Fluid circulation related to post‐Messinian extension,Thassos Island,North Aegean

In the North Aegean Domain, Thassos Island contains a Plio‐Pleistocene basin controlled by a large‐scale flat‐ramp extensional system with a potential décollement located at depth within a marble unit. Numerous mineralizations associated with normal faults of Plio‐Pleistocene age are the sign of fluid circulation during extension. Two main generations of fluid flow are recognized, related to Plio‐Pleistocene extension. A first circulation under high‐temperature conditions (about 100–200°C) resulted in dolomitization of marbles near the base of the Plio‐Pleistocene basin. The dolomites are characterized by low δ¹⁸O values (down to 11‰ versus Standard Mean Ocean Water). Some cataclastic deformation affected the dolomites. Hydrothermal quartz that crystallized in extension veins above a blind ramp also has low δ¹⁸O values (about 13‰). This shows that high‐temperature fluids moved up from the décollement level toward the surface. A second downward circulation of continental waters at near‐surface temperature is documented by calcite veins in fault zones and at the base of the Plio‐Pleistocene basin. These veins have O isotope values relatively constant at about 23–25‰ and C isotope values intermediate between the high δ¹³C value of the carbonate host rock (about 1–3‰ versus Peedee Belemnite) and the low δ¹³C value of soil‐derived carbon (?10‰). The calcites associated with the oxidative remobilization of primary sulphide Zn–Pb mineralization of Thassos carbonates have comparable O and C isotope compositions. Hot fluids, within the 100–200°C temperature range, have likely contributed to the weakening of the lower marble unit of Thassos and, thus, to the process of décollement.     

Pleistocene frogs from the Darling Downs,southeastern Queensland,and their palaeoenvironmental significance

Systematic collecting from fluvial late Pleistocene deposits from the Darling Downs, southeast Queensland, Australia, has led to the recovery of the first fossil frogs from the region, ail from the Myobatrachidae, a family of ground dwelling and burrowing frogs. The most common species recovered, Limnodynastes tasmaniensis, is extant on the Darling Downs. The fossil taxa include species whose extant populations inhabit arid zones(Limnodynastes sp. cf. L. spenceri), montane forests (Kyarranus spp.), and open woodlands (Neobatrachus sudelli), and indicate the existence of a mosaic of habitats during the Pleistocene. The absence of the Hylidae (tree frogs), a family common throughout the Darling Downs today, may be explained by a taphonomic bias that favours non-arboreal forms. Alternatively, hylids may have been rare or absent on the Darling Downs during the Pleistocene.     

Mitochondrial DNA haplotypes of Devensian hyaenas from Creswell Crags,England

Spotted hyaena (Crocuta crocuta) remains have been recovered from British Middle and Upper Pleistocene sites at intervals within the period 700–730 ka BP. Morphological studies have suggested that hyaenas of the Last Interglacial sensu stricto (Ipswichian: Marine Isotope Stage [MIS] 5e, 130–115 ka BP) and Last Glacial (Devensian: MIS 3, 61–24 ka BP) were two distinct populations, the Ipswichian hyaenas becoming extinct in Britain during MIS 5 and the Devensian ones arriving via a subsequent migration from continental Europe. However, the apparent presence of hyaenas in later MIS 5 deposits has led to the alternative suggestion that there was a southern relict population from which the Devensian hyaenas originated. We obtained ancient DNA (aDNA) sequences from four Devensian hyaena specimens from Creswell Crags, Derbyshire, dated to around 45 ka ¹⁴C BP. Each of these four specimens belonged to the mitochondrial DNA (mtDNA) clade A. This clade is not thought to have been present in Europe until ~360 ka BP, after the initial arrival of hyaenas in Britain. The DNA results, therefore, suggest that there were at least two waves of hyaena dispersals into Britain. The results are consistent with the repeated dispersals into Britain of another Pleistocene social carnivore, Homo sapiens.     

