Lark Quarry revisited: a critique of methods used to identify a large dinosaurian track-maker in the Winton Formation (Albian–Cenomanian), western Queensland,Australia

Thulborn, R.A., 2013. Lark Quarry revisited: a critique of methods used to identify a large dinosaurian track-maker in the Winton Formation (Albian–Cenomanian), western Queensland, Australia. Alcheringa, http://dx.doi.org/10.1080/03115518.2013.748482

A remarkable assemblage of dinosaur tracks in the Winton Formation (Albian–Cenomanian) at Lark Quarry, a site in western Queensland, Australia, has long been regarded as evidence of a dinosaurian stampede. However, one recently published study has claimed that existing interpretation of Lark Quarry is incorrect because the largest track-maker at the site was misidentified and could not have played a pivotal role in precipitating a stampede. That recent study has identified the largest track-maker as an ornithopod (bipedal plant-eating dinosaur) similar or identical to Muttaburrasaurus and not, as formerly supposed, a theropod (predaceous dinosaur) resembling Allosaurus. Those iconoclastic claims are examined here and are shown to be groundless: they are based partly on misconceptions and partly on fabricated data that have been assessed uncritically using quantitative measures of questionable significance. Such ill-founded claims do not reveal any substantial flaw in the existing interpretation of the Lark Quarry dinosaur tracks.     

On the integration of Ordovician conodont and graptolite biostratigraphy: new examples from Gansu and Inner Mongolia in China

Wang, Z.H., Bergström, S.M., Zhen, Y.Y., Chen, X. & Zhang, Y.D., 2013. On the integration of Ordovician conodont and graptolite biostratigraphy: New examples from Gansu and Inner Mongolia in China. Alcheringa 37, 510–528. ISSN 0311-5518.

Few Ordovician successions in the world contain both biostratigraphically highly diagnostic conodonts and graptolites permitting an integration between standard biozones based on these fossil groups. The Sandbian Guanzhuang section in the vicinity of Pingliang in the Gansu Province has an outstanding graptolite record through most of the Nemagraptus gracilis and Climacograptus bicornis graptolite biozones. Calcareous interbeds in the succession yield biostratigraphically important conodonts, including some species used for biozonations in Baltoscandia and the North American Midcontinent. Likewise, the middle–upper Darriwilian Dashimen section in the Wuhai region of Inner Mongolia hosts both diverse graptolites of the Pterograptus elegans, Didymograptus murchisoni and lowermost Nemagraptus gracilis biozones, and conodonts of Midcontinent and Baltoscandic types. The distribution patterns of these index fossil groups provide an unusual opportunity to closely correlate conodont and graptolite biozones in the middle to upper Darriwilian to Sandbian interval. For instance, the base of the C. bicornis Biozone is approximately coeval with the base of the Baltoscandic B. gerdae Subbiozone and a level near the middle of the North American P. aculeata Biozone.

Zhi-hao Wang [zhwang@nigpas.ac.cn] Xu Chen [xu1936@gmail.com], and Yuan-dong Zhang [ydzhang@nigpas.ac.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; Stig M. Bergström [stig@geology.ohio-state.edu], School of Earth Sciences, Division of Earth History, The Ohio State University, Columbus, OH 43210, USA; Yong Yi Zhen [yongyi.zhen@austmus.gov.au], Australian Museum, 6 College Street, Sydney NSW 2010, Australia.     

Lower Ordovician trilobites from the Septembersø formation,North-East Greenland

McCobb, L.M.E., Boyce, W.D., Knight, I. & Stouge, S., 2014. Lower Ordovician trilobites from the Septembersø formation, North-East Greenland. Alcheringa 38, 575–598. ISSN 0311-5518.

The informally named Septembersø formation is a 76 m thick succession of microbial and peritidal shelf carbonates deposited on the North-East Greenland shelf of Laurentia. The formation, assigned to the lower part of the Cape Weber Formation in all previous studies, lies disconformably upon the Skullrockian Antiklinalbugt Formation (revised) and conformably below the Tulean to Blackhillsian Cape Weber Formation (revised) in the Fimbulfjeld Group. With the exception of Randaynia, the modest trilobite fauna recovered from the Septembersø formation consists exclusively of bathyurids, and all represent new species. Both Chapmanopyge knudseni sp. nov. and Punka adamsi sp. nov. are represented by sufficient material to merit specific names. The remaining taxa, belonging to Bolbocephalus, Peltabellia, Randaynia and Chapmanopyge are left in open nomenclature. The trilobite genera present suggest that the Septembersø formation is referable to the Tulean Stage of the Ibexian Series, latest Tremadocian/earliest Floian in Global Standard terms.

Lucy M. E. McCobb [lucy.mccobb@museumwales.ac.uk], Department of Natural Sciences, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK; W. Douglas Boyce [dougboyce@gov.nl.ca] and Ian Knight [ianknight@gov.nl.ca], Geological Survey, Newfoundland and Labrador Department of Natural Resources, PO Box 8700, St. John’s, NL, Canada A1B 4J6; Svend Stouge [svends@snm.ku.dk], Natural History Museum of Denmark (Geological Museum), Øster Voldgade 5–7, DK-1350 Copenhagen K, Denmark.     

