Parotosuchus from the Early Triassic of Queensland and Western Australia

Three members of the labyrinthodont Family Capitosauridae are described. Two new species from the Arcadia Formation of the Rewan Group, Queensland, show affinities with more primitive capitosaurs and with pre-capitosaurid groups. A third skull from the Blina Shale of Western Australia is of indeterminate species. These are the first recorded capitosaurs from Queensland and Western Australia and the first Parotosuchus from a Lystrosaurus Zone fauna.     

A horned member of the labyrinthodont super-family Brachyopoidea from the Early Triassic of Queensland

A horned, short-skulled labyrinthodont amphibian from the Arcadia Formation of the Rewan Group near Bluff, Queensland, is placed in the family Chigutisauridae of the superfamily Brachyopoidea. This is the first occurrence of this family outside Argentina. The material, consisting of two skulls and three mandibles is described as Keratobrachyops australis gen. et sp. nov., and used as a basis for an examination of the relationship between the families Brachyopidae and Chigutisauridae.     

Early Cretaceous (Neocomian) microplankton from the Carpentaria Basin,northern Queensland

The Neocomian upper Helby Beds, basal Rolling Downs Group and Gilbert River Formation of the Carpentaria Basin contain a dinoflagellate sequence divisible into three zonal intervals; the zones represent the oldest continuous marine Cretaceous record for eastern Australia. Zone DK1 (dated latest Jurassic to Berriasian) commences with the first appearance of four species, zone DK2 (dated Valanginian) with that of four or five species, and zone DK3 (dated Hauterivian to Barremian) with that of three species.

Biostratigraphic aspects are discussed of species selected as possible zone marker fossils in so far as they (a) are relatively poorly known, or represent new species, and (b) have no previously published Neocomian record. New combinations are proposed of one acritarch species and six dinoflagellate species. One new acritarch genus and species, Pseudofromea collaris, and four new dinoflagellate species — Canningia crassicingulata, Cleistosphaeridium australe, Cyclonephelium asymmetricum, and Muderongia testudinaria — are proposed and described.     

Occurrence of Euowenia grata (De Vis, 1887) (Diprotodontidae,Marsupialia) from the Pliocene Spring Park Local Fauna,northeastern Queensland

Mackness, B.S., Black, K.H. & Price, G.J., 1.10.2014. Occurrence of Euowenia grata (De Vis, 1887 De Vis, C.W. [In Anon.] 1887. Untitled. The Brisbane Courier 9224 (44) (8 August), 6. [Google Scholar]) (Diprotodontidae, Marsupialia) from the Pliocene Spring Park Local Fauna, northeastern Queensland. Alcheringa 39, 000?000. ISSN 0311-5518

Ten specimens including several dentaries and maxillae, recovered from the Pliocene Spring Park Local Fauna, northern Australia, are referred to the diprotodontine Euowenia grata (De Vis). The fossils exhibit minimal dental wear and reveal new characters that are unrecognizable in the holotype. The remains represent at least three animals, effectively doubling the previous number of individuals known for this rare megaherbivore. The new records also provide a significant northern geographic range extension for the species and allow an assessment of intraspecific variation, sexual dimorphism and phylogenetic relationships for the species. Euowenia grata is most similar in morphology to the monotypic Pliocene diprotodontid Meniscolophus mawsoni.

Brian S. Mackness [deceased] and Karen H. Black [k.black@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, NSW, 2052, Australia; Gilbert J. Price [g.price1@uq.edu.au], Department of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia.     

