Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia,China

SHI, C.F., WANG, Y.J., YANG, Q. & REN, D., September 2012. Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China. Alcheringa 36, 311–320. ISSN 0311-5518.

A new grammolingiid genus, Chorilingia containing four new species (C. euryptera, C. parvica, C. translucida and C. peregrina) is described and illustrated from the Jiulongshan Formation at Daohugou, Inner Mongolia, China. The new genus is differentiated mainly on the first branch of vein Rs (Rs1) separating distal to the forks of both CuA and CuP. A key to species of the genus is provided.     

Paracordyline aureonemoralis (Lomandraceae): an Eocene monocotyledon from South Australia

A monocotyledonous leaf macrofossil taxon from Golden Grove in Adelaide, South Australia is recognised as being close to several extant Australasian species of Cordyline, especially those in the C. stricta (Sims) Endl. / C. fruticosa (L.) A. Chev. complex. The fossil is assigned to the form genus Paracordyline, known previously from the Oligocene Kerguélen Islands. However, as the Golden Grove taxon differs markedly from the Kerguélen species, it is considered to be a new species, P. aureonemoralis Conran & Christophel.     

First occurrence of Janospira from the Early Ordovician of Australia

Cavanosteus, a new genus of homostiid arthrodire, with C. australis (McCoy) as type species, is erected for new material from the Early Devonian (Emsian) of the Burrinjuck area, NSW. This new material compares closely with the holotype from Buchan, Victoria, identified as the central plates from the skull; it includes lower jawbones (infragnathals) lacking denticulation also closely similar to those of Homostius from the Middle Devonian of the Baltic Province and Scotland. The only bone from the dermal trunk armour referred to the new genus is an anterior dorsolateral plate. Relationships of other large Early Devonian arthrodires assigned to the family Homostiidae are discussed. Cavanosteus gen. nov. is probably the closest relative to the type genus known from the Early Devonian.     

A new giant python from the Pliocene Bluff Downs Local Fauna of northeastern Queensland

Liasis dubudingala n. sp., described on the basis of isolated vertebrae from the Early Pliocene Bluff Downs Local Fauna, is the largest snake known from Australia. Dependance of vertebral proportions on intracolumnar position indicates that the fossil taxon can be excluded from the Morelia/Python clade. High neural spines suggest possible affinity with Liasis olivacea, whereas a posterior dentary fragment with small teeth is unlike L. olivacea and more similar to Liasis mackloti or species of Bothrochilus and Leiopython. As these extant species have all recently been treated as members of Liasis, the new species is assigned to that genus.     

A new fossil genus of forest wallaby (Marsupialia,Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea

A new macropodine genus and species, Silvaroo bila, is described from the Pliocene Chinchilla Sand of Queensland. The generic concept of Protemnodon is reviewed, and it is concluded that two Pliocene species previously placed in that genus (bandharr and buloloensis) belong in Silvaroo. Species of Silvaroo resemble the modern forest wallabies of Papua New Guinea (species of Dorcopsis and Dorcopsulus) and also bear close phenetic resemblance to the late Miocene Dorcopsoides fossilis. Forest wallabies are not known from mainland Australia after the middle Pliocene, but occur in the late Pliocene and Pleistocene of New Guinea.     

Redescription of Parexus recurvus,an Early Devonian acanthodian from the Midland Valley of Scotland

Parexus Agassiz was one of the first Early Devonian ‘spiny sharks’ to be described. The genus is readily recognized by the large size and ornament of its anterior dorsal fin spine. Although two species were erected, reappraisal of all known specimens indicate they should be synonymized in the type species Parexus recurvus. Farnellia tuberculata Traquair, originally described as a vertebral column, is actually tooth rows of jaw dentition, and is also now considered to be a junior synonym of P. recurvus. Parexus has a perichondrally ossified scapulocoracoid of typical acanthodian shape, and diagnostic features of the family Climatiidae, but has distinctive scales comprising appositional growth zones that closely resemble those of the putative stem chondrichthyan Seretolepis elegans Karatajute-Talimaa.     

