Two new cowries (Gastropoda: Cypraeidae) from the middle Miocene of South Australia

Yates, A.M., December, 2008. Two new cowries (Gastropoda: Cypraeidae) from the middle Miocene of South Australia. Alcheringa 32, 353–364. ISSN 0311-5518.

The South Australian specimens of the cypraeids Umbilia leptorhyncha (McCoy, 1877) and Lyncina (Austrocypraea) contusa (McCoy, 1877) are re-examined. Umbilia caepa sp. nov. differs from U. leptorhyncha in its smaller size, more strongly pyriform shape, weaker and less extensive apertural dentition, plate-like columellar margin of the posterior canal and more extensive basal flanges. True U. leptorhyncha is also recorded from the Cadell Formation of South Australia, demonstrating that the two species were sympatric in the Murray Basin. The specimens originally referred to Cypraea contusa var. from the Cadell Formation have had a confusing taxonomic history and they are here named as a new species Lyncina (Austrocypraea) cadella sp. nov. The new species differs from true L. (A.) contusa in its smaller size, less extensive malleations of the dorsal surface, fewer apertural teeth and a projecting internal margin of the fossula. These two new species boost a small but growing list of species that were endemic to the Murray Basin during the middle Miocene.     

Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted,early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia

Chamberlain, P.M., Travouillon, K.J., Archer, M. & Hand, S.J., November 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa 40, XX–XX. ISSN 0311-5518.

A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.

Philippa M. Chamberlain [philippa.chamberlain@uq.net.au], School of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia; Kenny J. Travouillon [kenny.travouillon@museum.wa.gov.au; kennytravouillon@hotmail.com], Western Australian Museum, Locked Bag 49, Welshpool DC, WA, 6986, and School of Earth Sciences, University of Queensland, St Lucia, Queensland, 4072, Australia; Michael Archer [m.archer@unsw.edu.au] and Suzanne J. Hand [s.hand@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, New South Wales, 2052, Australia.     

Campanile (Mollusca: Gastropoda): a new record from the early Miocene of Victoria,Australia

Campanile rupicolum sp. nov. is described from the early Miocene Upper Maude Limestone Member of Maude, Victoria. The genus has not hitherto been recorded from Victoria, a part of the Southeast Australian biogeographic Province, and was thought to be characteristic of the Austral-Indopacific Province. This is the oldest occurrence of the genus so far recorded from Australia.     

Wombats (Vombatidae: Marsupialia) from the Pliocene Chinchilla Sand,southeast Queensland,Australia

Louys, J., 23.3.2015. Wombats (Vombatidae: Marsupialia) from the Pliocene Chinchilla Sand, southeast Queensland, Australia. Alcheringa 39, XXX–XXX. ISSN 0311-5518

The Chinchilla Local Fauna is one of the richest Pliocene vertebrate fossil assemblages in Australia. However, Vombatidae material preserved in the Chinchilla Sand is very poorly known, and no systematic examination of the wombats from Chinchilla has been conducted. Here I review the cranio-dental and mandibular wombat remains derived from Chinchilla. This material includes both adults and pouch-young specimens. At least five species of wombats are preserved in the fluviatile Chinchilla deposits, although a lack of stratigraphically controlled excavations makes it impossible to determine whether all five species were sympatric. Several wombat taxa are revised: Sedophascolomys gen. nov. is formally erected to replace the invalid ‘Phascolomys’; Vombatus mitchelli (Owen) is recognized as a species distinct from Vombatus ursinus (Shaw), and is recorded for the first time from Chinchilla. In addition to Vombatus mitchelli, the Chinchilla Sand also preserves evidence of Phascolonus gigas, Ramsayia magna, Ramsayia lemleyi and Sedophascolomys medius.

Julien Louys [julien.louys@anu.edu.au], Department of Archaeology and Natural History, School of Culture, History, and Languages, ANU College of Asia and the Pacific, The Australian National University, Canberra, ACT 2601, Australia.     

