Cainozoic euphorbiacean wood from the Canning Basin,Western Australia

A new species of fossil wood is described, Bridelioxylon canningense Bamford & McLoughlin, belonging to the Phyllanthoideae group of the Euphorbiaceae. The woods occur in indurated sandstones and conglomerates preserved in palaeochannels incised into the Upper Permian Condren Sandstone in the Landrigan Cliffs, northern Canning Basin, Western Australia. Based on the presence of the dicotyledonous woods and the geomorphological setting of the palaeochannel, a Palaeogene age is suggested for the host rocks. This is the first record of fossil euphorbiacean wood in Australia; pollen has been previously recorded. The fossil wood is most closely comparable to extant members of Bridelia that currently occupy the warmer and wetter parts of Australia.     

Middle Ordovician (Darriwilian) conodonts from the Goldwyer Formation of the Canning Basin,Western Australia

Zhen, Y.Y., Normore, L.S., Dent, L.M. & Percival, I.G., 11 July 2019. Middle Ordovician (Darriwilian) conodonts from the Goldwyer Formation of the Canning Basin, Western Australia. Alcheringa 44, 25–55. ISSN 0311-5518

Middle Ordovician conodonts attributed to 46 species were recovered from a stratigraphic interval spanning the Willara, Goldwyer and Nita formations in core sections from the Sally May-2 and Theia-1 petroleum exploration wells in the Canning Basin, Western Australia. The Histiodella serrata, Histiodella holodentata and Eoplacognathus pseudoplanus biozones are recognized in the lower and middle part of the Goldwyer Formation, indicative of an early–middle Darriwilian age. This revised conodont biostratigraphy enables more precise correlation with North America and North and South China. Several biogeographically distinctive conodont species, most likely of North Chinese origin, are recorded from the Goldwyer Formation. Their presence signals a strong palaeobiogeographic connection between the Sino-Korean Craton and the Canning Basin on the western margin of eastern Gondwana during the late Middle Ordovician.

Y.Y. Zhen* [yong-yi.zhen@planning.nsw.gov.au], W.B. Clarke Geoscience Centre, Geological Survey of New South Wales, 947–953 Londonderry Road, Londonderry NSW 2753, Australia; L.S. Normore [leon.normore@dmirs.wa.gov.au]; L.M. Dent [louisa.dent@dmirs.wa.gov.au], Department of Mines, Industry Regulation and Safety, Mineral House, Geological Survey of Western Australia, 100 Plain Street, East Perth, WA 6004, Australia; I.G. Percival [ian.percival@planning.nsw.gov.au], W.B. Clarke Geoscience Centre, Geological Survey of New South Wales, 947–953 Londonderry Road, Londonderry NSW 2753, Australia;     

Carboniferous platyceratid gastropods from Western Australia and a possible alternative lifestyle adaptation

Cook, A.G. &; Jell, P.A., September 2015. Carboniferous platyceratid gastropods from Western Australia and a possible alternative lifestyle adaptation. Alcheringa 40, XX–XX. ISSN 0311-5518

Platyceratid gastropods, common and in many cases abundant as elements of middle Palaeozoic gastropod faunas worldwide, are rare or absent in Australian Devonian faunas. In Australia, the earliest abundant platyceratids occur in the Lower Carboniferous (Tournaisian) echinoderm-rich Septimus Limestone and Enga Sandstone in the Bonaparte Gulf Basin, Western Australia. Four taxa, each with significant morphological plasticity, are recognized. In Platyceras (Platyceras) tubulosus (de Koninck, 1883 Koninck, L.G. De, 1883. Faune du calcaire Carbonifère de La Belgique, Quatrième Partie, Gastropodes (suite et fin). Annales du Musée Royal D’Histoire Naturelle de Belgique Tome VIII, Text, 1–240, pls 1–54. [Google Scholar]), three rows of long radially arranged spines and common pentameral symmetry of re-entrants on the aperture suggest an alternative possibility that a relationship between echinoderms and platyceratids developed, and that this may be with archaeocidaroids that are commonly preserved with the gastropods. Similarly in the singly spinose Platyceras (Platyceras) emmemmjae sp. nov., re-entrants suggest an echinoderm relationship. It is proposed that an echinoderm–Platyceras relationship possibly developed in Australia only after a suitable echinoid host had evolved allowing an alternative way for a gameto- or coprophagous habit to be exploited fully.

Alex G Cook [alex.Cook@y7mail.com] and Peter A. Jell [p.jell@uq.edu.au], School of Earth Sciences, The University of Queensland, Queensland 4072, Australia.     