Presence of the African Saber-toothed FelidMegantereon whitei(Broom, 1937) (Mammalia, Carnivora, Machairodontinae) in Apollonia-1 (Mygdonia Basin, Macedonia, Greece)

We have made a multivariate morphometric study of the fossils ofMegantereonfrom the Apollonia-1 site at the Mygdonia Basin (Macedonia, Greece), using discriminant functions. Results obtained indicate that these specimens belong to the African speciesMegantereon whiteiBroom, which is also found in the Lower Pleistocene of Dmanisi (East Georgia, Caucasus) and Orce (Southern Spain). Morphofunctional studies ofM. whiteisuggest that this saber-toothed felid was an ambush predator with great killing capability in relation to its food requirements, thus leaving enough meat to be scavenged by both hyaenas and hominids.M. whiteimay have played an important ecological role making the first dispersal ofHomoto Eurasia the Lower Pleistocene possible.     

New Human Remains of Archaic Homo sapiens and Lower Palaeolithic Industries from Visogliano (Duino Aurisina, Trieste, Italy)

New human remains have been found during recent excavations conducted in the lower section of Visogliano dolina (Trieste, Italy). They are represented by teeth and a fragment of particularly robust mandible that for their morphological and morphometrical characters can be generically attributed to archaic Homo sapiens. The lithic industry is formed by flakes, carinated points, scrapers, choppers and chopping tools and is characteristic of the Italian Pebble Culture. Sediments, pollen and faunal remains indicate that this part of the deposit was formed during interglacial climatic conditions in the early, but not initial phase of Middle Pleistocene (isotopic stadium 13 or 11).     

The first record of palaeopathology from the zooarchaeological record of late Pleistocene,southwest Tasmania

Several bones from the Late Pleistocene archaeological site, Kutikina Cave, in southwest Tasmania, show pathological conditions. The majority of these specimens represent Bennett's or red‐necked wallaby (Macropus rufogriseus), with a single specimen of the Common wombat (Vombatus ursinus) recorded. While there has been extensive work in this region of Australia since the early 1980s, this is the first record of palaeopathology from the late Pleistocene of Tasmania, and the first from a human accumulation of predominately macropod material. The palaeopathology raises several questions concerning the mobility of these animals which might have increased their susceptibly to predation, while some of the effected elements are fragmented, suggesting that people utilised these bones despite their deformities. Copyright © 2008 John Wiley & Sons, Ltd.     

Pleistocene foraminifera from Botany Bay,New South Wales

Unconsolidated sediments from a borehole in Botany Bay have been analyzed for their foraminiferal content. Faunas from between 41 and 54 m below sea level are interpreted as Pleistocene in age. The foraminiferal assemblage from 54 m indicates marine conditions of moderately deep water (25–30 m), and a climate warmer than at present. The higher assemblages indicate a more brackish environment

The foraminifera, belonging to 123 species in 67 genera, are tabulated and 24 are discussed and illustrated. Three species, Oolina bifidocostata, Fissurina alatoquadrata and Elphidium botaniensis and one subpecies, Quinqueloculina affinis atrata, are described as new.     

Composite hunting technologies from the Terminal Pleistocene and Early Holocene,Niah Cave,Borneo

Renewed archaeological investigation of the West Mouth of Niah Cave, Borneo has demonstrated that even within lowland equatorial environments depositional conditions do exist where organic remains of late glacial and early post-glacial age can be preserved. Excavations by the Niah Cave Research Project (NCP) (2000–2003) towards the rear of the archaeological reserve produced several bone points and worked stingray spines, which exhibit evidence of hafting mastic and fibrous binding still adhering to their shafts. The position of both gives strong indication of how these cartilaginous points were hafted and gives insight into their potential function. These artefacts were recovered from secure and ¹⁴C dated stratigraphic horizons. The results of this study have implications for our understanding the function of the Terminal Pleistocene and Early Holocene bone tools recovered from other regions of Island Southeast Asia. They demonstrate that by the end the Pleistocene rainforest foragers in Borneo were producing composite technologies that probably included fishing leisters and potentially the bow and arrow.     

