Placocystella in the Early Devonian (Lochkovian) of central Victoria

Jell, P.A., 2013. Placocystella in the Early Devonian (Lochkovian) of central Victoria. Alcheringa, 567–569. ISSN 0311–5518.

The South African allanicytidiid mitrate carpoid Placocystella africana (Reed) is recorded for the first time from Australasia having been collected from a Lochkovian bed of the Humevale Formation at Mooroolbark in eastern Melbourne. The Southern Hemisphere Allanicytidiidae incorporating five monospecific genera in Brazil, South Africa, Tasmania, Victoria and New Zealand is now known to have a species in common between South Africa and Victoria. The previously suggested synonymy of these five is revised to accept Placocystella, Tasmanicytidium, Allanicytidium and Australocystis (but not Notocarpos) as synonymous so that the family contains Placocystella with four species and monospecific Notocarpos.

Peter A. Jell [p.jell@uq.edu.au], School of Earth Sciences, The University of Queensland, St Lucia Queensland 4072, Australia. Received 10.4.2013; revised 6.6.2013; accepted 17.6.2013.     

Deciduous Nothofagus leaves from the Miocene of Cornish Head,New Zealand

Pole, M.S., 1994:03:28. Deciduous Nothofagus leaves from the Miocene of Cornish Head, New Zealand. Alcheringa 18, 79–83. ISSN 0311-5518.

Nothofagus leaves with plicate vernation, indicating a deciduous habit, are recorded from the Late Miocene of New Zealand. This is the first unequivocal record in New Zealand of deciduous Nothofagus, deciduous species now being restricted to Tasmania and South America.     

Notocarpos garratti gen. et sp. nov., a new Silurian mitrate carpoid from Victoria

Notocarpos garratti gen. et sp. nov. is described from the middle Ludlovian Humevale Formation of the Clonbinane district, Victoria. It is compared with similar anomalocystitid carpoids and is found to resemble most closely Allanicytidium flemingi Caster & Gill 1968 from the Early Devonian Reefton Beds of New Zealand. N. garratti provides evidence that anomalocystitids rested with the flattened thecal surface against the sea floor (i.e., an orientation opposite to that proposed by Jefferies, 1968). It is further suggested that the stele was adapted to provide a rearward mode of locomotion.     

Costulatotheca schleigeri (Hyolitha: Orthothecida) from the Walhalla Group (Early Devonian) at Mount Pleasant,central Victoria,Australia

Earp, C., 29 January 2019. Costulatotheca schleigeri (Hyolitha: Orthothecida) from the Walhalla Group (Early Devonian) at Mount Pleasant, central Victoria, Australia. Alcheringa. Alcheringa 43, 220–227. ISSN 0311-5518

A number of hyolith fragments (including one operculum), found in Early Devonian marine turbidites at Mt Pleasant, near Alexandra, central Victoria, are described as Costulatotheca schleigeri gen. et sp. nov., the first confirmed record of the order Orthothecida in the Devonian of Australia. Index fossils found at this locality (Uncinatograptus sp. cf. U. thomasi and Nowakia sp. ex gr. N. acuaria) indicate an age of Pragian or earliest Emsian. The taphonomy of rare rafted shelly fossils indicates that flysch deposition occurred in a very-low-energy environment into which there were occasional bursts of high-energy turbidites carrying allochthonous fossils from shallower water.

Clement Earp [omaiosys@yahoo.com.au], 1 De Havilland Place, Onerahi, Whangarei 0110, New Zealand.     

First occurrence of Janospira from the Early Ordovician of Australia

Cavanosteus, a new genus of homostiid arthrodire, with C. australis (McCoy) as type species, is erected for new material from the Early Devonian (Emsian) of the Burrinjuck area, NSW. This new material compares closely with the holotype from Buchan, Victoria, identified as the central plates from the skull; it includes lower jawbones (infragnathals) lacking denticulation also closely similar to those of Homostius from the Middle Devonian of the Baltic Province and Scotland. The only bone from the dermal trunk armour referred to the new genus is an anterior dorsolateral plate. Relationships of other large Early Devonian arthrodires assigned to the family Homostiidae are discussed. Cavanosteus gen. nov. is probably the closest relative to the type genus known from the Early Devonian.     

Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand)

Conran, J.G., Bannister, J.M. & Lee, D.E., 2013. Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand). Alcheringa 37, 496–509. ISSN 0311-5518.

Laurelia otagoensis sp. nov. Conran, Bannister & D.E. Lee (Laurales: Atherospermataceae) is described from the earliest Miocene Foulden Maar diatomite deposit, Otago, New Zealand. The new species is represented by mummified fossil leaves with well-preserved cuticle and associated clusters of achenes bearing persistent, long plumose styles. This basal angiosperm family is of significance because of its classic southern disjunctions and ecological importance in extant Gondwana-type rainforests, but has a very sparse fossil record. The present study describes one of very few convincing leaf fossils for Atherospermataceae and the only definitive fossil fruits. The presence of fossil Laurelia in Oligo–Miocene New Zealand combined with fossil leaf impressions from the late Eocene, Miocene dispersed cuticle and pollen from the Oligocene to Holocene shows that the family has had a long history in Cenozoic New Zealand. These new fossils also support palaeoclimatic data suggesting warmer conditions in the earliest Miocene of New Zealand.

John G. Conran [john.conran@adelaide.edu.au], Australian Centre for Evolutionary Biology & Biodiversity and Sprigg Geobiology Centre, School of Earth and Environmental Sciences, Benham Bldg DX 650 312, The University of Adelaide, SA 5005 Australia; Jennifer M. Bannister [jennifer.bannister@xtra.co.nz], Department of Botany, University of Otago, PO Box 56, Dunedin, New Zealand; Daphne E. Lee [daphne.lee@otago.ac.nz], Department of Geology, University of Otago, PO Box 56, Dunedin, New Zealand. Received 17.12.2012; revised 9.4.2013; accepted 15.4.2013.     

Thelodont fish and conodonts from the Early Devonian of Reefton,New Zealand

Thelodont scales and associated fragments of conodont elements identified as Turinia sp. cf. Turinia australiensis and Ozarkodina sp. cf. Ozarkodina buchanensis respectively are described for the first time from the Early Devonian Murray Creek Formation of Reefton, New Zealand. The conodont and thelodont components of the fauna suggest a late Pragian to early Emsian age for the assemblage.     

Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America

Camilleri, T.A., Warne, M.T., Holloway, D.J. & Weldon, E.A., 10 May 2019. Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America. Alcheringa XXX, X–X. ISSN 0311-5518.

Known occurrences of the ostracod genus Velibeyrichia are restricted to a number of Silurian to Lower Devonian geological strata in North America: the McKenzie Member of the Mifflintown Formation of Maryland and West Virginia; the Tonoloway Limestone of Maryland, West Virginia, Virginia and Pennsylvania; the Bloomsburg Formation of Maryland, Virginia and Pennsylvania; the Manlius Limestone of New York; and the Decker Limestone of New Jersey and New York. The genus includes six species: V. moodeyi (type species), V. mesleri, V. paucigranulosa, V. reticulosaccula, V. tonolowayensis and V. tricornia. The diagnostic combination of characters for this genus are: distinct deflection of the velum where it crosses the crumina in heteromorphs (adult female specimens), dorsal nodes on lobes L1 and L3, sexual dimorphism of the velum, and in tecnomorph specimens, either a shallow sulcus on lobe L3 or a zygal ridge (in adult tecnomorph specimens) extending from lobe L2 to lobe L3. The presence of one or the other of the latter two characters defines two distinct species groups.

Tamara T.A. Camilleri* [tamara.camilleri@deakin.edu.au], Mark T. Warne* [mark.warne@deakin.edu.au] and Elizabeth A. Weldon [l.weldon@deakin.edu.au], Deakin University, Geelong, School of Life and Environmental Sciences & Centre for Integrative Ecology (Melbourne Campus), 221 Burwood Highway, Burwood, Victoria 3125, Australia; David J. Holloway [dhollow@museum.vic.gov.au], Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia. *Also affiliated with: Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia.     