Middle Devonian gastropods from Xiangzhou and Wuxuan,Guangxi, South China

Eighteen taxa of Middle Devonian (Givetian) gastropods, including two new genera and five new species, are présent in the Tungkangling and Yingtang Formations, Wuxuan and Xiangxhou counties, Guangxi Province, China. Pingtianispira tuberculata gen. et sp. nov., Wuxuanella nodusa gen. et sp. nov. and Wuxuanella luifengshanensis sp. nov., Crenulazone wuxuanensis sp. nov. and Murchisonia luifengshanensis sp. nov. are erected from this distinct fauna. The fauna is dominated by nodose and unusual murchisonioids and has strong European and other Old World realm affinities.     

Revision of two phragmodontid species (Conodonta) from the Darriwilian (Ordovician) of the Canning Basin in Western Australia and phylogeny of the Cyrtoniodontidae

Zhen, Y.Y. 9 July 2019. Revision of two phragmodontid species (Conodonta) from the Darriwilian (Ordovician) of the Canning Basin in Western Australia and phylogeny of the Cyrtoniodontidae. Alcheringa XX, xxx–xxx. ISSN 0311-5518

Based on re-examination of the material used in the original study from the subsurface Goldwyer and Nita Formations (middle Darriwilian, Middle Ordovician) of the Canning Basin, Western Australia, two phragmodontid species (Phragmodus polystrophos Watson and Ph. spicatus Watson) are revised as having a septimembrate apparatus including geniculate (Ph. polystrophos) or nongeniculate (Ph. spicatus) M, triform alate Sa, modified tertiopedate (Ph. polystrophos) or tripennate (Ph. spicatus) Sb, modified bipennate Sc, modified quadriramate Sd, carminate Pa and pastinate Pb elements. Characterized by a carminate Pa element in their respective species apparatuses, these two species demonstrate a close phylogenetic relationship with Phragmodus cognitus Stauffer from the Late Ordovician (Sandbian) of North America. These distinctive shared characters have allowed their accommodation within a new genus, Protophragmodus gen. nov., which represents an evolutionary lineage separated from species of Phragmodus Branson & Mehl (sensu stricto). In addition, it is postulated that the Cyrtoniodontidae might have originated in the early–middle Darriwilian from ‘Plectodina’ in shallow-water settings, with Phragmodus (sensu stricto), the most derived part of the family, perhaps directly evolving from Protophragmodus gen. nov. in the late Darriwilian and then becoming cosmopolitan, deeper-water dwellers in the Late Ordovician.

Y.Y. Zhen [yong-yi.zhen@planning.nsw.gov.au], Geological Survey of New South Wales, W.B. Clarke Geoscience Centre, 947–953 Londonderry Road, Londonderry, NSW 2753, Australia.     

Ordovician acritarchs from the Canning Basin,Western Australia

A left mandibular toothplate of a chimaeroid fish collected from the Toolebuc Formation (Early Cretaceous, Albian), central Queensland is described. Comparisons with other chimaeroid genera show the toothplate to be sufficiently distinct to warrant the erection of a new genus. Ptyktoptychion tayyo gen. et sp. nov. is distinguished by the size and shape of its three tritors, the form of the symphysial facet and the overall shape of the toothplate.     

A second incisoscutid arthrodire (Pisces,Placodermi) from the Late Devonian Gogo Formation,Western Australia

Long, J. A., 1994:03:28. A second incisoscutid arthrodire (Pisces, Placodermi) from the Late Devonian Gogo Formation. Western Australia. Alcheringa 18. 59–69. ISSN 0311-5518.

A new incisoscutid arthrodire is described from the Upper Devonian Gogo Formation of Western Australia as Gogosteus sarahae gen. et sp. nov. It is characterised by its narrow headshield with cheek unit firmly attached to skull roof, crushing gnathal plates, deep postnasal plates, elongate anterior lateral plate with deeply embayed postbranchial notch and squarely cut posterior margin, and fine dermal ornamentation. The family Incisoscutidae Denison 1984 is redefined to include Incisoscutum ritchiei Dennis & Miles 1981 and Gogosteus gen. nov., and the superfamily Incisoscutoidea nov. defined to include Incisoscutidae and Camuropiscidae.     

A new Late Devonian acanthodian fish from the Hunter Formation near Grenfell,New South Wales

The ischnacanthid acanthodian Grenfellacanthus zerinae gen. et sp. nov. is described on the basis of two large jaw bones from the Late Devonian (late Famennian) Hunter Formation, near Grenfell, N.S.W. The new species is the youngest known ischnacanthid, and the largest ischnacanthid from Gondwana. As for many ischnacanthids, the structure of the jaws and teeth indicate that Grenfellacanthus was probably an ambush predator.     

