Trilobite faunas and palaeoenvironmental setting of the Silurian (early Ludlow) Melbourne Formation,central Victoria

Trilobites are common faunal elements in the Melbourne Formation, a unit of early Ludlow (upper nilssoni Biozone) age, which crops out extensively in the Darraweit Guim Province of the Melbourne Zone, central Victoria. New diagnoses are given for species previously described, including Maurotarion euryceps (McCoy, 1876; = Cyphaspis spryi Gregory, 1901), Raphiophorus jikaensis (Chapman, 1912; = Ampyx yarraensis Chapman, 1912), Cromus simpliciculus (Talent, 1964), Cromus spryi (Chapman, 1912), Sthenarocalymene kilmorensis (Gill, 1945; = Gravicalymene hetera Gill, 1945) and Trimerus harrisoni (McCoy, 1876). A new phacopid genus, Orygmatos is described, represented by the species O. yanyeani gen. et sp. nov. Other species newly described include Cromus melbournensis sp. nov., Arcticalymene australis sp. nov., “Ananaspis” woiwurrungi sp. nov. and Kettneraspis hollowayi sp. nov.

Species composition of the trilobite fauna varies spatially, and a number of distinct assemblages can be defined. Abundant trilobite moult configurations are conclusive for interpretation of the benthic fauna as autochthonous, inferring depth estimations based on benthic community distribution to be valid. A depth-related succession of communities is recognised and indicate the Melbourne Formation was deposited at relatively shallow depths on a broad, eastwardly deepening shelf, with deposition dominated by storm processes. The palaeoenvironment comprised a BA-1 community including the Arcticalymene australis trilobite assemblage, restricted to very shallow depths (～20 m) on the SW coastal margin of the shelf and preserved in proximal tempestite lithologies; and a BA-5 community group containing three distinct trilobite assemblages dominated by species of Cromus and a deeper water fauna, preserved in distal tempestite lithologies and ranging widely over the shelf at depths in the range of maximum storm wave base (～50 – 80 m).     

The Pleistocene of Rabat (Morocco): Mollusks,Coastal Environments and Human Behavior

The Rabat-Témara region of the Moroccan Atlantic coast reveals a succession of Quaternary palaeobeaches. This coastal area is dotted with numerous prehistoric caves. The study of the Upper Pleistocene coastal landscape associated with these caves is of paramount importance in the knowledge of human population subsistence. During the Upper Pleistocene, the ocean level changes drastically influenced the coastal geomorphology as well as the fauna assemblages. The chrono-lithostratographical analysis of the coastal sedimentary formations allows the distinction of three sequences rich in marine fauna. These sequences date from MIS 11 to MIS 5. The identification of malacofauna species from these deposits revealed 39 species, along with Bryozoans, Crustaceans, and Echinoids. These assemblages show a constant fauna cortege highlighted by the dominance of the amphi-Atlantic species Stramonita haemastoma. This species shows an increase in the number of specimens in the uppermost part of marine deposits, probably in relation with a climate warming in the MIS 5. This fauna of both intertidal rocky substrates and sandy substratum indicates environmental conditions close to the present-day Rabat-Témara coastline. As in other coastal locations of Africa from MIS 5, the Middle Stone Age Homo sapiens population benefitted from a littoral environment rich in coastal resources. Comparison between thanatocenoses and archaeological records allows us to identify both species available for Middle Stone Age population and those preferred for human use.     

Acanthodian,onychodontid and osteolepidid fish from the middle-upper Taemas Limestone (Early Devonian), Lake Burrinjuck,New South Wales

A meagre ischnacanthid acanthodian fauna, known from fragmentary dentigerous jawbones from Lake Burrinjuck, Yass district, New South Wales, consists of three taxa: Taemasacanthus narrengullenensis, Taemasacanthus cooradigbeensis and an indeterminate ischnacanthid. The fossils are from three Early Devonian (Emsian) units at Taemas: the Bloomfield Limestone Member, the Currajong Limestone Member and the Warroo Limestone Member, which constitute the middle-upper parts of the Taemas Limestone (Murrumbidgee Group); and from Unit 6 of the ‘Upper Reef formation’ at Wee Jasper. Whereas ischnacanthid acanthodian remains are increasingly rarer in the higher levels of the Taemas Limestone, remains of onychodontid and osteolepidid fishes are relatively abundant; Onychodus yassensis sp. nov., Onychodus sp. and isolated osteolepidid teeth are locally common.     

Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.     