Miralinidae (Marsupialia: Phalangeroidea) from northern Australia,including the youngest occurrence of the family

Five isolated molars from two localities in the Northern Territory, the middle Miocene Bullock Creek and late Oligocene Kangaroo Well sites, are assigned to the new miralinid genus Barguru, which includes the three new species: Barguru kayir, Barguru maru and Barguru kula. The Miralinidae was previously thought to be restricted to the late Oligocene and early Miocene, but the occurrence at Bullock Creek extends the time range of this family into the middle Miocene. Analysis of metaloph development in the Miralinidae suggests that loph formation in this family followed a different trajectory to that of phalangerids.     

The sthenurine affinity of the Late Miocene kangaroo,Hadronomas puckridgi Woodburne (Marsupialia,Macropodidae)

Previous studies of the dentition and cranial base of Hadronomas puckridgi Woodburne suggested that this large, Late Miocene kangaroo might be related to the Sthenurinae. A recently recovered specimen, in which the neurocranium, rostrum and some of the incisors are preserved, reveals a more definite affinity with the sthenurines in its possession of laterally expanded postorbital crests of the frontals, reduced I² accompanied by enlarged I³, the presence of a well-developed posterior mental foramen, a deep, posteriorly situated digastric eminence and elevated pterygoid fossa of the dentary.     

A new Miocene carnivorous marsupial,Barinya kutjamarpensis (Dasyuromorphia), from central Australia

Binfield, P., Archer, M., Hand, S.J., Black, K.H., Myers, T.J., Gillespie, A.K. & Arena, D.A., June 2016. A new Miocene carnivorous marsupial, Barinya kutjamarpensis (Dasyuromorphia), from central Australia. Alcheringa 41, xx–xx. ISSN 0311-5518.

A new dasyuromorphian, Barinya kutjamarpensis sp. nov., is described on the basis of a partial dentary recovered from the Miocene Wipajiri Formation of northern South Australia. Although about the same size as the only other species of this genus, B. wangala from the Miocene faunal assemblages of the Riversleigh World Heritage Area, northwestern Queensland, it has significant differences in morphology including a very reduced talonid on M₄ and proportionately wider molars. Based on the structural differences and the more extensive wear on its teeth, the central Australian species might have consumed harder or more abrasive prey in a more silt-rich environment than its congener, which hunted in the wet early to middle Miocene forests of Riversleigh.

Pippa Binfield [pippa.binfield@outlook.com], Michael Archer [m.archer@unsw.edu.au], Suzanne J. Hand [s.hand@unsw.edu.au], Karen H. Black [k.black@unsw.edu.au], Troy J. Myers [t.myers@unsw.edu.au] Anna K. Gillespie [a.gillespie@unsw.edu.au] and Derrick A. Arena [rick@sitc.net.au], PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales 2052, Sydney, Australia.     

A new fossil genus of forest wallaby (Marsupialia,Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea

A new macropodine genus and species, Silvaroo bila, is described from the Pliocene Chinchilla Sand of Queensland. The generic concept of Protemnodon is reviewed, and it is concluded that two Pliocene species previously placed in that genus (bandharr and buloloensis) belong in Silvaroo. Species of Silvaroo resemble the modern forest wallabies of Papua New Guinea (species of Dorcopsis and Dorcopsulus) and also bear close phenetic resemblance to the late Miocene Dorcopsoides fossilis. Forest wallabies are not known from mainland Australia after the middle Pliocene, but occur in the late Pliocene and Pleistocene of New Guinea.     

Phylogenetic implications of macropodid (Marsupialia: Macropodoidea) posteranial remains from Miocene deposits of Riversleigh,northwestern Queensland

A new sthenurine taxon, Rhizosthenurus flanneryi gen. et sp. nov. from middle late Miocene deposits of Riversleigh, northwestern Queensland, is established on the basis of material previously assigned to Bulungamayinae. Postcranial remains belonging to an indeterminate basal macropodid are described. Cladistic analysis of 50 discrete postcranial characters coded for 16 ingroup and two outgroup taxa suggests that R. flanneryi is the most plesiomorphic member of a clade containing the late Miocene macropodid Hadronomas puckridgi and crown-group sthenurines. The indeterminate basal macropodid is placed as the immediate sister taxon to the bulungamayine Ganguroo bilamina and a monophyletic clade containing macropodines and sthenurines. This arrangement supports consideration of bulungamayines as ancestral to other macropodids and suggests that macropodines and sthenurines are monophyletic. However, the possibility of a diphyletic origin for Macropodinae and Sthenurinae from within Bulungamayinae cannot be dismissed.     