Shackleton Limestone archaeocyaths

A new monotypic genus of cheirurine trilobite, Azyptyx, is described based on the new species A. toongabbiensis from Early Devonian (late Lochkovian) limestones of the Wurutwun Formation near Toongabbie, Victoria. A feature of the genus that is unusual for the subfamily is a continuous basal glabellar furrow (S1) that does not normally meet the occipital furrow medially.     

Pterygotid eurypterid chelicera from the Lower Devonian of Victoria

A single eurypterid (Arthropoda: Chelicerata) chelicera, assigned to Acutiramus sp. cf. A. bohemicus, is described from the Wilson Creek Shale, Turtons Creek inlier, north of Foster, Victoria, Australia. The specimen comprises the proximal portion of both rami. This pterygotid chelicera supports an Early Devonian (?Lochkovian) age for the stratum at this locality, by comparison with occurrences of A. bohemicus from the Czech Republic and closely related species in northern Gondwana.     

The record of Araucariaceae macrofossils in New Zealand

Pole, M., December, 2008. The record of Araucariaceae macrofossils in New Zealand. Alcheringa 32, 405–426. ISSN 0311-5518.

The Araucariaceae have a long record in New Zealand, extending back to the Jurassic at least, and Araucaria extends back to at least the Late Cretaceous. This paper reviews the macrofossil record of the family and presents new information based largely on the leaf cuticle record. Agathis, which is the only genus of the family currently growing in New Zealand, has no record before the Cenozoic. All specimens previously identified from pre-Cenozoic strata clearly belong to other taxa or do not show characteristic features of the genus. Araucariaceae macrofossils are virtually ubiquitous in the Cretaceous assemblages of New Zealand but are conspicuous by their absence or rarity in Palaeocene deposits. Their demise may be an expression of events at the Cretaceous–Palaeogene boundary.     

An early Miocene ant (subfam. Amblyoponinae) from Foulden Maar: the first fossil Hymenoptera from New Zealand

Kaulfuss, U., Harris, A.C., Conran J.G. & Lee, D.E., 2014. An early Miocene ant (subfam. Amblyoponinae) from Foulden Maar: the first fossil Hymenoptera from New Zealand. Alcheringa 38, 568–574. ISSN 0311-5518.

The ant subfamily Amblyoponinae is presently represented in New Zealand by one endemic species in the cosmopolitan genus Stigmatomma and an introduced Australian species of Amblyopone. The fossil record of the group is restricted to two species of Stigmatomma from late Eocene Baltic Amber. Here, we describe the third fossil record, an Amblyopone-like specimen from the early Miocene of Otago, southern New Zealand, based on a winged male that resembles the extant A. australis Erichson in size, general habitus and characters of wing venation, but also shares features with the African amblyoponine genus Zymmer. This represents the first fossil record of Amblyoponinae from the Southern Hemisphere and the first example of Hymenoptera among the few pre-Quaternary insect fossils known from New Zealand. It suggests a long history of Amblyoponinae in New Zealand and Australia.

Uwe Kaulfuss [uwe.kaulfuss@otago.ac.nz] and Daphne E. Lee [daphne.lee@otago.ac.nz], Department of Geology, University of Otago, PO Box 56, Dunedin 9016, New Zealand; Anthony C. Harris [anthony.harris@otagomuseum.govt.nz], Otago Museum, PO Box 6202, Dunedin 9059, New Zealand; John G. Conran [john.conran@adelaide.edu.au], ACEBB & SGC, School of Earth and Environmental Sciences, The University of Adelaide, Benham Bldg, DX 650 312, Adelaide SA 5005, Australia. Received 18.3.2014; revised 15.5.2014; accepted 23.5.2014.     