New,early ostracods from the Ordovician (Tremadocian) of Iran: systematic,biogeographical and palaeoecological significance

New Tremadocian ostracod material from the Alborz Mountains of Iran confirms the early and widespread occurrence of the Ordovician genus Nanopsis, and the apparently simultaneous first appearance of ostracods in the fossil record at the level of the P. deltifer conodont biozone (485.5 Ma) from China to Argentina. Nanopsis pairidaeza sp. nov. adds to the pool of species diversity for the Early Ordovician, though documented Tremadocian ostracod generic diversity remains low, with only four genera. The presence of Early Ordovician ostracods in Alborz, their occurrence elsewhere in palaeocontinental Gondwana, Baltica and China coupled to their marked absence from the Tremadocian of Laurentia and Siberia, supports the notion of the earliest occurrence of ostracods centred on Gondwana/Baltica.     

Australian Cainozoic Bryozoa, 1: Nudicella gen. nov. (Onychocellidae,Cheilostomata): taxonomy,palaeoenvironments and biogeography

The new bryozoan genus Nudicella (Onychocellidae, Cheilostomata) is proposed to accommodate the common and widespread Australian Cainozoic cheilostome bryozoan Eschara clarkei Tenison Woods, which is redescribed and subdivided into four species: N. clarkei (Tenison Woods), N. cribriforma sp. nov., N. latiramosa sp. nov. and N. tenuis sp. nov. Cellaria gigantea Maplestone is also reassigned to Nudicella. Colonies of this genus display a wide variety of growth forms, including cribrate fenestrate, flat robust branching, foliose, delicate branching and encrusting; their occurrences correlate with changes in sedimentary facies and palaeoenvironments. The distinctive cribrate style of fenestrate growth form has evolved convergently in unrelated bryozoan groups at various geological intervals. It is found in a wide variety of sedimentary facies, as in other coexisting opportunistic genera such as Celleporaria, indicating a wide ecological tolerance. The oldest recorded occurrence of Nudicella is in the Paleocene of north western Australia. From there it appears to spread south in the Eocene and then east towards the Otway Basin in southeastern Australia, where it occurs in the Oligocene and Miocene; no post-Miocene representatives of this genus are yet known.     

A new species of Loricera Latreille from Eocene Baltic amber (Coleoptera: Carabidae: Loricerinae)

Cai, C., Liu Y. & Huang, D., February 2017. A new species of Loricera Latreille from Eocene Baltic amber (Coleoptera: Carabidae: Loricerinae). Alcheringa xx, xxx-xxx. ISSN 0311-5518.

Loricerinae is a small, distinctive subfamily of ground beetles, comprising only one genus Loricera Latreille. Only one fossil species is known to date. Here, we describe a new species, Loricera groehni sp. nov., belonging to Loricera based on a well-preserved adult in Eocene Baltic amber. Loricera groehni is tentatively attributed to the obsoleta group of the subgenus Loricera s.str. based on the relatively long antennomere 3 and punctate elytral interval 7. The discovery of a new species morphologically close to the extant Loricera species from western China and northern India suggests that the obsoleta group was more widespread in the Eocene than it is at present. The distribution pattern of Chinese Loricera is probably relictual. The fossil species, possessing conspicuous stiff setae on the basal antennomeres, was probably a specialized predator of springtails.

Chenyang Cai [cycai@nigpas.ac.cn], Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; Ye Liu [liuye198282@126.com], Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China; Diying Huang [dyhuang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.     

A numerical taxonomic study of Late Carboniferous Spiriferidina from eastern Australia

Species of Late Carboniferous spiriferidines belonging to the genera Alispirifer, Licharewia, Neospirifer, Spirifer, Liriplica, and Spinuliplica are analyzed on the basis of fifty-two different morphological characters. A computer program for cluster analysis is used which ensures that each character is given equal weighting, and the degrees of similarity between species are measured objectively.

The resulting classification is similar to the conventional one, with three exceptions. It is recommended that Spinuliplica spinulosa Campbell and Liriplica alta Campbell be assigned to one genus, Liriplica; Licharewia bootiensis Thompson be assigned to the genus, Alispirifer; and Spirifer pristinus (Maxwell) be reassigned to the genus Neospirifer.