A second species of the possum Durudawiri (Marsupialia: Miralinidae) from the early Miocene of Riversleigh,northwestern Queensland

Durudawiri anfractus sp. nov. (Marsupialia: Miralinidae) is described from Riversleigh. This, the second described species of the genus, is very similar in morphology to, but much larger than, D. inusitatus. Durudawiri anfractus and D. inusitatus are found at similar sites, all early Miocene. The Miralinidae remains one of the most time-restricted families of marsupial, being found so far in only the late Oligocene and early Miocene.     

New record of Warendja wakefieldi (Vombatidae; Marsupialia) from Wombeyan Caves,New South Wales

Transverse sections of Chirodipterus australis and Griphognathus whitei are used to demonstrate the histological structures of the rostral and symphysial tubuli in Devonian lungfish. The walls of the tubuli are composed of bony tissue indistinguishable from the cancellar bone found in dermal bone. It has many spaces and perforations, and is contiguous in places with the cancellar bone of the external dermal bone. The walls of lateral-line canals have the same histological structure as the walls of the tubuli; they also intercommunicate with them. The tubuli and the lateral-line canals open to the surface through large perforations on the external surface. In Griphognathus the walls of the tubes under the external dermal bone have two layers: the outer one is dense bone, but the inner one has the appearance of calcified fibrous connective tissue of the kind associated with the tissue surrounding the lateral-line canals in living lungfish. The tubuli are closely related to the pore-canal system via canals that penetrate the dermal bone; this same relationship has been observed in Dipnorhynchus and Speonesydrion. The intimate connection between the tubuli and the lateral-line system suggests that they were formed by the sinking of neuromasts into and under the external dermal bone, with their walls surrounded by cancellar bone.     

A new theropod dinosaur from South Australia

The Oligocene vegetation at Pioneer was closed temperate rainforest dominated by Nothofagus johnstonii Hill, which probably produced N. menziesii-type pollen. However, other angiosperms (Quintinia, Cupaniae, Ilex, Cunoniaceae, Myrtaceae, Proteaceae and Winteraceae) were also present, as well as several conifers (Athrotaxis, Phyllocladus, Podocarpus, Dacrydium, Dacrycarpus and Araucariaceae). This rainforest was floristically more complex that the modern Tasmanian Nothofagus cunninghamii rainforests but contained many taxonomically related elements. One major difference was that a fern similar to extant Cyathea filled the riparian niche now largely occupied by the tree-fern Dicksonia antarctica. There is indirect evidence that species producing Nothofagus brassii-type pollen may have occurred upstream of the site of deposition, suggesting that the Nothofagus species were altitudinally zoned or edaphically restricted. The current absence of many of these Nothofagus species in Tasmania may be due to their inability to survive the low temperatures of the Quaternary glaciations. The high degree of similarity of the Pioneer palynoflora to that recorded in Oligocene sediments in onshore (Partridge, 1971) and offshore (Stover &; Partridge, 1973; Stover &; Evans, 1973) Gippsland Basin strongly suggests that there was little regional differentation in southeastern Australia at that time.     

A new species of Scirrotherium Edmund & Theodor, 1997 (Xenarthra,Cingulata, Pampatheriidae) from the late Miocene of South America

Góis, F., Scillato-Yané, G.J., Carlini, A.A. & Guilherme, E. 2013. A new species of Scirrotherium Edmund & Theodor, 1997 (Xenarthra, Cingulata, Pampatheriidae) from the late Miocene of South America. Alcheringa 37, 175–186. ISSN 0311-5518.