Revision of the Early Devonian tabulate coral Pleurodictyum bifidum from New South Wales

Plusquellec, Y. &; Wright, A.J., October 2017. Revision of the Early Devonian tabulate coral Pleurodictyum bifidum from New South Wales. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

The tabulate coral Pleurodictyum bifidum Jones, 1944 Jones, O.A., 1944. Tabulata and Heliolitida from the Wellington district, N.S.W. Journal and Proceedings of the Royal Society of New South Wales 77, 33–39. [Google Scholar], from the Early Devonian (Pragian or lower Emsian) Garra Formation of central New South Wales, Australia is revised on the basis of the holotype and three other specimens. It is selected as the type species of the new monotypic genus Bifidomeria (Family Roemeriidae), which differs from Roemeria in its strictly cerioid corallum, its bifid septal spines and aspects of its microstructure. Study of the detailed microstructure of two other tabulate corals from the Devonian of New South Wales has led to the following revised generic assignments: Michelinia progenitor Chapman, 1921 Chapman, F., 1921. New or little known fossils in the National Museum. Part XXV—some Silurian tabulate corals. Proceedings of the Royal Society of Victoria 33, 212–225. [Google Scholar], previously assigned to Roemeripora, is assigned to Roemeria, and Holacanthopora clarkei Wright &; Flory, 1980 Wright, A.J. &; Flory, R.A., 1980. A new Early Devonian tabulate coral from the Mount Frome Limestone, near Mudgee, New South Wales. Proceedings of the Linnean Society of New South Wales 104, 211–219. [Google Scholar] is assigned to Michelinia.

Yves Plusquellec [yves.plusquellec@univ-brest.fr], Université de Bretagne Occidentale, CNRS-UMR 6538 ‘Domaines océaniques’, Laboratoire de Paléontologie, UFR Sciences et Techniques, 6 avenue Le Gorgeu, CS 98837, F-29283 Brest, France; Anthony Wright [tony.wright@optusnet.com.au], GeoQuEST Research Centre, School of Earth &; Environmental Sciences, University of Wollongong, Wollongong NSW 2522, Australia.     

A new Early Devonian lungfish from Guangxi,China, and its palaeogeographic significance

Qiao, T. & Zhu, M., 13.4.2015. A new Early Devonian lungfish from Guangxi, China, and its palaeogeographic significance. Alcheringa 39, xxx–xxx. ISSN 0311-5518

A new species of Cathlorhynchus (Dipnorhynchidae, Dipnoi) is described based on a mandible from the marine Yukiang Formation (early Emsian, Early Devonian) of Guangxi, southern China. It resembles the type species of Cathlorhynchus, C. trismodipterus, in that the anterior portion of the internal median septum terminates abruptly and does not contact dermal bones ventrally. The new Chinese form, together with Erikia jarviki from the Emsian of Yunnan, southern China, confirms the occurrence of the Dipnorhynchus lineage outside Australia. Coupled with the distribution of Westollrhynchus, Ichnomylax and Jessenia, we propose that the Dipnorhynchus lineage was dispersed widely during the early Emsian, corroborating the trans-Panthalassic distribution of early sarcopterygians.

Tuo Qiao [qiaotuo@ivpp.ac.cn] and Min Zhu [zhumin@ivpp.ac.cn], Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, PO Box 643, Beijing 100044, PR China.     

A Tremadocian asterozoan from Tasmania and a late Llandovery edrioasteroid from Victoria

Jell, P.A., 2014. A Tremadocian asterozoan from Tasmania and a late Llandovery edrioasteroid from Victoria. Alcheringa 38, 528–540. ISSN 0311-5518.

An asterozoan, Maydena roadsidensis gen. et sp. nov., is described from the mid-Tremadocian (La1b Zone) Florentine Valley Formation in southwestern Tasmania and is the oldest known asterozoan in the world. Although only a single, largely dissociated, specimen is available, enough is preserved to recognize distinctive ambulacral plates similar to those of Archegonaster Jaekel from the Llanvirn of the Czech Republic. Reciprocodiscus transambus n. gen., n. sp. is an isorophid edrioasteroid from the uppermost late Llandovery Springfield Formation exposed in the bed of Deep Creek, near Springfield, 65?km north northwest of Melbourne. It occurs with a low-diversity trilobite fauna indicating a deepwater, subphotic environment. This edrioasteroid has in each ambulacrum a single series of floor plates that are not visible on the oral surface, indicating its isorophid affinity and retains certain apparently primitive features that are not seen in post-Cambrian edrioasteroids, such as the very large plates of the marginal circlet, plated aboral surface and ambulacral tips extending onto the marginal circlet plates.