Pragian and lower Emsian (Lower Devonian) conodonts from Liujing,Guangxi, South China

LU, J.-F., QIE, W.-Q. & CHEN, X.-Q., July 2016. Pragian and lower Emsian (Lower Devonian) conodonts from Liujing, Guangxi, South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

Lower Devonian (Pragian and Emsian) conodonts are described from the Liujing section in Guangxi, South China, highlighting the phylogeny of early polygnathids and establishing the basis for faunal correlations with deposits throughout the world. Records of Eognathodus kuangi sp. nov. and E. nagaolingensis Xiong increase the biodiversity of the Pragian eognathodids in South China and represent intermediate stages and probably the latest eognathodids in the evolutionary succession from Eognathodus to Polygnathus. During this evolutionary succession, the degeneration of the sulcus (or the flattening of the upper platform surface) is the most important morphological change, especially in the eognathodid lineage. Polygnathus trilinearis, P. pireneae and P. sokolovi are recognized together for the first time in South China. The contemporaneous occurrences of the kitabiformis and sokoloviformis morphs of P. pireneae with P. sokolovi and their respective similarities to P. kitabicus and P. sokolovi suggest that the latter two species are phylogenetically linked with P. pireneae. Evidence from the Liujing section also favours Polygnathus probably having evolved from Eognathodus in a tropical or subtropical area.

Jian-feng Lu* [lujfivan@sina.cn] and Xiu-qin Chen [xqchen@nigpas.ac.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Wen-Kun Qie [wkqie@nigpas.ac.cn], Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China. *Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China.     

First records of Geinitziidae (Insecta: Grylloblattida) from the Upper Triassic of Argentina (Mendoza)

Lara, M.B. & Aristov, D., August 2016. First records of Geinitziidae (Insecta: Grylloblattida) from the Upper Triassic of Argentina (Mendoza). Alcheringa 41, xxx–xxx. ISSN 0311-5518

A new grylloblattid (Permoshurabia argentina sp. nov.: Geinitziidae) is described and illustrated from the Upper Triassic of Argentina. The material represents the first record of this family from Argentina and expands the geographic distribution of this group during the Triassic.

María Belén Lara [lara.maria.belen@live.com.ar], Area Paleontología (Centro de Ecología Aplicada del Litoral-Universidad Nacional del Nordeste-Consejo Nacional de Investigaciones Científicas y Técnicas), Casilla de Correo 128, 3400 Corrientes, Argentina; Danil Aristov [danil_aristov@mail.ru], Borissak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya str. 123, Moscow, 117997, Russia.     

A new petauroid possum (Marsupialia,Pseudocheiridae) from the Pliocene Bluff Downs Local Fauna,northern Queensland

Pseudochirops winteri n. sp. is described from the Pliocene Bluff Downs Local Fauna on the basis of a left dentary fragment with M₂. It is considered to be most similar to P. archeri although its phylogenetic position must remain tentative until further material is found. The presence of this presumably rainforest-dwelling possum, the first from this fauna, may indicate a limited closed forest component. Given the paucity of additional rainforest taxa recovered from this site despite extensive screen washing, however, such a component must have been restricted to gullies or riparian fringes.     

A new Triassic Palaeolimnadiopsis (Crustacea: Conchostraca) from the Sydney Basin,New South Wales

Palaeolimnadiopsis bassi sp. nov. is described from the mid-Triassic Hawkesbury Sandstone of the Sydney Basin, New South Wales. It is distinguished from other species of the genus by its paired growth lines, moderately large size, nonterminal umbo, and small larval valve. Macrolimnadiopsis, Pteriograpta, and Limnadia (Falsisca) are considered to be synonymous with Palaeolimnadiopsis, and Belgolimnadiopsis with Euestheria.     

A short-snouted trematosauroid (Tetrapoda,Temnospondyli) from the Early Triassic of Australia: the oldest known trematosaurine

The trematosauroid temnospondyl Tirraturhinus smisseni gen. et sp. nov. from the Arcadia Formation of central Queensland, Australia, is described on the basis of its rostrum. This is the first trematosaurine (short-snouted) trematosauroid from Australia, and is considered to be most closely related to Tertrema acuta from Spitzbergen. Tirraturhinus smisseni occurs alongside lonchorhynchine (long-snouted) trematosauroids in the Arcadia Formation; the co-occurrence of both trematosauroid morphotypes in that fauna is repeated in a number of non-marine Early Triassic faunas elsewhere in Pangaea. The Arcadia Formation is probably Griesbachian (earliest Triassic), so that T. smisseni is the oldest known trematosaurine.     

Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted,early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia

Chamberlain, P.M., Travouillon, K.J., Archer, M. & Hand, S.J., November 2015. Kutjamarcoot brevirostrum gen. et sp. nov., a new short-snouted, early Miocene bandicoot (Marsupialia: Peramelemorphia) from the Kutjamarpu Local Fauna (Wipajiri Formation) in South Australia. Alcheringa 40, XX–XX. ISSN 0311-5518.

A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.

Philippa M. Chamberlain [philippa.chamberlain@uq.net.au], School of Earth Sciences, University of Queensland, St Lucia, Queensland 4072, Australia; Kenny J. Travouillon [kenny.travouillon@museum.wa.gov.au; kennytravouillon@hotmail.com], Western Australian Museum, Locked Bag 49, Welshpool DC, WA, 6986, and School of Earth Sciences, University of Queensland, St Lucia, Queensland, 4072, Australia; Michael Archer [m.archer@unsw.edu.au] and Suzanne J. Hand [s.hand@unsw.edu.au], School of Biological, Earth and Environmental Sciences, University of New South Wales, New South Wales, 2052, Australia.     

New ontogenetic information on Duyunaspis duyunensis Zhang & Qian in Zhou et al., 1977 (Trilobita,Corynexochida) from the Cambrian and its possible sexual dimorphism

Lei, Q.P., September .2015. New ontogenetic information on Duyunaspis duyunensis Zhang & Qian in Zhou et al., 1977 (Trilobita, Corynexochida) from the Cambrian and its possible sexual dimorphism. Alcheringa 40, XXX–XXX. ISSN 0311-5518.

The hypostomal condition and the complete ontogenetic development of Duyunaspis duyunensis Zhang & Qian in Zhou et al., 1977 are restudied on the basis of abundant specimens from the Balang Formation (Cambrian, Qiandongian) in Zila Village, Paiwu Township, Huayuan County, Hunan Province. The relatively complete ontogenetic series (degree 0 to 9) provides new evidence that the holaspis of this species has nine rather than seven thoracic segments as proposed by McNamara et al. (2006). Comparisons between Duyunaspis duyunensis and two associated species, Arthricocephalus chauveaui and Changaspis elongata, support its placement in the Subfamily Oryctocarinae. Moreover, based on observations of a large number of specimens, the presence of an anterior indentation on the pygidial end is possibly representative of sexual dimorphism.

Qianping Lei [cicelyapple@126.com], Natural Department, Changzhou Museum, Changzhou, Jiangsu Province, 213022, PR China; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China.     

Phylogenetic analysis of a new fossil Notonectidae (Heteroptera: Nepomorpha) from the Late Jurassic of China

Zhang W.T., Yao Y.Z. & Ren D., June 2012. Phylogenetic analysis of a new fossil Notonectidae (Heteroptera: Nepomorpha) from the Late Jurassic of China. Alcheringa, 239–250. ISSN 0311-5518.

A new fossil species Notonecta vetula sp. nov. is described and illustrated using nymph and adult fossil specimens collected from the Upper Jurassic Chijinqiao Formation, Yumen City, Gansu Province, China. A phylogenetic analysis, based on a combination of fossil and extant backswimmers, was conducted to confirm the position of the new fossil within the Notonectidae.

Wei-ting Zhang [zhangweitinghao@163.com], Yun-zhi Yao* [yaoyz100@gmail.com] and Dong Ren [rendong@mail.cnu.edu.cn], Key Lab of Insect Evolution and Environmental Changes, Capital Normal University, Beijing 100048, PR China; *Corresponding author; also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS), Nanjing 210008, PR China. Received 13.7.2011; revised 19.9.2011, accepted 27.9.2011.     