The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914)—implications for the identification of Pseudisograptus manubriatus subspecies

VandenBerg, A.H.M. & Maletz, J., April 2016. The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914)—implications for the identification of Pseudisograptus manubriatus subspecies. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The holotype of Pseudisograptus manubriatus manubriatus (Hall, 1914) has been rediscovered in the collections of Museum Victoria and is recognized to belong to Pseudisograptus manubriatus harrisi as described by Cooper & Ni (1986). This taxon becomes a junior synonym of P. m. manubriatus. The selected neotype of Pseudisograptus manubriatus manubriatus (Hall, 1914) can be referred to Pseudisograptus manubriatus texanus Cooper & Ni, 1986. The proximal development in all known relief specimens of Pseudisograptus and their descendants indicates that a symmetrical development of the manubrium must be regarded as the rule, and the interpretation of a strong asymmetry in Pseudisograptus manubriatus texanus is rejected.

Alfons H.M. VandenBerg [avandenberg@museum.vic.gov.au, lanceolatus@hotmail.com], Geology, Museum Victoria, GPO Box 666, Vic 3001, Australia; Jörg Maletz [yorge@zedat.fu-berlin.de], Department of Earth Sciences, FU Berlin, Malteserstrasse 74-100, D-12249 Berlin, Germany.     

A new assemblage of graptolites,rhabdopleuran hemichordates and chitinous hydroids from the late Arenig (Ordovician) of the Banestan area,east-central Iran

A rich benthic and planktonic graptolite fauna is associated with encrusting rhabdopleuran hemichordates and chitinous hydrozoans in the late Arenig (Ordovician) part of the Katkoyeh Formation at Banestan, east-central Iran.

New taxa described here are the dendroid graptolites Callograptus huckriedei sp. nov. and Dictyonema bitubulata sp. nov. Other dendroid graptolites Dendrograptus sp. cf. D. flexuosus J. Hall, 1865; Acanthograptus divergens Skevington, 1963; and Thallograptus ?succulentus Ruedemann, 1904 are described. The tuboid graptolite Galeograptus sp., the rhabdopleuran hemichordates Rhabdopleura sp. aff. R. primaevus (Kozlowski, 1967) and Kystodendron sp., and the chitinous hydroid Palaeotuba sp. are also described. Graptoloid graptolites described here are Aulograptus? sp., Didymograptus incertus Perner, 1895, Undulograptus formosus (Mu & Lee, 1958), and Yutagraptus sp. cf. Y. mantuanus Riva, 1994. All but two species-level taxa are described from Iran for the first time.

The fauna is confirmed as being late Arenig or, less likely, early Llanvirn and thus probably correlates with the early part of the Darriwilian stage. It resembles the Atlantic cold water faunas in lacking isograptids and sinograptids.     

Rare encrusted lingulate brachiopods from the Cambrian–Ordovician boundary beds of Baltica

Vinn, O., December 2015. Rare encrusted lingulate brachiopods from the Cambrian–Ordovician boundary beds of Baltica. Alcheringa 40, xx–xx. ISSN 0311-5518

Encrustation is rare on late Cambrian and Tremadocian brachiopods of Baltica. The encrusting fauna is represented by a single taxon, Marcusodictyon. Only Schmidtites celatus is encrusted in the Furongian of Estonia. The Marcusodictyon–Schmidtites association is the earliest example of syn vivo encrustation and symbiosis from the Baltica palaeocontinent. The encrusting faunas of the late Cambrian and Tremadocian of Baltica were unusual presumably owing to palaeogeographic reasons because the other known examples of early encrustation originate from lower palaeolatitudes.

Olev Vinn [olev.vinn@ut.ee], Department of Geology, University of Tartu, Ravila 14A, 50411 Tartu, Estonia.     

An allosaurid theropod dinosaur from the Early Cretaceous of Victoria,Australia

The discovery of an astragalus of an allosaurid theropod in southeastern Australia extends the geographic range of the Allosauridae to include all continents except Antarctica and the Indian subcontinent. In addition, it confirms the presence of this family during the Early Cretaceous.     

Early Ordovician conodonts from Zhejiang Province,southeast China and their biostratigraphic and palaeobiogeographic implications

Forty-two conodont species are documented from the Liuxia, Shijiatou and Jingshan formations in Zhejiang Province of southeast China, located palaeogeographically on the Jiangnan Slope offshore to the Yangtze Platform. From these faunas, eight successive conodont biozones of Tremadocian to middle Floian (Early Ordovician) age are recognized, including the Cordylodus lindstromi Biozone, Cordylodus angulatus Biozone, Chosonodina herfurthi Biozone, Paltodus deltifer Biozone, Paroistodus proteus Biozone, Triangulodus bifidus Biozone, Serratognathus diversus Biozone and Prioniodus elegans Biozone. Several zonal index species of the Baltoscandian succession—Paltodus deltifer, Paroistodus proteus and Prioniodus elegans—are described and illustrated in detail for the first time from South China. Co-occurrence of P. proteus and Serratognathus bilobatus in several samples below the appearance of P. elegans also confirms correlation of the S. diversus Biozone (basal Floian) with the upper P. proteus Zone of the Baltoscandian succession. These Zhejiang faunas are dominated by pandemic forms, and are similar to those of the Open-Sea Realm elsewhere, inhabiting deep, offshore environments.     

Earliest known pelecypod on Earth — a new Early Cambrian genus from South Australia

A new genus and species of fordilloid pelecypod, Pojetaia runnegari gen. et sp. nov., is described from the lower part of the Parara Limestone, Yorke Peninsula, South Australia. Establishing the age of the sample on an international scale is difficult, as with almost all Early Cambrian formations, but evidence tends to suggest that this is the oldest pelecypod so far recorded. Clearly defined hinge teeth and sockets on both valves and on internal moulds help to confirm the pelecypod placement of fordilloids.