Peter A. Jell [p.jell@uq.edu.au] School of Earth Sciences, The University of Queensland, St Lucia Queensland 4072, Australia.     

Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia

Benson, R.B.J., Fitzgerald, E.M.G., Rich, T.H. & Vickers-Rich, P., 2013. Large freshwater plesiosaurian from the Cretaceous (Aptian) of Australia. Alcheringa 37, 1–6. ISSN 0311-5518

We report a large plesiosaurian tooth from the freshwater early–middle Aptian (Early Cretaceous) Eumeralla Formation of Victoria, Australia. This, combined with records of smaller plesiosaurian teeth with an alternative morphology, provides evidence for a multitaxic freshwater plesiosaurian assemblage. Dental and body size differences suggest ecological partitioning of sympatric freshwater plesiosaurians analogous to that in modern freshwater odontocete cetaceans. The evolutionarily plastic body plan of Plesiosauria may have facilitated niche differentiation and helped them to exclude ichthyosaurs from freshwater environments during the Mesozoic. However, confirmation of this hypothesis requires the discovery of more complete remains.

Roger B.J. Benson [roger.benson@earth.ox.ac.uk], Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK; Erich M.G. Fitzgerald [efitzgerald@museum.vic.gov.au], Thomas H. Rich [trich@museum.vic.gov.au], Museum Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia; Thomas H. Rich and Patricia Vickers-Rich [pat.rich@monash.edu], School of Geosciences, Monash University, Clayton, Victoria 3800, Australia. Received 30.10.2012; revised 27.1.2013; accepted 31.1.2013.     

New evidence for a Silurian (Ludlow) age for the earliest Baragwanathia flora

In the Silurian-Devonian marine clastic strata of the Yea District, central Victoria, two distinct flora and faunal assemblages containing elements of the Baragwanathia flora occur separated stratigraphically by 1700m of siltstone. The upper assemblage is Early Devonian (Praguian) in age, whilst the lower assemblage is considered to be Late Silurian (Ludlow).     

Bryozoans from the late Cretaceous Kahuitara Tuff of the Chatham Islands,New Zealand

Taylor, P.D., & Gordon, D.P., December, 2007. Bryozoans from the Late Cretaceous Kahuitara Tuff of the Chatham Islands, New Zealand. Alcheringa 31, 339-363. ISSN 0311-5518.

Fourteen bryozoan species are described from the Campanian – Maastrichtian Kahuitara Tuff of Pitt Island, substantially increasing the known diversity in this deposit from the two species recorded previously and making it the most diverse bryozoan biota yet described from the Cretaceous of Australasia. Nine of the Kahuitara Tuff bryozoans are cyclostomes, four are cheilostomes, and one is a shell-boring ctenostome. Seven new species are described: Ceriocava hakepaensis sp. nov., Tholopora australis sp. nov., Crisidmonea lanauzeorum sp. nov., Cookobryozoon cretacea sp. nov., Chiplonkarina preeceorum sp. nov. Chiplonkarina bifoliata sp. nov. and Aechmella rangiauriensis sp. nov. The remaining species are left in open nomenclature because of preservational deficiencies or lack of taxon-diagnostic gonozooids. The ctenostome family Cookobryozoidae is subsumed in the Terebriporidae. The new family Chiplonkarinidae is proposed for anascan cheilostomes previously assigned to the paraphyletic Electridae and distinguished by having primarily erect colonies with long, tubular zooids reminiscent of stenolaemates. None of the Kahuitara Tuff bryozoan species is known elsewhere, but all apart from one genus occur in roughly coeval deposits. No families regarded as particularly characteristic of the austral post-Cretaceous are evident. The relatively large number (three) of co-occurring species of Chiplonkarina is notable, as is the dominance of cyclostomes and the first record of Tholopora in the Southern Hemisphere.

Paul D. Taylor [p.taylor@nhm.ac.uk], Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, UK; Dennis P. Gordon [d.gordon@niwa.co.nz], National Institute of Water and Atmospheric Research, Private Bag 14-901, Kilbirnie, Wellington, New Zealand; received 7.3.2006, revised 3.9.2006.