A new species of Scirrotherium Edmund & Theodor (Pampatheriidae) is described: S. carinatum. This genus was previously represented by a single species, S. hondaensis Edmund & Theodor, 1997, recorded from the middle Miocene (Laventan) of Colombia. Pampathere remains are common in the ‘Conglomerado osífero’ (late Miocene) of the Ituzaingó Formation in the Paraná River cliffs of Entre Ríos, Chubut (Argentina) and Acre (Brazil). All of them were referred to Kraglievichia paranense, and they are mostly osteoderms. However, only a few of these specimens are strictly consistent with that species. The new species described herein differs from S. hondaensis in having very thin osteoderms with more elaborate ornamentation. The movable band (or imbricate) osteoderms have a sculptured exposed surface and a single transverse row of anterior foramina; the lateral margins are wider and also bear foramina, and the longitudinal central elevation is long and proximally wide, and forms a posteriorly raised crest. This elevation is delimited by a fairly broad and shallow depression on each side, separating the marginal elevation from the longitudinal central one. The exposed surfaces of fixed (or fused) osteoderms bear large and deep anterior foramina, the longitudinal central elevation and delimiting depressions are very pronounced. This new species increases the known diversity of the pampatheres from the ‘Conglomerado osífero’. Additionally, a new nomenclatural scheme for Pampatheriidae osteoderms is proposed.

Flávio Góis [goisf@fcnym]unlp.edu.ar], Gustavo Juan Scillato-Yané [scillato@fcnym.unlp.edu.ar], Alfredo Armando Carlini [acarlini@fcnym.unlp.edu.ar], Departamento Científico Paleontología Vertebrados, Museo de La Plata, Paseo del Bosque, s/n, 1900, La Plata, Buenos Aires, Argentina and CONICET; Edson Guilherme [guilherme@ufac.br], Universidade Federal do Acre, Laboratório de Pesquisas Paleontológicas (LPP), BR 364, Km 04, 69.915-900. Rio Branco, AC, Brazil. Received 11.5.2012; revised 31.7.2012; accepted 4.9.2012 .     

Ngapakaldia bonythoni (Marsupialia,Diprotodontidae): new material from Riversleigh,northwestern Queensland,and a reassessment of the genus Bematherium

A latest Permian (late Changhsingian) radiolarian fauna is recorded from the upper Talung Formation, Hushan, Nanjing, Jiangsu Province, South China. This fauna includes 24 species belonging to 16 genera; new species are Albaillella hushanensis, Copicyntroides stellatus and Trilonche crassus. The presence of the radiolarian fauna and its taxonomic composition reveal that the Eastern Qinling-Dabie deep sea, which was located along the northern margin of the northeastern Yangtze Basin, persisted at least until the end of the Palaeozoic and that the collision between the North China and South China plates had not occurred by the end of the Permian.     

A new Pliocene tree kangaroo species (Marsupialia,Macropodinae) from the Chinchilla Local Fauna,southeastern Queensland

A new species of very large tree kangaroo, Bohra wilkinsonorum, is described from a maxillary fragment from the Pliocene Chinchilla Sands of southeastern Queensland. Allocation to Bohra, which has previously been known from postcranial material only, is suggested on the basisof its similar size and stage of evolution to Bohra paulae. Both species of Bohra are plesiomorphic with respect to species of Dendrolagus, and are much larger than any known species of Dendrolagus. This new taxon from Chinchilla has expanded the tree kangaroo record from the Pliocene of southeastern Australia, supporting the hypothesis that the group originated in the late Miocene of ‘mainland’ Australia, finding refuge in north-eastern Queensland and New Guinea as climate became drier in the Quaternary. Fossil tree kangaroos are unknown from the Pliocene of Papua New Guinea where most living species now occur.     

A new petauroid possum (Marsupialia,Pseudocheiridae) from the Pliocene Bluff Downs Local Fauna,northern Queensland

Pseudochirops winteri n. sp. is described from the Pliocene Bluff Downs Local Fauna on the basis of a left dentary fragment with M₂. It is considered to be most similar to P. archeri although its phylogenetic position must remain tentative until further material is found. The presence of this presumably rainforest-dwelling possum, the first from this fauna, may indicate a limited closed forest component. Given the paucity of additional rainforest taxa recovered from this site despite extensive screen washing, however, such a component must have been restricted to gullies or riparian fringes.     