Morphological analysis of Ozarkodina prolata Mawson and Ozarkodina midundenta (Wang & Ziegler) (Emsian conodonts) from South China

Lu, J.-F., November 2015. Morphological analysis of Ozarkodina prolata Mawson and Ozarkodina midundenta (Wang & Ziegler) (Emsian conodonts) from South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

Investigations of conodonts from Emsian (Lower Devonian) strata at Bahe, Liujing and Daliantang in Guangxi and Yunnan, South China, provide new data on the morphological variability and phylogenetic affinity of Ozarkodina midundenta, a species initially assigned to Pandorinellina but transferred to Ozarkodina herein. Morphological analysis suggests that O. midundenta probably developed from O. prolata by progressive fusion of denticles in the middle third of the blade above the basal cavity in the Pa element.

Jian-feng Lu [lujfivan@sina.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China. Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China.     

Mecochirus Germar (Decapoda: Glypheoidea) in the Lower Cretaceous of Queensland

JELL, P.A., WOODS, J.T. & COOK, A.G., May 2017. Mecochirus Germar (Decapoda: Glypheoidea) in the Lower Cretaceous of Queensland. Alcheringa 41, 514–523 ISSN 0311-5518.

Three new species of glypheoid decapod crustaceans, Mecochirus mcclymontorum, M. bartholomaii and M. lanceolatus, are described from the late Aptian of the Eromanga, Carpentaria and Maryborough basins, respectively. The first two occur in the Doncaster Member of the Wallumbilla Formation and the last in the Maryborough Formation. This is the first record of Mecochirus Germar, 1827 or the Mecochiridae Van Straelen, 1925 in Australia and one of only a few Cretaceous occurrences of this largely Jurassic genus.

Peter A. Jell [amjell@bigpond.com], Jack T. Woods and Alex G. Cook [alex.cook@y7mail.com], School of Earth Sciences, The University of Queensland, St Lucia Queensland 4072, Australia.     

The taxonomic placement of the Miocene Patagonian frog Wawelia gerholdi (Amphibia: Anura)

Nicoli, L. Muzzopappa, P. & Faivovich, J., July 2016 The taxonomic placement of the Miocene frog Wawelia gerholdi (Amphibia: Anura). Alcheringa 40, xxx–xxx. ISSN 0311-5518.

The Miocene frog Wawelia gerholdi, from northern Patagonia, was previously attributed to Ceratophryidae, the clade of extant South American horned frogs, making it one of the oldest records known for this group. However, the only known specimen has juvenile traits and is strongly similar to the australobatrachian Calyptocephalella. We re-examined the morphology of W. gerholdi and provide additional anatomical information and comparisons with juvenile individuals of Ceratophryidae and Calyptocephalella gayi, the only extant species of Calyptocephalella. We reject a close relationship between W. gerholdi and Ceratophryidae. Furthermore, the holotype is indistinguishable from juveniles of C. gayi and shares with all the species of this genus a character state combination that seems unique among anurans. Therefore, we consider Wawelia to be a junior synonym of Calyptocephalella. The problematic taxonomic status of numerous fossil species of Calyptocephalella, together with its untested monophyly, also prompts us to provisionally consider the extinct species as a species inquirenda included in this genus, pending a comprehensive revision of all available remains.

Laura Nicoli [lnicoli@macn.gov.ar] and Julián Faivovich [julian@macn.gov.ar], División Herpetología, and Paula Muzzopappa [pmuzzopappa@macn.gov.ar], Sección Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’-CONICET, Av. Angel Gallardo 470, C1405DJR, Buenos Aires, Argentina.     

