A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods

Kear, B.P., Fordyce, R.E., Hiller, N. & Siversson, M., December 2017. A palaeobiogeographical synthesis of Australasian Mesozoic marine tetrapods. Alcheringa 42, 461-486. ISSN 0311-5518.

THE LAST 15 years has witnessed a blossoming of research on Australasian Mesozoic marine tetrapod fossils. Much of this work has focused on amniotes, particularly those from the prolific Lower Cretaceous (Aptian–Albian) Lagerstätten of the Eromanga Basin in central and eastern Australia, and Upper Cretaceous (Campanian–Maastrichtian) sequences of the North and South islands of New Zealand. However, rare and less popularized remains have also been found in Lower Triassic–mid-Cretaceous rocks from Australia, New Zealand and the Chatham Islands, and on the tectonically proximal landmasses of New Caledonia and Timor. Currently identified taxa include estuarine–paralic rhytidostean, brachyopid, capitosaurian and trematosaurian temnospondyls from the earliest Triassic (Induan–Olenekian), Middle–Late Triassic (Anisian–Norian) eosauropterygians, and mixosaurian, shastasaurian and euichthyosaurian ichthyosaurians, Early–Middle Jurassic (Sinemurian–Bajocian) ichthyosaurians, together with plesiosauroid and rhomaleosaurid-like plesiosaurians, and diverse Early (Aptian–Albian) through to Late Cretaceous (Campanian–Maastrichtian) elasmosaurid, leptocleidid, polycotylid, probable cryptoclidid and pliosaurid plesiosaurians, as well as ophthalmosaurid ichthyosaurians, sea turtles incorporating protostegids, and mosasaurid squamates. This faunal succession evidences almost continuous occupation of southern high-palaeolatitude seas, and repeated endemic diversifications (including nascent members of some key lineages) amongst emigrant cosmopolitan clades. The primary dispersal routes are likely to have been peri-Gondwanan, with coastal migrations along the western Tethys and polar margins of the Panthalassan Ocean. However, augmentation by increasing continental fragmentation and seaway corridor connectivity probably occurred from the Middle Jurassic to Late Cretaceous. Latest Cretaceous mosasaurid and elasmosaurid taxa also reveal regional affinities with the emergent western Pacific and Weddellian austral bioprovinces. The extreme rarity, or complete absence, of many major groups prevalent elsewhere in Gondwana (e.g., tanystropheids, Triassic sauropterygians, bothremydid marine turtles, thalattosuchians and dyrosaurid crocodylomorphs) is conspicuous, and might be related to stratigraphical/collecting biases, or the predominantly higher-palaeolatitude, cooler-water Mesozoic palaeogeography of the Australasian region. Although the burgeoning record is substantial, much still awaits discovery and adequate documentation; thus Australasia is still one of the most exciting prospects for future insights into the global history of Mesozoic marine tetrapods.

Benjamin P. Kear* [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, SE-752 36 Uppsala, Sweden; R. Ewan Fordyce [ewan.fordyce@otago.ac.nz] Department of Geology, University of Otago, Post Box 56, Dunedin 9054, New Zealand; Norton Hiller [norton.hiller@gmail.com] Canterbury Museum, Rolleston Avenue, Christchurch 8013, New Zealand; Mikael Siversson [mikael.siversson@museum.wa.gov.au] Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106.     

First record of Carinodens (Squamata,Mosasauridae) from the uppermost Maastrichtian of Stevns Klint,Denmark

Milàn, J., Jagt, J.W.M., Lindgren, J. & Schulp, A.S., November 2017. First record of Carinodens (Squamata, Mosasauridae) from the uppermost Maastrichtian of Stevns Klint, Denmark. Alcheringa 42, 597-602. ISSN 0311-5518.

Here we report on an important addition to the Late Cretaceous fossil record of marine reptiles from Denmark: a tooth crown of the rare durophagous mosasaur Carinodens minalmamar found in the uppermost Maastrichtian strata at the UNESCO World Heritage Site Stevns Klint. The tooth was found within the uppermost few metres of the Maastrichtian chalk placing it within the latest 50.000 years prior to the K/Pg boundary. The new find is a shed crown probably representing a tooth from the 11th to 13th position in the jaw. The tooth represents the northernmost occurrence of the genus Carinodens. Previous mosasaur finds from Denmark have all been from the hypercarnivorous mosasaurids Mosasaurus hoffmannii and Plioplatecarpus sp., thus our specimen adds a new trophic niche exploited by marine tetrapods in the food web of the latest Maastrichtian of Denmark.

Jesper Milàn* [jesperm@oesm.dk], Geomuseum Faxe/Østsjællands Museum, Østervej 2, 4640 Faxe, Denmark; John W.M. Jagt [john.jagt@maastricht.nl], Natuurhistorisch Museum Maastricht, De Bosquetplein 6–7, 6211 kJ Maastricht, the Netherlands; Johan Lindgren [johan.lindgren@geol.lu.se], Department of Geology, Lund University, Sölvegatan 12, 223 62 Lund, Sweden. Anne S. Schulp? [anne.schulp@naturalis.nl], Natuurhistorisch Museum Maastricht, De Bosquetplein 6–7, 6211 kJ Maastricht, the Netherlands. *Also affiliated with: Natural History Museum of Denmark, Øster Voldgade 5–7, 1465 Copenhagen K, Denmark. ?Also affiliated with: Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands, and Faculteit Bètawetenschappen, Vrije Universiteit Amsterdam, De Boelelaan 1085, 1081 HV Amsterdam, the Netherlands.     