Neoproterozoic microfossils from the Officer Basin,Western Australia

Microfossil assemblages are described from the early Neoproterozoic Madley and Browne Formations, western Officer Basin. One chert and eleven siliciclastic samples yielded microfossils. Myxococcoides cantabrigiensis occurs as pustular mats in the chert sample and Eomicrocystis malgica, Pterospermopsimorpha granulata, Skiagia sp. cf. S. pusilla, and undetermined species of Obruchevella, Heliconema, and Trachystrichosphaera are present in acid macerated samples. Leiosphaeridia spp. and Siphonophycus spp. are also found in fine-grained siliciclastic samples, with clusters of Synsphaeridium sp. in some samples. These findings enable a more substantial reconstruction of the palaeoenvironment of Supersequence 1 in the western Centralian Superbasin. The acanthomorph acritarchs are considered to be planktonic eucaryotes washed into environments which ranged from coastal sabkha through to tidal flats, which may be the source of the prokaryotic, benthic, matforming cyanobacteria.     

Late Devonian (Famennian) brachiopod Yunnanella from the Junggar Basin,northern Xinjiang,China

The Late Devonian (Famennian) brachiopod Yunnanella is reported from the western Junggar Basin, northern Xinjiang, on the basis of Yunnanella hanburii (Davidson) and Yunnanella sp.     

Ordovician biostratigraphy of the Canning Basin,Western Australia

Six biostratigraphically distinct faunas based largely on trilobites and graptolites are defined from the Lower to Middle Ordovician limestone, sandstone and shale sequence of the Canning Basin. They range in age from the Tremadoc (fauna 1), through the Arenig (faunas 2, 3) to the Llanvirn (faunas 4–6).     

Microfossils from Neoproterozoic Supersequence 1 of the Officer Basin,Western Australia

The discovery of a rich assemblage of microfossils from the Neoproterozoic western Officer Basin (Centralian Superbasin) provides a more complete understanding of the biostratigraphy of this Basin. The microfossils are found in Supersequence 1 (～800 Ma) in Western Australia. The assemblages are comprised of acritarchs and cyanobacteria isolated by acid maceration from siliciclastics of the Browne (Madley), Hussar, Kanpa and Steptoe Formations. The distinctive acritarchs Cerebrosphaera buickii, Satka colonialica, Stictosphaeridium sinapticuliferum and Pterospermopsimorpha insolita are of particular interest in the Neoproterozoic. These taxa are found in similar depositional environments in Spitsbergen, Arizona, Canada and Siberia. This evidence, together with lithostratographic correlations, isotope chemostratigraphy, and sequence analysis contributes to the continuing development of Neoproterozoic stratigraphy.     

Late Devonian thelodonts (Agnatha) from the Gneudna Formation,Carnarvon Basin,Western Australia

Dermal denticles of two thelodont agnathans, Australolepis seddoni gen. et sp. nov., and a possible nikoliviid gen. et sp. indet. are described from the Gneudna Formation, a marine sequence of interbedded limestone and shale exposed on the eastern edge of the Carnarvon Basin, Western Australia. The age and significance of these new forms are discussed. The Gneudna Formation is Late Devonian, probably early Frasnian, which, if confirmed, makes these the youngest thelodonts known to date.     

Systematic position of two Early Devonian sinistral heterostrophic gastropods from the Garra Limestone,New South Wales

Two Early Devonian gastropod genera, Garraspira gen. nov. and Anoriostoma from the GarraLimestone, with sinistrally heterostrophic shells, share several shell features and are placed in the new tribe Anoriostomatini within the subfamily Agnesiinae. In contrast to the other members of the latter subfamily these genera represent a lineage in which the apertural slit was lost during evolution. This fact supports the opinion that the presence or absence of the apertural slit does not necessarily have significance for high-level taxonomy. Morphology of the gerontic whorl in Anoriostoma sinistra and Garraspira imtsitata suggests their limited mobility during the last ontogenetic stage.     

Ostracods of the Late Devonian Frasnian/Famennian transition from Western Junggar,Xinjiang, NW China

Song, J., Crasquin, S. & Gong, Y., September 2016. Ostracods of the Late Devonian Frasnian/Famennian transition from western Junggar, Xinjiang, NW China. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

Ostracods are described for the first time from the Late Devonian of western Junggar in Xinjiang, NW China. Fifty-two species belonging to 30 genera are recognized, and seven are new: Arcuaria hebukesarensis sp. nov., Bairdia shaerbuertiensis sp. nov., Cribroconcha honggulelengensis sp. nov., Microchelinella bulongourensis sp. nov., M. hoxtolgayensis sp. nov., Pribylites wulankeshunensis sp. nov. and P. junggarensis sp. nov. The ostracod fauna indicates a probable late Frasnian age for the lower member of the Hongguleleng Formation, and the Frasnian/Famennian boundary may exist in the basal part of the formation. The ostracod assemblages are referable to the Eifelian Mega-Assemblage, incorporating both the palaeocopid and smooth-podocopid associations. The fauna implies deposition in a nearshore–offshore environment during a transgression when the lower member of the Hongguleleng Formation was being deposited in western Junggar.