A new Jungle-fowl from the Pleistocene of Europe

The domesticated Red Jungle-fowl G. gallus is believed to have been dispersed by man from India during the Holocene. The distal end of a radius from the Ipswichian Interglacial deposits at Crayford, Kent, was indistinguishable from that of the wild form of Red Jungle-folw. A coracoid from the early Middle Pleistocene of N Norfolk was also very similar to that species but showed differences comparable with those found between different species of Gallus. At least one species referable to Gallus is known from the Pliocene of South-eastern Europe. Extrapolating from zoogeographical speciation patterns it would be possible for a Gallus species to have evolved through Pleistocene speciation in the European region. The species might have become extinct during a glaciation or have been exterminated by early man. A new species Gallus europaeus is described with the coracoid as a holotype, and the radius from Kent is tentatively referred to it.     

On the occurrence of the scimitar-toothed cat, Homotherium latidens (Carnivora; Felidae), at Kents Cavern,England

Teeth of Homotherium latidens recovered from late Pleistocene sediments, Kents Cavern, England have long been the source of controversy. H. latidens is conspicuously absent from other late Pleistocene cave deposits in Britain, and is widely thought to have been extirpated from the region during the isotope stage 10 glacial period. Here we present high spatial resolution analyses of fluorine and uranium uptake profiles in teeth of three species from the same cave. The H. latidens tooth is clearly distinguished from the unambiguously provenanced Late Pleistocene hyaena and Middle Pleistocene cave bear teeth. These results are consistent with the theory that the H. latidens teeth originated at an exogenous location, were probably transported to Kents Cavern as Palaeolithic trade goods, and were buried in Kents Cavern in Palaeolithic times.     

Lagostomus maximus (Desmarest) (Rodentia,Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay

Ubilla, M. & Rinderknecht, A., April 2016. Lagostomus maximus (Desmarest) (Rodentia, Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The extant plains vizcacha, Lagostomus maximus, is described from the Late Pleistocene (Dolores Formation) of Uruguay based on an almost complete articulated skeleton. It is compared with the nominally extinct Pleistocene species of the genus. An AMS ¹⁴C taxon-age is determined for L. maximus at 11 879 ± 95 years BP (cal. BP 13 898–13 941). Lagostomus maximus is absent from modern mammal communities in Uruguay, and no Holocene evidence is available. Because L. maximus exhibits remarkable sexual and ontogenetic dimorphism, we examined a range of juvenile, sub-adult and adult male and female specimens. It is not possible to differentiate the articulated Pleistocene fossil from sub-adult specimens of L. maximus based on this sample. Moreover, the skull characteristics, including a broad vertical ramus of the zygomatic arch, semi-circular temporal crest and short robust sagittal crest, most closely resemble extant female individuals. Molar size is non-predictive for ontogenetic stage or body mass because it reaches stability during early adulthood. Other fragmentary skull remains are herein assigned to L. sp. cf. L. maximus. Lagostomus cavifrons from the Pleistocene of Argentina is considered synonymous with L. maximus. Its character states concur with the range of intraspecific variability and it has been established on a young adult or adult male individual. Likewise, the other Argentinean Pleistocene species based on incomplete mandibles, L. angustidens, L. striatus, L. heterogenidens, L. egenus, L. minimus and L. debilis, concur with ontogenetic morphs and are here assigned to Lagostomus sp. The ecological preferences of extant L. maximus infer open arid or semi-arid landscapes for the latest Pleistocene of southern Uruguay. This hypothesis is reinforced by the coeval presence of Microcavia, Galea and small camelids in the same strata. The Last Glacial Maximum likely promoted this environmental setting. Latest Pleistocene or early Holocene climatic change might have facilitated local extinctions and/or range shifts among this mammal fauna.

Martín Ubilla [martinubilla@gmail.com], Facultad de Ciencias, Universidad de la Republica, Iguá 4225, 11400, Montevideo, Uruguay; Andrés Rinderknecht [apaleorinder@yahoo.com], Museo Nacional de Historia Natural, Paleontología, 11000, Montevideo, Uruguay.     

Population relationships of later Pleistocene hominids: A multivariate study of available crania

The Neanderthal populations of the Upper Pleistocene have been regarded by a number of anthropologists as direct ancestors to modern man. Results of multivariate analyses conducted in this study suggest that early Neanderthal crania were morphologically more like modern Homo sapiens than were the later Neanderthals. The implications of these results are discussed in the light of archaeological evidence and comparative studies of Pleistocene crania from outside Europe.     