A method of graphical analysis using contour diagrams is also presented as an alternative method of illustrating the relationships between the different species of spiriferidines.     

The distribution of the Mesozoic ostracod genus Procytherura Whatley: palaeogeographical implications with special reference to Argentina

A review is undertaken of the nine species of Procytherura known to occur in Argentina and a new species, Procytherura serangodes sp. nov. is described. The global distribution of the genus indicates that it was more or less equally diverse and widely distributed in both hemispheres during most of the Lower and Middle Jurassic, but that in the Upper Jurassic and in the Lower Cretaceous, it became progressively restricted to the Southern Hemisphere. The widespread distribution of several Jurassic species of Procytherura, which occur in both Great Britain and Northwest Europe and in Argentina, is shown to be related to the availability of important migration routes, including the Tethys and the Hispanic Corridor. In the Lower Cretaceous, the very widespread distribution of Procytherura in the Southern Hemisphere, is used to support the existence of important routes along the eastern and western seaboards of Africa, the latter associated with the opening of the South Atlantic.     

A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China

LI, Y.J., HAN, G., NEL, A. REN, D., PANG, H. & LIU. X.L., September 2012. A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China. Alcheringa 36, 321–324. ISSN 0311-5518.

The new petalurid species Pseudocymatophlebia boda is described from Lower Cretaceous strata of Inner Mongolia, China. It provides additional morphological characters for this genus, which has been previously recorded from the Lower Cretaceous of England. Together with Aktassia, it is the second aktassiid genus with a very wide distribution, even though this family remains known only from Eurasia. Furthermore, a new name, Brachaktassia gen. nov., is proposed to replace the brachiopod genus Aktassia Popov, 1976.     

Tertiary Nothofagus macrofossils from Cethana,Tasmania

Two new Tertiary species of Nothofagus from the Early Eocene-Oligocene deposit at Cethana represent the first reports of fossil species which are not closely related to the extant Australian species N. moorei and N. cunninghamii. N. cethanica sp. nov. is most closely related to the extant New Zealand species N. fusca and N. truncata and gives further evidence of the relatively slow evolution within this genus. The other specimen is indistinguishable from extant adult N. gunnii leaves, and has been assigned to that species. This fossil shows that the deciduous habit was probably already present in N. gunnii by the Oligocene, and this may have helped N. gunnii to survive the Late Tertiary/Quaternary glaciations. Juvenile N. gunnii foliage gives some insight into the possible origins of this species, which may have been from the same ancestral stock as N. fusca, N. truncata, and N. cethanica.     

The southernmost records of Anhingidae and a new basal species of Anatidae (Aves) from the lower–middle Miocene of Patagonia,Argentina

New bird fossils from the Santa Cruz Formation (lower–middle Miocene), Santa Cruz Province, Patagonia, Argentina, are described. They represent an indeterminate species of the extinct anhingid Macranhinga and a new genus and species of basal Anatidae Ankonetta larriestrai. The record of the giant darter Macranhinga constitutes the southernmost record for the family, and expands the known stratigraphic range of the genus, previously restricted to the upper Miocene. Based on an analysis of the fossil anhingid record from South America, we hypothesize that giant darters disappeared from South America in the early Pliocene due to climatic deterioration, regression of marine and freshwater environments, the arrival of placental carnivorous mammals, and also probably by competition with phalacrocoracid cormorants. The new anatid Ankonetta is based on an incomplete but informative tarsometatarsus, with superficial similarities to extant Dendrocygna. A brief overview of several fossil ducks from the Patagonian Cenozoic concludes that most pre-Pliocene examples belong to non-anatine taxa, indicating that plesiomorphic ducks were the dominant anseriforms in those times, a pattern also evident on other continents.     

