An Early Permian brachiopod fauna of Gondwanan affinity from the Baoshan block,western Yunnan,China

The present paper describes and illustrates an Early Permian brachiopod fauna collected from two localities from the upper part of the type Dingjiazhai Formation near Youwang, 30 km south of Baoshan in the Baoshan block, western Yunnan, China. The brachiopod fauna is dominated by Stenoscisma sp. and Elivina yunnanensis sp. nov. and exhibits strong generic and some specific links with faunas from the Bisnain assemblage of Timor and the Callytharra Formation of Western Australia and, to a lesser extent, faunas from the Jilong Formation of southern Tibet, the Tashkazyk Formation of southeastern Pamir, the lower Toinlungkongba Formation of northwestern Tibet, the upper Pondo Group of central Tibet, and the Jimba Jimba Calcarenite of the Carnarvon Basin, Western Australia. Based on these correlations, a Late Sakmarian (Sterlitamakian) age is preferred for the Dingjiazhai brachiopod fauna. Two new species are proposed: Globiella youwangensis sp. nov. and Elivina yunnanensis sp. nov.     

Ngapakaldia bonythoni (Marsupialia,Diprotodontidae): new material from Riversleigh,northwestern Queensland,and a reassessment of the genus Bematherium

A latest Permian (late Changhsingian) radiolarian fauna is recorded from the upper Talung Formation, Hushan, Nanjing, Jiangsu Province, South China. This fauna includes 24 species belonging to 16 genera; new species are Albaillella hushanensis, Copicyntroides stellatus and Trilonche crassus. The presence of the radiolarian fauna and its taxonomic composition reveal that the Eastern Qinling-Dabie deep sea, which was located along the northern margin of the northeastern Yangtze Basin, persisted at least until the end of the Palaeozoic and that the collision between the North China and South China plates had not occurred by the end of the Permian.     

Late Silurian and Early Devonian biostratigraphy in the Hill End Trough and the Limekilns area,New South Wales

Precordilleran Late Palaeozoic brachiopod genera considered for this palaeobiogeographical analysis belong to four faunal associations: the low diversity Early Carboniferous (Tournaisian) Protocanites scalabrinii-Azurduya chavelensis Zone; the Late Carboniferous (Bashkirian-Moscovian) Levipustula fauna, which appears in sequences associated with glacial intervals; the latest Carboniferous-earliest Permian (Gzhelian-Asselian) Rhipidomella-Micraphelia and Tuberculatella-Aseptella associations, and the Early Permian (mid to late Asselian) Tivertonia jachalensis-Streptorhynchus inaequiornatus Biozone. The brachiopod affinities of the Precordillera are compared with the biogeographical Late Palaeozoic regions previously suggested by other authors using cluster analysis. During the Bashkirian-Moscovian, the Precordilleran faunas show a high affinity with the Austral Realm because of the presence of the Levipustula faunal elements in Eastern Australia. At the same time the affinity with the central and North American Regions is very low. The Early Permian Precordilleran faunas demonstrate highest affinity with the Gondwanan Realm with several genera also linked to the Tethyan and Boreal Realms. After the Late Carboniferous glacial episodes that affected the southwestern Gondwanan margin, the Precordilleran region underwent climatic amelioration. Subsequently, the Early Permian Precordilleran brachiopod assemblages are characterized by typical cool to cold water genera widely developed in the core Gondwanan Realm, with fewer genera (such as Kochiproductus, Rhynchopora and Neochonetes) suggesting a warmer water influence. The Precordilleran faunas demonstrate an increase in brachiopod diversity from the Carboniferous to the Permian, related to water temperature changes and to the palaeogeographical evolution of the southwestern Gondwanan margin and the movement of Gondwana across the South Pole.     

Conodont fauna and biostratigraphy of the Honghuayuan Formation (Early Ordovician) of Guizhou,South China

Serratognathus diversus An, Cornuodus longibasis (Lindström), Drepanodus arcuatus Pander, and eleven other less common conodonts, including Cornuodus? sp., Oistodus lanceolatus, Protopanderodus gradatus, Protoprioniodus simplicissimus, Juanognathus variabilis, Nasusgnathus dolonus, Paltodus? sp., Scolopodus houlianzhaiensis, Semiacontiodus apterus, Semiacontiodus sp. cf. S. cornuformis and Serratognathoides? sp., are described and illustrated from the Honghuayuan Formation in Guizhou, South China, concluding revision of the conodont fauna from this unit, which comprises 24 species in total. The most distinctive species in the fauna, S. diversus, consists of a trimembrate apparatus, including symmetrical Sa, asymmetrical Sb and strongly asymmetrical Sc elements. This species concept is supported by the absence of any other element types in a large collection represented by nearly 500 specimens of this species. The fauna indicates a late Tremadocian to mid-Floian age (Early Ordovician) for the Honghuayuan Formation, which was widely distributed on the Yangtze Platform in shallow water environments. Previously published biostratigraphic zonations for the Honghuayuan Formation are reviewed, and revised on the basis of our knowledge of the entire conodont fauna, supporting the establishment of three biozones, Triangulodus bifidus, Serratognathus diversus, and Prioniodus honghuayanensis biozones in ascending order. Species of Serratognathus enable correlation between Ordovician successions of South China, North China (North China Platform and Ordos Basin), Tarim Basin, and further afield into Malaysia and northwestern Australia.     