Junjun Song [hnlisa@126.com], State Key Laboratory of Biogeology and Environmental Geology, School of Earth Sciences, China University of Geosciences, Wuhan 430074, PR China; CR2P, MNHN-UPMC- CNRS, Sorbonne Universités, Université Pierre et Marie Curie, T46-56, E.5, case 104, 75252 Paris cedex 05, France; Sylvie Crasquin [sylvie.crasquin@upmc.fr], CR2P, MNHN-UPMC-CNRS, Sorbonne Université, Universités Pierre et Marie Curie, T46-56, E.5, case 104, 75252 Paris cedex 05, France; Yiming Gong* [ymgong@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, School of Earth Sciences, China University of Geosciences, Wuhan 430074, PR China.     

Late Silurian vertebrate microfossils from the Carnarvon Basin,Western Australia

Burrow, C.J., Turner, S., Trinajstic, K. &; Young, G.C., 27 February 2019. Late Silurian vertebrate microfossils from the Carnarvon Basin, Western Australia. Alcheringa 43, 204–219. ISSN 0311-5518.

A core sample from the offshore Pendock 1A well, Carnarvon Basin, Western Australia yielded microvertebrate residues at an horizon in the lower part of the Hamelin Formation, dated as late Silurian, ? Ludlow, based on associated conodonts. The fish fauna comprises loganelliiform thelodont scales, the ? stem gnathostome Aberrosquama occidens nov. gen. et sp., the acanthodian Nostolepis sp. aff. N. alta, and the ? stem osteichthyan Andreolepis sp. aff. A. petri. Because of the paucity of the material, and some differences between the Pendock scales and those of established species, a precise age can not be confirmed; however, the composition of the fauna at generic level most closely resembles that of late Silurian (Ludlow) assemblages from northern Eurasia.

Carole J. Burrow* [carole.burrow@gmail.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Susan Turner [palaeoich@yahoo.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Kate Trinajstic [k.trinajstic@curtin.edu.au], School of Molecular and Life Sciences, Curtin University, Perth, WA 6102, Australia; Gavin C. Young [gavin.young@anu.edu.au], Research School of Physics and Engineering, Australian National University, Canberra, ACT 2000, Australia.     

Asteropygine trilobites from the Late Devonian of the Kerman region,Iran

The discovery of two new Frasnian species of the subfamily Asteropyginae, Bradocryphaeus kermanensis and Radiopyge hamedii, in the Haruz section north of Kerman, east-central Iran, demonstrates for the first time the presence of North Gondwanan trilobites in this region. The two new species provide a link between faunas from Afghanistan to the east and the Alborz Mountains in north Iran. The recognition of R. hamedii confirms the validity of Radiopyge Farsan, hitherto known only from Afghanistan, as well as the presence in some Frasnian Asteropyginae of six pairs of lateral pygidial spines.     

Symbionts in a stromatoporoid-chaetetid association from the Middle Devonian Burdekin Basin,north Queensland

A spectacular association between a chaetetid, a stromatoporoid (Salairella sp.), and straight, vertical tubes interpreted to have housed symbiotic worms is reported from the Givetian Burdekin Formation, Burdekin Basin, north Queensland. The final growth surface of Salairella sp. shows skeletal distortion characterised by long, rather continuous coenosteles and coenostroms with upturned edges. This distorted interface was probably the result of spatial competitive interaction between the encrusting chaetetid and the underlying stromatoporoid. Neither sediment infilling, erosion or surface breakage occurs on the final growth surface of the stromatoporoid skeleton. As such, the growth of the Salairella sp. specimen appears to have been supressed by the encrusting chaetetid. Worm tubes are continuous through the distorted interface without interruption. The skeletal association also suggests that the round, straight, vertical tubes had a symbiotic intergrowth relationship initially with the stromatoporoid and, subsequently, the chaetetid.     

Healed injuries in Early Cambrian trilobites from South Australia

Examples of an undescribed species of the trilobite Redlichia from the Emu Bay Shale (Early Cambrian), Kangaroo Island, South Australia, show damage to the exoskeleton attributed to the action of predators. Injury was probably not lethal. The identity of the predators is unresolved, notwithstanding soft-part preservation within the fossil assemblage. Possible culprits include either a rare and presumably large animal such as an arthropod or conceivably cannibalism by Redlichia itself. This report provides new data on the occurrence of Cambrian predators, and casts further doubt on earlier suggestions that macrophagous predation was insignificant at this time. Aspects of trilobite predation during the Palaeozoic are reviewed, with emphasis placed on their ability to withstand substantial injuries and the possible repair mechanisms that promoted wound healing and survival.