The craniology and relationships of four species of bos 1. Basic craniology: Bos taurus L. and its absolute size

This paper is the first of a series on the craniology of four species of Bos. Here (and in the next paper) Bos taurus in Britain is used to establish a basic craniology for the genus. In the initial sections notes are made on the nomenclature, material available, history, and age assessment of B. taurus. The subsequent section deals with the effects of sex, age and breed on absolute size.The results show that Bos taurus is very variable, particularly in the length of the horncore. Sex influences certain dimensions very greatly, particularly those of breadth. Although the horns of cows are often longer than those of bulls of the same breed the horncores may be shorter in cows. In studying the effects of age it can be seen that growth rates are initially very high, slowing down at about three years of age. Tooth row length decreases with age and hence it is a particularly unreliable indicator of size. Obviously breed affects size and it has been possible to divide the skulls studied into size groups which correspond well with known body weights.     

Metaceratodus kaopen comb. nov. and M. wichmanni comb. nov., two Late Cretaceous South American species of an austral lungfish genus (Dipnoi)

Cione, A.L. & Gouiric-Cavalli, S., June 2012. Metaceratodus kaopen comb. nov. and M. wichmanni comb. nov., two Late Cretaceous South American species of an austral lungfish genus (Dipnoi). Alcheringa 36, 203–216. ISSN 0311-5518.

Metaceratodus wollastoni, an Australian species, was reported from Upper Cretaceous beds of Patagonia in 1997. Later, three new species (Ceratodus wichmanni, Ptychoceratodus kaopen and Ptychoceratodus cionei), based on scarce material, were described from the same region. Two of these species were later referred to Ferganoceratodus. After examining much more abundant and better-preserved material, we conclude that neither the occurrence of Metaceratodus wollastoni nor those of Ptychoceratodus and Ferganoceratodus in the Cretaceous of South America are supported. We consider that C. wichmanni and P. cionei are synonyms and we reassign the three putative species to Metaceratodus under two new combinations: M. kaopen comb. nov. and M. wichmanni comb. nov. Both differ from the other species of the genus in having pits over most of the occlusal surface and a different occlusal profile of the tooth plate, and most have four ridges in the lower and upper tooth plates. Metaceratodus wichmanni differs from M. kaopen in oclussal profile, inner angle, and symphysis development among other features. Metaceratodus kaopen is known from the upper Santonian–lower Campanian Anacleto Formation of Río Negro province and M. wichmanni from upper Campanian–lower Maastrichtian units of Chubut, Río Negro, Neuquén and Mendoza provinces, Argentina. The occurrence of Metaceratodus in southern South America corroborates a close biogeographical relationship with Australia in the latest Cretaceous.

Alberto Luis Cione [acione@museo.fcnym.unlp.edu.ar] and Soledad Gouiric-Cavalli [sgouiric@museo.fcnym.unlp.edu.ar], División Paleontología Vertebrados, Museo de La Plata, Paseo del Bosque s/n, W1900FWA La Plata, Argentina. Received 23.11.2010, revised 11.7.2011, accepted 7.8.2011.

     

Umkomasia (megasporophyll): part 1 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed

The genus Umkomasia, a megasporophyll, belonging to the pteridosperms (seed ferns) in the family Umkomasiaceae (Corystospermaceae), is reassessed comprehensively worldwide. All previous records are analysed. Certain fertile structures previously attributed are reclassified. Umkomasia is shown to be restricted to the Triassic of Gondwana where it is associated with the genus Pteruchus, a microsporophyll, and the genus Dicroidium, a vegetative leaf. It is well represented from Argentina, Australia and southern Africa where the Molteno Formation is by far the most comprehensively sampled with eight species described. Two specimens from the upper Permian of India attributed to Umkomasia are reclassified as cf. Arberiopsis sp. A whorled fertile structure from Antarctica, previously assigned to Umkomasia, is reclassified in a new genus as Axsmithia uniramia. Another compression fossil and the permineralized Umkomasia resinosa remain as valid records from Antarctica. The material described as Umkomasia from the Triassic of China is reclassified as Stenorachis asiatica. The Lower Jurassic record from Germany is placed in a new genus as Kirchmuellia franconica. The records of Umkomasia sp. from the Rhaetic of Germany are reclassified as cf. Kirchmuellia sp. and the single specimen from the Jurassic of Libya as genus et sp. indet. The Lower Cretaceous record from Mongolia has been reclassified by other researchers as Doylea mongolica. A pictorial key to Umkomasia species is provided, geographic and stratigraphic distributions are tabulated.