A new Late Permian brachiopod fauna from Tanggula,Qinghai-Tibet Plateau and its palaeogeographical implications

Weihong He, Jianjun Bu, Zhijun Niu & Yang Zhang, June, 2009. A new Late Permian brachiopod fauna from Tanggula, Qinghai-Tibet Plateau and its palaeogeographical implications. Alcheringa 33, 113–132. ISSN 0311-5518.

A brachiopod fauna described from the Late Permian of the Gongri and Lizai villages, northwest of the Qoima Co Lake, Tanggula area, southern Qinghai, northwest China, includes ten species in nine genera. This fauna from the (Qiangtang Block) has a strong affinity to coeval faunas of South China, sharing 74% of its species. In addition, the Late Permian Tanggula brachiopod assemblages also demonstrate a clear link with the Middle Permian brachiopod faunas of neighbouring microcontinents including the Indochina block (Malaysia and Cambodia) and the Shan-Thai block (Thailand), as suggested by the presence of Caricula sp. cf. C. salebrosa, Transennatia termierorum and Strophalosiina. This phenomenon indicates that there were relatively narrow seaways between these microcontinents to enable ready interchange of brachiopods during the Permian, and that brachiopods tended to retreat towards the palaeoequatorial region throughout the period.     

Early Carboniferous brachiopod faunas from the Baoshan block,west Yunnan,southwest China

38 brachiopod species in 27 genera and subgenera are described from the Yudong Formation in the Shidian-Baoshan area, west Yunnan, southwest China. New taxa include two new subgenera: Unispirifer (Septimispirifer) and Brachythyrina (Longathyrina), and seven new species: Eomarginifera yunnanensis, Marginatia cylindrica, Unispirifer (Unispirifer) xiangshanensis, Unispirifer (Septimispirifer) wafangjieensis, Brachythyrina (Brachythyrina) transversa, Brachythyrina (Longathyrina) baoshanensis, and Girtyella wafangjieensis. Based on the described material and constraints from associated coral and conodont faunas, the age of the brachiopod fauna from the Yudon Formation is considered late Tournaisian (Early Carboniferous), with a possibility extending into earlyViséan.     

Pennsylvanian (Carboniferous) brachiopods from the Itaituba Formation of the Amazon Basin,Brazil

A new brachiopod fauna is described from the lower Itaituba Formation at the Caima Quarry 2 section in the Itaituba area, Amazon Basin, Brazil. The Amazonoproductus amazonensis-Anthracospirifer oliveirai Assemblage is proposed for this fauna, which is considered early Pennsylvanian (Morrowan) as constrained by associated conodont and fusulinacean faunas. Nine brachiopod taxa are described herein, including Amazonoproductus amazonensis gen. et sp. nov., and Buxtonioides itaitubensis sp. nov. and Linoproductus caima sp. nov. The new tribe Marginovatini of the Linoproductoidea (the Productida) is also proposed.     

A new early Silurian brachiopod genus,Thulatrypa, from Norway and South China,and its palaeobiogeographical significance

Huang, B., Baarli, B.G., Zhan, R.B. & Rong, J.Y., October 2015. A new early Silurian brachiopod genus, Thulatrypa, from Norway and South China, and its palaeobiogeographical significance. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

The smooth atrypoid brachiopod Thulatrypa gen. nov. incorporates two species, a younger (T. gregaria) from Norway, and an older (T. orientalis) from South China, which collectively span the middle Rhuddanian through Aeronian. In Baltica, the genus thrived just below the storm wave base in a tropical BA4 setting extending slightly into BA3 and BA5 respectively, whereas in South China, its representative occurs in a much shallower assemblage (BA2–3). Their palaeobiogeographical implications are carefully investigated. This study supports the arguments that Thulatrypa may have originated in South China in the middle Rhuddanian and extended its range to eastern Baltica in the late Rhuddanian. Larvae may have drifted along a channel from the east to the southwest of Baltica, which supports the reconstructions of palaeocurrents in the early Silurian in previous palaeogeographical studies.