Llanvirn-early Caradoc trilobite biofacies of western Hubei and Hunan,China

On the basis of representative collections made from seven measured sections in western Hubei and Hunan, four early Llanvirn and five late Llanvirn-early Caradoc trilobite biofacies are distinguished along a bathymetric gradient from shallow to deep outer shelf. The diversity and relative abundance of mesopelagic cyclopygids gradually increased southwards or seawards. According to the facies association exhibited in the region, the upper limit of living depth for different forms is inferred as follows: 1) more than 70 m (M. (Microparia)); 2) less than 100 m (M. (Quadratapyge), Pricyclopyge, Aspidaeglina); 3) at approximately 100 m (Cyclopyge); and 4) more than 100 m (Symphysops, Psilacella). Benthic forms were absent in the deep outer-shelf basin because of stagnant conditions. In shallower sites of the outer shelf, genera of nileids in the carbonate facies and raphiophorids in the clastic facies were relatively abundant, but some endemic forms, such as the asaphid Birmanites and remopleuridid Hexacopyge, became dominant during the late Llanvirn-early Caradoc. Rorringtoniids are mainly associated with deeper outer-shelf facies. Panderia is common in an early Llanvirn nileid fauna at Yichongqiao, Cili area, and may well indicate development of shelf carbonate buildups. A transgressive event is suggested by the sudden appearance of rich Cyclopyge in the early Caradoc at Jiuxi, Changde.     

Late Ordovician linguliformean microbrachiopods from north-central Kazakhstan

An assemblage of well-preserved Devonian trilobites is described from limestones in Satun Province, southern Thailand. The biogeographic implications are briefly discussed. The fauna, which is of Early Devonian age, probably Emsian, includes Decoroproetus, Cornuproetus, Platyscutellum and two species of Reedops, one of which is new. No similar fauna has been described before from the Shan-Thai block. The fauna is a typical Hercynian assemblage with close specific comparisons with Turkey, Morocco and Bohemia.     

Oligocene-Miocene calcareous nannofossils from the Torquay Basin,Victoria, Australia

Oligocene-Miocene rocks of the Torquay Basin contain at least 55 different species of calcareous nannofossils. These species allow zonation of an offshore subsurface section (Nerita No. 1 Well) and two onshore sections (near Torquay). The Angahook Formation and the base of the Jan Juc Formation in the Nerita No. 1 Well belong to Martini's (1971) NP 22 calcareous nannofossil Zone (early Oligocene). The NP 23 Zone could not be identified with certainty, but Zones NP 24/25 (late Oligocene) to NN 2 (early Miocene) were recognized in the Jan Juc Formation. The Oligocene-Miocene boundary is present between 122 m and 135 m below the sea floor at this site. The Puebla Formation, exposed at the sea floor, contains NN 5 spanning the early-middle Miocene boundary.

Onshore along the Torquay coast, NP 24/25 (late Oligocene) is present in the Angahook Formation; the Jan Juc Formation at Bird Rock also contains NP 24/25, except the uppermost 2.5 m which belongs to NN 1. The NN 2 Zone begins at the contact between the Jan Juc and Puebla Formations and continues upward for at least 3 m. After a covered interval, NN 4 was identified 10 m above the contact. The Oligocene-Miocene boundary is placed at the base of the hard band capping Bird Rock in the type section of the Jan Juc Formation.

Nannofossil assemblages in the Torquay sections suggest deposition in cool, shallow continental shelf waters. The assemblages in Nerita No. 1 Well also suggest cool shelf waters, although deeper than at Torquay.     

History,Chronology and Techno-Typology of the Upper Paleolithic Sequence in the Shuidonggou Area,Northern China

The timing and behavioral markers of the Upper Paleolithic in different parts of the world are of great importance to research on modern human dispersals. The pattern of behavioral developments in the Upper Paleolithic in northern China differs in important ways from the patterns observed in West Eurasia, Africa, and South Asia. Shuidonggou (SDG), a cluster of Paleolithic sites in northern China, contains several of the most important Upper Paleolithic sites in the region. Various localities yield evidence of three major cultural components dated by ¹⁴C, uranium-series, and optically stimulated luminescence (OSL) methods to between roughly 46 ka and 10 ka. The oldest component, blade assemblages with western Eurasian early Upper Paleolithic characteristics, appears to be intrusive from Siberia and/or Mongolia, beginning at least 41 ka (e.g., SDG 1 and SDG 9). Advanced core and flake assemblages may mark the appearance of an indigenous Late Paleolithic of North China beginning at around 33 ka (e.g., SDG 2 and SDG 8). Finally, around 10.5 ka, microblade technology arrived in the area (SDG 12), although we are not sure of its origins at present. Other typical Upper Paleolithic cultural remains, such as bone tools and body decorations, have been found at various localities in the SDG area as well (e.g., ostrich eggshell beads from SDG 2, 7, and 8). Information from this cluster of occupations increases our understanding of cultural variability, adaptation, and demographic dynamics of modern humans in Late Pleistocene northern Asia.