A reappraisal of Neocalamites and Schizoneura (fossil Equisetales) based on material from the Triassic of East Antarctica

Sphenophytes are a common floral element in the Triassic of Gondwana. Most sphenophyte compression fossils have been conventionally assigned to a few, presumably very widespread species of Neocalamites based on vegetative features of the stems (or pith casts) and the foliage. During recent decades, however, new reports on morphological and anatomical details of some of these fossils have cast doubt on the systematic affinities of many Gondwanan Triassic sphenophytes. Here we describe Neocalamites suberosus (Artabe & Zamuner) nov. comb. et emend. and Schizoneura africana Feistmantel emend. from several Triassic deposits in the central Transantarctic Mountains and Victoria Land, East Antarctica. The material enables a critical reevaluation of morphological and anatomical features that have been historically used to define the two genera, including leaf-base morphology, degree of leaf fusion, stem vasculature and vallecular canals, and features of the nodal diaphragm. The diagnoses of Neocalamites and Schizoneura are emended so that they more accurately reflect recent advances in our understanding of the anatomy and ontogeny of these plants.

[Benjamin Bomfleur [bbomfleur@ku]edu], Rudolph Serbet [serbet@ku.edu], Edith L. Taylor [etaylor@ku.edu] and Thomas N. Taylor [tntaylor@ku.edu], Department of Ecology and Evolutionary Biology, and Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA; Ignacio H. Escapa [iescapa@mef.org.ar], CONICET—Museo Paleontológico Egidio Feruglio, Trelew, Chubut 9100, Argentina. Received 4.7.2012; revised 22.12.2012; accepted 7.1.2013.

Bomfleur, B., Escapa, I.H., Serbet, R., Taylor, E.L. & Taylor, T.N., 2013. A reappraisal of Neocalamites and Schizoneura (fossil Equisetales) based on material from the Triassic of East Antarctica. Alcheringa 37, 1–17. ISSN 0311-5518.     

Late Triassic megaspores from the Amery Group,Prince Charles Mountains,East Antarctica

Cantrill, D. J., & Drinnan, A. N., 1994:03:28. Late Triassic megaspores from the Amery Group, Prince Charles Mountains, East Antarctica. Alcheringa 18, 71–78. ISSN 0311-5518.

Megaspores referable to the genera Cabochonicus Batten & Ferguson 1987 and Minerisporites Potonié 1956 are a common component of the palaeoflora recovered from the Jetty Member within the Flagstone Bench Formation of the Amery Group. The known ranges of these two genera, in conjunction with the macrofloral remains, suggest a Late Triassic age. Two new species, Cabochonicus sinuosus and Minerisporites triangulatus, are described. Scanning electron microscopic examination of the spores indicates that standard palynological treatment of megaspores can result in sculptural degradation, possibly leading to incorrect generic assignment.     

The radiation of early Silurian spiriferide brachiopods,with new taxa from the Llandovery of Iran

Popov, L.E. & Cocks, L.R.M., 2013. The radiation of early Silurian spiriferide brachiopods, with new taxa from the Llandovery of Iran. Alcheringa 38, 560–566. ISSN 0311–5518.

Although there were Late Ordovician spiriferides in the superfamily Cyrtioidea, namely Eospirifer and Odakella, only the former genus survived the terminal Ordovician extinction, and only Eospirifer is known from the earliest Llandovery (Rhuddanian). However, in the succeeding mid-Llandovery (Aeronian), the spiriferides radiated to include not only more species within Eospirifer and Striispirifer in the Eospiriferidae, but also the new genus Iranospirifer described here, which is the earliest representative of the other family within the superfamily, the Hedeinopsidae. The Ordovician species were confined to the South China continent and the Boshchekul volcanic island arc in Kazakhstan, but by the Aeronian the superfamily had spread westwards to various other continents, including the main Gondwanan superterrane (which included Iran) in the early Aeronian, and Avalonia-Baltica and Laurentia in the late Aeronian. The new species Eospirifer ghobadiae and Iranospirifer qarabilensis are both described from the lower Aeronian of Iran, and there is a note on the Aeronian rhynchonellide Stegocornu, which is endemic to Iran and nearby areas.

Leonid E. Popov [lepbarry@yahoo.co.uk; leonid.popov@museumwales.ac.uk] Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK. L. Robin M. Cocks [r.cocks@nhm.ac.uk], Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK. Received 29.5.2013, revised 12.7.2013, accepted 21.7.2013.