrative Ecology (Melbourne Campus), 221 Burwood Highway, Burwood, Victoria 3125, Australia; David J. Holloway [dhollow@museum.vic.gov.au], Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia. *Also affiliated with: Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia.     

Bat cave and Hobbit hole,microbats of Liang Bua (Flores,Indonesia)

Liang Bua is a limestone cave on the island of Flores, Indonesia. Palaeontological and archaeological excavations at this site have unearthed one of the best insular records of fossil bats to date. The assemblage is numerically dominated by the horseshoe bat species Rhinolophus euryotis, followed by the leafnosed bat Hipposideros diadema. In addition to these species, remains of Rhinolophus simplex, Hipposideros sp. cf. H. sumbae, Murina sp. aff. M. florium, Taphozous sp., Kerivoula sp. and Miniopterus sp. have also been recovered. Two taxa found in Liang Bua, Rhinolophus euryotis and Miniopterus sp., are not apparent in the modern bat fauna of Flores. Some specimens, found during the more recent excavations, are known from specific depths. Two age-groups are represented. The stratigraphically oldest is 74–61 ka and includes Murina sp. aff. M. florium and Rhinolophus euryotis. Conversely, the younger, Holocene, assemblage is much more diverse. Despite the relatively small amount of microbat material collected thus far, it displays a higher diversity than the rodents from the site. This is presumably related to the better dispersal ability of bats relative to other, non-volant mammals.     

Late Triassic megaspores from the Amery Group,Prince Charles Mountains,East Antarctica

Cantrill, D. J., & Drinnan, A. N., 1994:03:28. Late Triassic megaspores from the Amery Group, Prince Charles Mountains, East Antarctica. Alcheringa 18, 71–78. ISSN 0311-5518.

Megaspores referable to the genera Cabochonicus Batten & Ferguson 1987 and Minerisporites Potonié 1956 are a common component of the palaeoflora recovered from the Jetty Member within the Flagstone Bench Formation of the Amery Group. The known ranges of these two genera, in conjunction with the macrofloral remains, suggest a Late Triassic age. Two new species, Cabochonicus sinuosus and Minerisporites triangulatus, are described. Scanning electron microscopic examination of the spores indicates that standard palynological treatment of megaspores can result in sculptural degradation, possibly leading to incorrect generic assignment.     

A second species of the possum Durudawiri (Marsupialia: Miralinidae) from the early Miocene of Riversleigh,northwestern Queensland

Durudawiri anfractus sp. nov. (Marsupialia: Miralinidae) is described from Riversleigh. This, the second described species of the genus, is very similar in morphology to, but much larger than, D. inusitatus. Durudawiri anfractus and D. inusitatus are found at similar sites, all early Miocene. The Miralinidae remains one of the most time-restricted families of marsupial, being found so far in only the late Oligocene and early Miocene.     

Late Silurian vertebrate microfossils from the Carnarvon Basin,Western Australia

Burrow, C.J., Turner, S., Trinajstic, K. &; Young, G.C., 27 February 2019. Late Silurian vertebrate microfossils from the Carnarvon Basin, Western Australia. Alcheringa 43, 204–219. ISSN 0311-5518.

A core sample from the offshore Pendock 1A well, Carnarvon Basin, Western Australia yielded microvertebrate residues at an horizon in the lower part of the Hamelin Formation, dated as late Silurian, ? Ludlow, based on associated conodonts. The fish fauna comprises loganelliiform thelodont scales, the ? stem gnathostome Aberrosquama occidens nov. gen. et sp., the acanthodian Nostolepis sp. aff. N. alta, and the ? stem osteichthyan Andreolepis sp. aff. A. petri. Because of the paucity of the material, and some differences between the Pendock scales and those of established species, a precise age can not be confirmed; however, the composition of the fauna at generic level most closely resembles that of late Silurian (Ludlow) assemblages from northern Eurasia.

Carole J. Burrow* [carole.burrow@gmail.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Susan Turner [palaeoich@yahoo.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Kate Trinajstic [k.trinajstic@curtin.edu.au], School of Molecular and Life Sciences, Curtin University, Perth, WA 6102, Australia; Gavin C. Young [gavin.young@anu.edu.au], Research School of Physics and Engineering, Australian National University, Canberra, ACT 2000, Australia.     

Microflora of the H.Y.C. Pyritic Shale Member of the Barney Creek Formation (McArthur Group), middle Proterozoic of northern Australia

The H.Y.C. Pyritic Shale Member of the Barney Creek Formation (ca 1 500 my old; northern Australia) contains several stratiform base metal sulfide deposits of economic significance. Black cherts within these mineral deposits preserve a diverse assemblage of bacterial and algal microfossils. The assemblage differs from most other Precambrian biotas so far described in that it was deposited in deep water, it is not associated with stromatolites or algal mats, and it is dominated by filamentous bacteria, most of which are pyritized. Analysis of the assemblage suggests that the depth of the depositional basin exceeded that of the photic zone, that the bacteria inhabited the basin floor where they maintained anoxic conditions through heterotrophic degradation of detrital organic matter, and that the algae inhabited overlying near surface waters. Most of the algal fossils have been assigned to the Cyanophyta, although two of the described species are potentially referable to the eukaryotic green or red algae. Differences between this assemblage and other biotas described from the McArthur Group suggest that a workable system of biostratigraphic zonation for the Group is feasible.

Fossils in the H.Y.C. assemblage are here referred to 21 species and 16 genera, of which 14 species and 6 genera are new. The new taxa are: Bacteria, Biocatenoides incrustata sp. nov., B. pertenuis sp. nov., Ramacia carpentariana gen. et sp. nov., Coleobacter primus gen. et sp. nov., Ferrimonilis variabile gen. et sp. nov.; Chroococcales (Cyanophyta), Nanococcus vulgaris gen. et sp. nov., Bisacculoides tabeoviscus gen. et sp. nov., B. vacua gen. et sp. nov., B. grandis gen. et sp. nov.; Nostocales (Cyanophyta), Oscillatoriopsis schopfii sp. nov., Cyanonema inflatum sp. nov., C. minor sp. nov.; Incertae sedis, Clonophycus elegans gen. et sp. nov., Globophycus minor sp. nov. In addition, the new combination Gunflintia septata (Schopf) is proposed.     

The oldest Ordovician stromatoporoids from Australia

The oldest known Australian Ordovician stromatoporoids are described from the Cashions Creek Limestone (formerly the Maclurites-Girvanella horizon) and correlatives of the Gordon Subgroup in Tasmania. The Cashions Creek Limestone and equivalents are correlated approximately with the North American Chazyan (Middle Ordovician). Representatives of Labechia, Stratodictyon and Stromatocerium are recorded from localities in the Mole Creek area, from the Florentine Valley and from Melrose. Three new species, Labechia banksi, Stratodictyon vetus and Stromatocerium bigsbyi are described. L. banksi comes from a slightly higher horizon in the succession at Mole Creek where it occurs in association with the earliest corals (Lichenaria). The distribution of the earliest Ordovician stromatoporoids — those appearing in the North American and Tasmanian successions — is reviewed, together with a discussion of their possible origins and interrelationships.