     

A history of changing aesthetic values in the Yarra Valley landscape,Victoria

Aesthetic responses to landscape may be emotional, sensory, or experiential, or a combination; it is a difficult value to identify and assess. This paper considers changing perceptions of the aesthetic values of the Yarra Valley landscape using a heritage methodology that involves examining artistic, creative, and published sources accounting for a lengthy period of changing land use. Scenic quality mapping, where available, is used as a support, and extensive reference is made to Australian impressionist art. Artworks for Yarra Valley landscapes may be considered as being among the cultural heritage indicators of significance and there is a new aesthetic of land art in private venues for the visiting public. The paper also illustrates how aesthetic values have been more broadly recognised in the post‐World War II era and become the domain of planning authorities as well as artists. It is for regional planners to be aware of and maintain these aesthetic values in the landscape.     

Thelodont fish and conodonts from the Early Devonian of Reefton,New Zealand

Thelodont scales and associated fragments of conodont elements identified as Turinia sp. cf. Turinia australiensis and Ozarkodina sp. cf. Ozarkodina buchanensis respectively are described for the first time from the Early Devonian Murray Creek Formation of Reefton, New Zealand. The conodont and thelodont components of the fauna suggest a late Pragian to early Emsian age for the assemblage.     

Acanthodian fish remains from the Lower Devonian Cavan Bluff Limestone (Murrumbidgee Group), Taemas district,New South Wales

A well-preserved acanthodian fish fauna from the Lower Devonian (early Emsian) Cavan Bluff Limestone (Murrumbidgee Group), Taemas, Yass district, New South Wales, consists of dentigerous jawbones, fin spines and scales. Four taxa belonging to the Order Ischnacanthida are recognized including Taemasacanthus erroli Long, 1986 and newly described genera and species Cavanacanthus warrooensis gen. et sp. nov., Cambaracanthus goodhopensis gen. et sp. nov. and Taemasacanthus porca sp. nov. An amended diagnosis is provided for T. erroli. The jawbone of C. warrooensis gen. et sp. nov. is of moderate size and bears a single row of teeth with a circular parabasal section. The jawbone extends in an anterior direction beyond the foremost tooth. C. goodhopensis gen. et sp. nov. is a small to moderate sized jawbone bearing two teeth rows separated by a longitudinal ridge. The teeth of the mesial tooth row, the main tooth row, are circular in parabasal section. The lateral tooth row is weakly developed, bearing one or two incipient teeth. T. porca sp. nov. is represented by a small curved jawbone (mesially concave) bearing two teeth rows separated by a prominent longitudinal ridge. The teeth of both tooth rows have a circular parabasal section. The anterior extension of this ridge beyond the foremost tooth represents approximately one quarter the length of the jawbone. These fishes inhabited a Lower Devonian carbonate platform consisting of patch reefs built upon a muddy substrate on a low energy shallow marine shelf which was subjected to frequent storm surges.     

Pterygotid eurypterid chelicera from the Lower Devonian of Victoria

A single eurypterid (Arthropoda: Chelicerata) chelicera, assigned to Acutiramus sp. cf. A. bohemicus, is described from the Wilson Creek Shale, Turtons Creek inlier, north of Foster, Victoria, Australia. The specimen comprises the proximal portion of both rami. This pterygotid chelicera supports an Early Devonian (?Lochkovian) age for the stratum at this locality, by comparison with occurrences of A. bohemicus from the Czech Republic and closely related species in northern Gondwana.     

Specialized Early Upper Palaeolithic Hunters in Southwestern France?

Paul Mellars has long used cave and rockshelter ungulate faunal assemblages from southwestern France to argue that the early Upper Palaeolithic people of this region focused their hunting on reindeer (Rangifer tarandus), and that such specialized hunting distinguishes the Upper from the Middle Palaeolithic in at least this region. We examine this argument quantitatively, using a sample of 133 Mousterian, Châtelperronian, and Aurignacian ungulate assemblages. We show that only five Aurignacian assemblages, from three sites, stand out in terms of the degree to which their ungulate faunas are dominated by a single taxon. We also show that some Mousterian cave and rock shelter ungulate assemblages are more heavily dominated by large bovids than Aurignacian assemblages are dominated by reindeer, and that Mellars' argument is highly dependent on the exclusion of open sites from the analysis and on the numerical threshold he has selected to indicate hunting specialization.     