Bing Huang [bhuang@nigpas.ac.cn], Ren-bin Zhan [rbzhan@nigpas.ac.cn] and Jia-yu Rong [jyrong@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, PR China; B. Gudveig Baarli [gudveig.baarli@williams.edu], Department of Geosciences, Williams College, Williamstown, MA 01267, USA.     

Foraminiferal biostratigraphy of the Bei’an Formation (Visean–Serpukhovian) in the Pengchong area of Liuzhou,Guangxi, South China

Shen, Y. & Wang, X.L., 24.6.2015. Foraminiferal biostratigraphy of the Bei’an Formation (Visean–Serpukhovian) in the Pengchong area of Liuzhou, Guangxi, South China. Alcheringa 39,559–572 ISSN 0311-5518

The type section of the Mississippian Bei’an Formation is well exposed on both sides of the Beihuan expressway at Liuzhou in Guangxi, South China, about 700 m east of the Pengchong GSSP (Global Boundary Stratotype Section and Point) for the base of the Visean Stage. Foraminifers are abundant and diverse (at least 59 species and 39 genera) in this section, and five foraminiferal zones are recognizable in the Bei’an Formation. In ascending order, these are the Pojarkovella nibelis, Koskinotextularia, Bradyina, Janischewskina and ‘Millerella’ tortula zones. Based on the foraminifers, the Bei’an Formation is regarded as being of middle Visean to Serpukhovian age, i.e., equivalent to biozones MFZ12–MFZ15 from the stratotypes of Europe, and the Visean–Serpukhovian boundary is placed at the base of the ‘Millerella’ tortula Zone.

Yang Shen [shenybj@126.com] (Corresponding author) and Xun-Lian Wang [wxl@cugb.edu.cn], School of Earth Sciences and Resources, China University of Geosciences (Beijing), 29 Xueyuan Road, Beijing 100083, PR China.     

A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida

Zhang, Y., He, W.H., Shi, G.R., Zhang, K.X. & Wu, H.T., 26.2.2015. A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida. Alcheringa 39, xxx–xxx. ISSN 0311-5518.

As the third and last part of a systematic palaeontological study of the brachiopod fauna from the Permian–Triassic boundary section at Zhongzhai in Guizhou Province (South China), this paper reports 15 species (including three new species: Tethyochonetes minor sp. nov., Neochonetes (Zhongyingia) transversa sp. nov., Paryphella acutula sp. nov.) in Order Productida. In addition, the morphological features and definitions of several key Changhsingian brachiopod taxa (e.g., Paryphella and Oldhamina interrupta) are clarified and revised.

Yang Zhang* [zhangy05@163.com] and G.R. Shi [guang.shi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; Weihong He [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Huiting Wu [ht_wu415@163.com], Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China. *Also affiliated with: Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

Late Devonian (Famennian) brachiopod Yunnanella from the Junggar Basin,northern Xinjiang,China

The Late Devonian (Famennian) brachiopod Yunnanella is reported from the western Junggar Basin, northern Xinjiang, on the basis of Yunnanella hanburii (Davidson) and Yunnanella sp.     

Late Carboniferous marine invertebrate zones of eastern Australia

Brachiopods of the Marginirugus barringtonensis and Levispustula levis Zones comprise the youngest of three major Carboniferous biostratigraphic units in eastern Australia. The remarkable change in the composition of marine invertebrates between the second and third of these units, a transition that only two brachiopod species are known to survive, has been attributed to the isolation of eastern Australia and to a drop in temperature. The Marginirugus barringtonensis and Levipustula levis Zones are redefined from reference sections in the Myall region, N.S.W.; the L. levis Zone now includes the Syringothyris bifida Zone of Campbell (1961). Faunal sequences show that these two zones are closely related and that there is no hiatus between them, as formerly proposed by McKellar (1965). Conodonts identified by Mr. D. Crane indicate a late Viséan to early Namurian age for the Marginirugus barringtonensis Zone, and an age of early Namurian for the base of the Levipustula levis Zone; brachiopod evidence indicates that the L. levis Zone extends into the Westphalian. The younger Auriculispina levis Zone from the Yarrol Basin may be coeval with the Trigonotreta campbelli Zone from N.S.W. Faunas from both the L. levis and A. levis Zones are present in Argentina and many species from the L. levis Zone have affinities with forms from the Baikal region, U.S.S.R. Correlations based on the zones have eliminated the need for a hiatus between the Branch Creek and Baywulla Formations in the Yarrol Basin.