Cut marks on the Middle Pleistocene elephant carcass of Áridos 2 (Madrid,Spain)

Áridos 1 and Áridos 2 (Madrid, Spain) are two Middle Pleistocene sites belonging to the isotopic stages 9–11. Both places contain partial carcasses of Elephas (Paleoxodon) antiquus associated to Acheulian stone tools. In this work, the taphonomic study of the elephant remains of Áridos 2 is presented. This study has documented several cut marks on different bones, which indicate bulk flesh and viscerae extraction by Middle Pleistocene hominins. Several arguments are provided to support that at least some of the cut marks were made with handaxes, further suggesting that some of these artifacts were butchering tools in this stage of human evolution. Although cut marks on elephant carcasses have been documented at some Middle Pleistocene sites, very few have been published in detail to allow consideration of their status as hominin-imparted marks. By doing so, the present study provides more evidence of large carcass exploitation by hominins during this period.     

Monograptus deubeli and praedeubeli (Wenlock,Silurian) in the Asian part of the former Soviet Union

Koren', T. N., & Suyarkova, A. A., 1994:03:28. Monograptus deubeli and praedeubeli (Wenlock, Silurian) in the Asian part of the former Soviet Union. Alcheringa 18, 000–000. ISSN 0311-5518.

During the last decade the sequence and composition of the later Homerian (Wenlock) graptolite fauna has become comparatively well known in many regions of the world. A complete and detailed graptolite biozonation has been elaborated in Europe, and partly confirmed in Australia and Arctic Canada. Valuable new data have been obtained from several widely scattered areas within the Asian part of the former U. S. S. R. M. praedeubeli Jaeger (rare) and M. deubeli (common) have been found in the Homerian sequences in the Novosibirsk Islands and Novaya Zemlya, as well as in the southern Urals and the Turkestan-Alai Mountains of central Asia. In sections studied here, especially in the Alai Range, both species have well defined stratigraphic ranges in a variety of pelagic and hemipelagic facies. Their well established taxonomy based on sicular shape and thecal structure allows their recognition in material of different preservation. They belong to a single phylogenetic line and can be used as important biostratigraphic markers for interregional correlation, especially considering the distinct differences in the composition of associated monograptids. Thus, members of the Pristiograptus idoneus—Lobographu? claudiae lineage parallelling the development of M. praedeubeli and M. deubeli in the Homerian of central Asia, Australia, Arctic Canada and Arctic Russia are unknown from European and northern African sections.     

Further Ektopodontidae (Phalangeroidea,Mammalia) from southwestern Victoria

The youngest members of the Ektopodontidae, an enigmatic family of Phalangeroidea, are reported from southwestern Victoria. Four isolated teeth are described as Ektopodon paucicristata sp. nov. (from the ?mid-?late Pliocene, and early Pliocene) and two isolated Pleistocene teeth are assigned to Darcius duggani Rich, 1986. Previously, the youngest known ektopodontid was early Pliocene.     

Reappraisal of ‘chronospecies’ and the use of Arvicola (Rodentia,Mammalia) for biochronology

The water vole, genus Arvicola, is characterised by a broad geographic distribution throughout Europe and is widespread during the late Middle and Upper Pleistocene. This genus is used as a major biostratigraphic tool within the Quaternary. Specific determinations using the Schmelzband-Differenzierung-Quotient or SDQ have identified many chronospecies within the fossil species Arvicola cantiana (Hinton, 1910). As SDQ calculation remains limited, this study reappraises the Arvicola genus in terms of morphodiversity and morphospace using outline analysis which takes into account the tooth as a whole. Outline analysis suggests that one single species of Arvicola, A. cantiana, was present during the Pleistocene. This species shows great variability with no trends or patterns in morphospace. Thus, these results call into question the reliability of SDQ for specific determinations and throw doubt on the biochronological framework based on Arvicola.     