The Upper Paleolithic in Mediterranean Spain: A Review of Current Evidence

In spite of an active and sophisticated archaeological research program, the Paleolithic of the Iberian peninsula remains comparatively little known to English-speaking prehistorians, with the exception of Cantabrian Spain. The rich data set compiled by Spanish prehistorians and their colleagues over the past several decades stands to make a valuable and unique contribution to our understanding of the Pleistocene prehistory of Europe. We present a detailed overview of Upper Paleolithic chronology, sites, and assemblages for Mediterranean Spain, an area of over 1,600,000 km ² that extends from the French border to the Straits of Gibraltar. To interpret these data, we employ a regional perspective that emphasizes studies of paleoeconomy (especially zooarchaeology) and settlement. The Middle–Upper Paleolithic transition and Upper Paleolithic art also receive detailed treatment, and the Upper Paleolithic of Mediterranean Spain is discussed in the broader context of the late Upper Pleistocene of western Europe and the Mediterranean Basin.     

Plumulites and the machaeridian problem

Plumulites richorum sp. nov. is represented by two complete sclerite assemblages from Early Devonian strata of the Humevale Formation in a small quarry northeast of Kinglake West, Victoria. The completeness of these sclerite assemblages necessitates a revision of terminology applied to machaeridians. Moreover, the sclerite assemblages make it necessary to erect a new Family Plumulitidae and allow several deductions about the animal itself, namely: 1, there was a discrete head possibly with soft anterior projections; 2, there was considerable modification of sclerites and their arrangement near the head, indicating possible sexual dimorphism; 3, sclerites were almost certainly not rigidly attached to the soft parts of the animal. The material shows that Plumulites was neither an arthropod nor an echinoderm, but rather a vagile benthic animal probably belonging to the Annelida.     

A second incisoscutid arthrodire (Pisces,Placodermi) from the Late Devonian Gogo Formation,Western Australia

Long, J. A., 1994:03:28. A second incisoscutid arthrodire (Pisces, Placodermi) from the Late Devonian Gogo Formation. Western Australia. Alcheringa 18. 59–69. ISSN 0311-5518.

A new incisoscutid arthrodire is described from the Upper Devonian Gogo Formation of Western Australia as Gogosteus sarahae gen. et sp. nov. It is characterised by its narrow headshield with cheek unit firmly attached to skull roof, crushing gnathal plates, deep postnasal plates, elongate anterior lateral plate with deeply embayed postbranchial notch and squarely cut posterior margin, and fine dermal ornamentation. The family Incisoscutidae Denison 1984 is redefined to include Incisoscutum ritchiei Dennis & Miles 1981 and Gogosteus gen. nov., and the superfamily Incisoscutoidea nov. defined to include Incisoscutidae and Camuropiscidae.     

Vertebrate microremains from the presumed earliest Carboniferous of the Mansfield Basin,Victoria

The Agnostida are the most important part of the fauna of the Abbey Shales, which is the best-known middle St David's (Middle Cambrian) sequence in Britain. Revision of Illing's (1916) collections allows a correlation of the Paradoxides hicksii fauna with Westergaard's (1946) Ptychagnostus atavus-Tomagnostus fissus Zone, and the Paradoxides davidis fauna with Westergaard's Hypagnostus parvifrons Zone and Ptychagnostus punctuosus Zone (based on the Swedish sequence). A Metadiscus sp. is described. Six species of Condylopygidae are discussed, of which three show a pygidial ‘fringe’. Eleven species of Quadragnostinae, three of Ptychagnostinae (one showing a hypostome), and four of Tomagnostinae (including young growth-stages) are illustrated or discussed. Two species of Diplagnostidae, one species of Phalacroma and two of Phalagnostus are illustrated.     

Palynological evidence for the age of the synorogenic Brewer Conglomerate,Amadeus Basin,central Australia

An assemblage of spores recovered from the Undandita Member of the Brewer Conglomerate, the uppermost unit of the Pertnjara Group in the Amadeus Basin of central Australia, establishes the age of that formation as Late Devonian. Comparison with spore assemblages from sedimentary basins in Western Australia indicates that the central Australian assemblage is dateable within the post-early Frasnian to pre-late Famennian interval. Fish remains and spores previously described from the Parke Siltstone at the base of the Pertnjara Group are of probable early Frasnian age, so it now appears that the entire group was deposited during the Late Devonian. The Brewer Conglomerate is of synorogenic origin, and was deposited during uplift associated with the Alice Springs Orogeny; the spore data thus indicate that this tectonic event began in the Late Devonian. Isotopic age determinations in the Arltunga Nappe Complex and the Strangways Range of the Arunta Block have yielded Early Carboniferous dates. These possibly reflect a later stage of deformation than that which was responsible for the deformation and folding of the Brewer Conglomerate.