Taxa described from the Marginirugus barringtonensis and Levipustula levis Zones in the Myall region include Yagonia gibberensis Roberts gen. et sp. nov., Bulahdelia myallensis Roberts gen. et sp. nov., Alispirifer yagonensis Thompson sp. nov., A. alatus Thompson sp. nov., Licharewia bootiensis Thompson sp. nov., Spirifer pristinus (Maxwell), Spiriferi sp., Neospirifer campbelli Maxwell and N. senilis Maxwell.     

Carboniferous postglacial faunas in the late Serpukhovian–Bashkirian interval of central-western Argentina

Cisterna, G.A., Sterren, A.F., López Gamundí, O. & Vergel, M.M., March 2017. Carboniferous postglacial faunas in the late Serpukhovian–Bashkirian interval of central-western Argentina. Alcheringa, ISSN 0311-5518.

Typical glacial–postglacial sequences associated with the Late Palaeozoic Ice Age (LPIA) are recognized in the Calingasta-Uspallata Basin, central-western Argentina, particularly in the Hoyada Verde and El Paso formations (late Serpukhovian–Bashkirian) at Barreal Hill (San Juan province). Brachiopods and bivalves accompanied by gastropods, conulariids, nautiloids, corals and ostracods constitute the marine assemblages of the El Paso Formation. They are assigned to the Aseptella–Tuberculatella/Rhipidomella–Micraphelia (AT/RM) fauna, characterized by two fossil assemblages: Aseptella–Tuberculatella, identified in the lower fossiliferous interval, and Rhipidomella–Micraphelia in the upper. The development of the different invertebrate assemblages within the El Paso Formation, and their relationship with coeval suite in the Hoyada Verde Formation, can be explained by a complex array of abiotic factors (substrate stability, turbidity, nutrient availability, variation in oxygen levels, poor circulation and salinity variations in the water column) that were directly related to glacial retreat dynamics and coastal configuration. A restricted palaeofjord setting is proposed for the depositional environment of the El Paso Formation in contrast to an exposed open marine coast with a gently sloping shelf for the Hoyada Verde Formation. The study of the postglacial fauna of the El Paso Formation and its relationship with the Levipustula fauna in the Calingasta-Uspallata Basin, help determine the main controls on the distribution of the postglacial faunas in other late Palaeozoic South American basins, such as the Tepuel Genoa Basin in Patagonia and the Tarija Basin in Bolivia.

Gabriela A. Cisterna [gabrielacisterna@conicet.gov.ar], CONICET-UNLAR, Av. Dr. Luis M. de la Fuente s/n, La Rioja, 5300, Argentina; Andrea F. Sterren [asterren@unc.edu.ar], CICTERRA (CONICET – Universidad Nacional de Córdoba), Av. Vélez Sarsfield 1611, X5016GCA, Córdoba, Argentina; Oscar López Gamundí [orlg2003@yahoo.com], P1C Consultants 1121 Banks Street, Houston, TX 77006, USA; María del Milagro Vergel [vergelmar@tucbbs.com.ar], CONICET – INSUGEO – Facultad de Ciencias Naturales e Instituo Miguel Lillo (UNT), Miguel Lillo 205, 4000, San Miguel de Tucumán, Argentina.     

Late Silurian pentameride brachiopods from Yass and Molong,New South Wales

The five known species of pentameride brachiopods from the Yass Syncline Ludlow (LateSilurian) succession, belonging to the superfamilies Pentameroidea, Gypiduloidea and Clorindoidea, are fully revised; no new species are recognised. The pentameroids Conchidium sp. cf. hospes and Aliconchidium yassi are confined to the Bowspring Limestone Member (Silverdale Formation). The gypiduloid Ascanigypa glabra and externally homeomorphic clorindoids Barrandina wilkinsoni and Clorinda minor replace them in the overlying Barrandella Shale Member, the last two extending into the Yarwood Siltstone Member (Black Bog Shale). Clorinda minor is also possibly present in the Rainbow Hill Member (Rosebank Shale). All except C. minor are uncommon to rare components of the Yass brachiopod fauna. Clorinda molongensis, a species of uncertain mid- to late Silurian age from the Molong Limestone, is also revised. Aliconchidium and Barrandina are known only from Yass, whereas Clorinda is cosmopolitan. Conchidium alsois widespread, but C. hospes is a species from the Prague Basin probably also known from the Urals and the Tien Shan. Ascanigypa is another Prague Basin taxon, recently recognised in Arctic Canada.     