Miocene wood from the LaTrobe Valley coal measures,Victoria, Australia

An initial study of a collection of fossil conifer wood is reported from the late early Miocene Yallourn Clays, an interseam unit intergrading into the base of the early to middle Miocene Yallourn seam of the LaTrobe Valley, Victoria in southeastern Australia. The fossil wood shares characteristics with the modem genera Dacrycarpus and Dacrydium. On the basis of contiguous, uniseriate tracheid pitting and 1–2 podocarpoid cross field pits, it is placed in the form genus Podocarpoxylon, and the new species P. latrobensis. The wood is compared with extant Podocarpaceae and other Australian fossil woods. Its ring anatomy is consistent with low temperature or rainfall seasonality in the early Miocene.     

A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales,Australia)

?erňanský, A. & Hutchinson, M.N., 2012. A new large fossil species of Tiliqua (Squamata; Scincidae) from the Pliocene of the Wellington Caves (New South Wales, Australia). Alcheringa, 1–6. ISSN 0311-5518.

We describe an isolated frontal bone referable to a new species, Tiliqua laticephala (Scincidae), from the Pliocene Big Sink doline of the Wellington Caves, central eastern New South Wales, Australia. The bone is very robust, is unusually broad and thick, especially around the bases of the subolfactory processes and represents a large and heavily built lizard. The fossil has multiple fragmentary osteoderms in the frontal region, showing asymmetry in shape and thickness that do not correspond to the more regularly arranged anterior head shields of other scincoids. The specimen shares two unusual character states with extant Tiliqua, especially the large armoured species, T. rugosa. Other large skinks related to T. rugosa (other Tiliqua spp., Corucia zebrata, Egernia cunninghamii, Bellatorias major, Liopholis kintorei) are less similar in terms of frontal shape, thickness, sculpture, osteoderm ornamentation, and positioning of adjacent bones.     

Fossil marsupial predators of South America (Marsupialia,Borhyaenoidea): bite mechanics and palaeobiological implications

Borhyaenoids were marsupial predators that inhabited South America during the Cenozoic. They were very significant because no other mammals rivaled them as terrestrial hunters of large prey. Here we estimate the bite force of three species of borhyaenoids by two different methods to infer predatory behaviour in extinct taxa. One of the methods uses mainly the skull and only some simple measurements of the mandible; the other uses several measurements within the dentary. The results show that bite forces are very high in comparison to predators of the order Carnivora, a feature manifest by several other living and extinct marsupial predators. Differences in size, bite mechanics and special adaptations among the borhyaenoids suggest a very wide range of predatory behaviours that rival those represented in the extant families of the Order Carnivora.     

A probable ankylosaurian (Dinosauria,Thyreophora) from the Early Cretaceous of New South Wales,Australia

Bell, P.R., Burns, M.E. & Smith, E.T. October 2017. A probable ankylosaurian (Dinosauria, Thyreophora) from the Early Cretaceous of New South Wales, Australia. Alcheringa 42, 120–124. ISSN 0311-5518.

We describe an isolated osteoderm from the Albian Griman Creek Formation where it is exposed near the town of Lightning Ridge in central-northern New South Wales, Australia. Several lines of evidence allow referral of this element to the Ankylosauria—a group that epitomises body armour and ubiquitous osteodermal coverage among dinosaurs. Despite the abundant record of fossil vertebrates from this interval, ankylosaurians have not been previously reported, although, they have been described from penecontemporaneous deposits in western Queensland and Victoria. This discovery, therefore, provides an important link between the northerly faunas (including the Griman Creek Formation) that flourished at the edge of the epeiric Eromanga Sea, with those from the sub-polar rift-valley system of Victoria during the mid-Cretaceous.

Phil R. Bell [pbell23@une.edu.au], School of Environmental and Rural Science, University of New England, Armidale 2351, NSW, Australia; Michael E. Burns [mburns3@jsu.edu], Department of Biology, Jacksonville State University, 700 Pelham Rd N., Jacksonville, AL 36265-2138, USA; Elizabeth T. Smith [elizabethtsmith@exemail.com.au], Australian Opal Centre, Lightning Ridge 2834, NSW, Australia.