Comparative Skeletal Pathology and the Case for Conspecific Care in Middle Pleistocene Hominids

Lebel et al. (2001, Proc Natl Acad Sci98, 11097) claim that the pathological alterations to the Aubesier 11 partial mandible demonstrate that Middle Pleistocene hominid populations, “had achieved a level of sociocultural elaboration sufficient to maintain debilitated individuals and to provide the motivation to do so”. This hypothesis is tested here by comparing the pathological processes seen in the Aubesier 11 hominid specimen with those evident in wild non-human primate skeletal material. A variety of wild non-human primates have similar pathological processes of equal or greater severity than that seen in Aubesier 11. There is also no evidence of any condition in Aubesier 11 which would have prevented that individual from providing for himself/herself. Therefore the Aubesier 11 partial mandible cannot be taken as evidence for an increased level of conspecific care in Middle Pleistocene hominids, contra Lebel et al. (2001). This illustrates the importance of a comparative approach to palaeopathology.     

Subsistence continuity,change, and environmental adaptation at the site of Nugljanska,Istria, Croatia

This paper investigates changes in subsistence strategies at the upland cave site of Nugljanska (Croatia) at the end of the last ice age, during a time of rapid sea level rise and changing environment. We analysed the faunal assemblage from archaeological levels spanning the Pleistocene–Holocene transition (15,000–8000 years BP) and evaluated changes in relative abundance of species, measured species diversity, and compared the representation of terrestrial mammal and marine resources. We found a significant shift in the most abundant prey species exploited (from red deer, Cervus, to wild boar, Sus). There was some correlation between dietary diversification and periods of increased moisture availability and the spread of deciduous forest in the area. Our results suggest that there was a continuing reliance on terrestrial resources throughout time and that changes in dietary patterns were likely due to local environmental change and potentially, changing seasonal mobility strategies, at the Pleistocene–Holocene transition.     

The first record of Galea Meyen, 1832 (Rodentia,Hystricognathi, Caviidae) in the late Pleistocene of southern Brazil and its palaeobiogeographic implications

The first caviine rodent referable to Galea Meyen, 1832 is described from the late Pleistocene of southern Brazil based on a left dentary with the p4–m3 series. The specimen derives from the Ponte Velha I locality in the Touro Passo Creek (Touro Passo Formation, upper Pleistocene), western Rio Grande do Sul State. The main characters used to assign this specimen to Galea are: anterior area of horizontal crest at the level of prism I of p4; deep anterior area of masseteric fossa; incisor alveolus on the medial face of the dentary extended up to the level of prism II of m2; and presence of cement in the hypoflexid. Currently, the genus has a disjunct distribution, with a group in Argentina, Bolivia and Peru, and another in northern and northeastern Brazil. The presence of this taxon in Pleistocene deposits of Rio Grande do Sul State, Uruguay and the Argentine Mesopotamian, where there are no extant representatives of the genus, indicates its wider distribution during the late Pleistocene.     

First fossil fish (Teleostei,Siluriformes) from the Late Pleistocene of Santa Fe Province,Argentina

Vallone, E.R., Vezzosi, R.I. & Cione, A.L. February 2017. First fossil fish (Teleostei, Siluriformes) from the Late Pleistocene of Santa Fe Province, Argentina. Alcheringa 00, 000–000. ISSN 0311-5518.

The record of fossil fish from the Pleistocene of Argentina is poor. Here we describe the first ichthyofauna from Late Pleistocene riverbank beds in the Salado River of Santa Fe Province, Argentina. The material consists of isolated pectoral and dorsal fin spines, together with skull fragments. Four species-level taxa referable to three families can be identified: Pterodoras granulosus (Doradidae), Pimelodus cf. maculatus and Pimelodus cf. albicans (Pimelodidae) and cf. Hypostomus sp. (Loricariidae). Specimens attributed to Pterodoras granulosus and Pimelodus maculatus represent a minimum age for origin of these taxa. The Salado River assemblage includes the richest record of Pleistocene catfishes yet documented from southern South America.

Evelyn Romina Vallone [evelynvallone@conicet.gov.ar] and Raúl Ignacio Vezzosi [vezzosiraul@gmail.com], Laboratorio de Paleontología de Vertebrados, CICYTTP-CONICET, Materi y España, (3105) Diamante, Entre Ríos, Argentina; Alberto Luis Cione [acione@museo.fcnym.unlp.edu.ar], División Paleontología de Vertebrados. Museo de La Plata. Paseo del Bosque s/n, (1900) La Plata, Argentina.