Ordovician trilobite faunas and depositional history of the Taebaeksan Basin,Korea: implications for palaeogeography

The Taebaeksan Basin occupies the central-eastern part of the Korean peninsula and was a low-relief shallow marine carbonate shelf on which the Cambro-Ordovician Choson Supergroup was deposited. In the Taebaeksan Basin trilobites are among the most dominant fossil groups in the Lower Ordovician, but they become less important in Middle Ordovician faunal assemblages. The Early Ordovician trilobite faunas of the Taebaeksan Basin are characterised by the common occurrence of pandemic genera such as Jujuyaspis, Leiostegium, Asaphellus, Protopliomerops, Hystricurus, Apatokephalus, Shumardia, Asaphopsoides, and Kayseraspis. Biogeographically significant trilobite taxa include Yosimuraspis, Dikelokephalina, Koraipsis, and Chosenia. These Ordovician trilobite faunas, which thrived in shallow marine environments, show a remarkable similarity with faunas from North China, implying that the Taebaeksan Basin was connected through contiguous shallow waters to North China. These Sino-Korean faunas exhibit a close biogeographic connection with Australian faunas, with which they share some endemic genera, whereas they are more distantly related to the faunas of South China, South America, and North America. Based on these palaeobiogeographical features, it is suggested that in the early Palaeozoic much of the present Korean peninsula including the Taebaeksan Basin belonged to the Sino-Korean block, while part of the peninsula was derived from the Yangtze block.     

Fauna and biostratigraphy of the Cambrian (Series 2, Stage 4; Ordian) Tempe Formation (Pertaoorrta Group), Amadeus Basin,Northern Territory

A new faunal assemblage is reported from the Tempe Formation (Cambrian Series 2, Stage 4; Ordian) retrieved from the Hermannsburg 41 drillcore, Amadeus Basin, central Australia. Two trilobite taxa, including one new species Gunnia fava sp. nov., four brachiopod taxa, including the age-diagnostic Karathele napuru (Kruse), Kostjubella djagoran (Kruse) and Micromitra nerranubawu Kruse, together with a bradoriid, helcionellids, hyoliths, echinoderms, chancelloriids, sponges and problematic tubes are described. The fauna has close links to those of the neighbouring Daly, Georgina and Wiso basins and suggests that the Tempe Formation correlates with the Australian Ordian stage (either the Redlichia forresti or Xystridura negrina assemblage zones). The Giles Creek Dolostone in the eastern Amadeus Basin, previously regarded as coeval with the Tempe Formation, has recently been reported to be of early Templetonian age in its type section. The described taxa from the Tempe Formation confirm that these two sedimentary units are not contemporaneous and that regional stratigraphic schemes should be amended.     

Permian brachiopods from the Tupe Formation,San Juan Province,Precordillera, Argentina

The brachiopod fauna from the Tupe Formation at La Herradura Creek, located on the west flank of Perico Hill, San Juan Province, Argentina, palaeogeographically belongs to the western sector of the Paganzo basin (‘Guandacol embayment’). The stratigraphical section of the Tupe Formation at La Herradura Creek is the stratotype of the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone, was previously regarded as being of Late Carboniferous age but here is assigned to the earliest Permian (Asselian). We describe and review the biozone assemblage, which consists of Streptorhynchus inaequiornatus, Tivertonia jachalensis, Kochiproductus sp., Costatumulus sp., Coronalosia argentinensis, Tupelosia paganzoensis, Trigonotreta pericoensis, Septosyringothyris sp. aff. Septosyringothyris jaguelensis and Crurithyris? sp. This brachiopod assemblage is related to Indian and Australian Early Permian faunas and its presence in the La Herradura Creek section provides new evidence in support of an Asselian (Early Permian) age for the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone. This assemblage is also important for intra- and inter-basinal correlation because several of its characteristic species have been identified from other sections of the Paganzo basin and the Río Blanco basin. The proposed age for this biozone is consistent with the age of palynological data from slightly above the marine faunas from the stratotype locality.     

Wenlock brachiopods from Canberra,Australia

Twenty-four brachiopod species are described from a well-preserved shelly fauna in the Walker Volcanics, collected from the Molonglo valley west of Canberra. There are three inarticulates and twenty-one articulates, of which nine are new, three are closely compared with previously described species and twelve are under open nomenclature. New are Skenidioides thrinax, Salopina mediocostata, Aegiria (Epelidoaegiria subgen. nov.) chilidifera, Morinorhynchus oepiki, Australina (Australina) kausi, Coelospira cavata, Endospirifer anxius, Reticulariopsis silurica and Vadum (gen. nov.) coppinsense. The fauna is shown to be cosmopolitan in character and falls within the scope of Benthic Assemblages 3 to 5 of Boucot (1975). It is of Wenlock age.