Two new skeletons of the enigmatic,rail-like avian taxon Songzia Hou, 1990 (Songziidae) from the early Eocene of China

Sachs, S. &; Kear, B.P. November 2018. A rare new Pliensbachian plesiosaurian from the Amaltheenton Formation of Bielefeld in northwestern Germany. Alcheringa 42, 487-500. ISSN 0311-5518.

We describe a new plesiosaurian from the upper Pliensbachian Amaltheenton Formation of Bielefeld in northwestern Germany. The taxon is based upon an incomplete associated skeleton comprising part of the right mandibular ramus, several teeth, a series of cervical, pectoral, dorsal and caudal vertebrae, as well as ribs, limb girdle elements including a nearly complete right scapula, and various distal limb bones. A unique character state combination serves to distinguish the Amaltheenton Formation remains from other previously documented Early Jurassic plesiosaurians. The most important features are the presence of a longitudinal notch incising the posterior rim of the glenoid fossa and retroarticular process, and a pronounced ventrolateral shelf on the scapula, both of which constitute derived states otherwise shared with Early Cretaceous leptocleidians. However, phylogenetic analysis using a ‘total group’ Plesiosauria data-set that specifically accommodates for Pliensbachian taxa unanimously placed the Amaltheenton Formation plesiosaurian among Early–Middle Jurassic pliosaurids. This discovery is significant because it reveals unexpected homoplasy, but also because it establishes what is only the third formally named plesiosaurian taxon thus far documented from Pliensbachian strata worldwide.

Sven Sachs^* [sachs.pal@gmail.com], Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany and Im Hof 9, 51766 Engelskirchen, Germany; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden.     

Early Cretaceous polar biotas of Victoria,southeastern Australia—an overview of research to date

Poropat, S.F., Martin, S.K., Tosolini, A.-M.P., Wagstaff, B.E, Bean, L.B., Kear, B.P., Vickers-Rich, P. &; Rich, T.H., May 2018. Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date. Alcheringa 42, 158–230. ISSN 0311-5518.

Although Cretaceous fossils (coal excluded) from Victoria, Australia, were first reported in the 1850s, it was not until the 1950s that detailed studies of these fossils were undertaken. Numerous fossil localities have been identified in Victoria since the 1960s, including the Koonwarra Fossil Bed (Strzelecki Group) near Leongatha, the Dinosaur Cove and Eric the Red West sites (Otway Group) at Cape Otway, and the Flat Rocks site (Strzelecki Group) near Cape Paterson. Systematic exploration over the past five decades has resulted in the collection of thousands of fossils representing various plants, invertebrates and vertebrates. Some of the best-preserved and most diverse Hauterivian–Barremian floral assemblages in Australia derive from outcrops of the lower Strzelecki Group in the Gippsland Basin. The slightly younger Koonwarra Fossil Bed (Aptian) is a Konservat-Lagerstätte that also preserves abundant plants, including one of the oldest known flowers. In addition, insects, crustaceans (including the only syncaridans known from Australia between the Triassic and the present), arachnids (including Australia’s only known opilione), the stratigraphically youngest xiphosurans from Australia, bryozoans, unionoid molluscs and a rich assemblage of actinopterygian fish are known from the Koonwarra Fossil Bed. The oldest known—and only Mesozoic—fossil feathers from the Australian continent constitute the only evidence for tetrapods at Koonwarra. By contrast, the Barremian–Aptian-aged deposits at the Flat Rocks site, and the Aptian–Albian-aged strata at the Dinosaur Cove and Eric the Red West sites, are all dominated by tetrapod fossils, with actinopterygians and dipnoans relatively rare. Small ornithopod (=basal neornithischian) dinosaurs are numerically common, known from four partial skeletons and a multitude of isolated bones. Aquatic meiolaniform turtles constitute another prominent faunal element, represented by numerous isolated bones and articulated carapaces and plastrons. More than 50 specimens—mostly lower jaws—evince a high diversity of mammals, including monotremes, a multituberculate and several enigmatic ausktribosphenids. Relatively minor components of these fossil assemblages are diverse theropods (including birds), rare ankylosaurs and ceratopsians, pterosaurs, non-marine plesiosaurs and a lepidosaur. In the older strata of the upper Strzelecki Group, temnospondyl amphibians—the youngest known worldwide—are a conspicuous component of the fauna, whereas crocodylomorphs appear to be present only in up-sequence deposits of the Otway Group. Invertebrates are uncommon, although decapod crustaceans and unionoid bivalves have been described. Collectively, the Early Cretaceous biota of Victoria provides insights into a unique Mesozoic high-latitude palaeoenvironment and elucidates both palaeoclimatic and palaeobiogeographic changes throughout more than 25 million years of geological time.

Stephen F. Poropat*? [sporopat@swin.edu.au; stephenfporopat@gmail.com], Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Sarah K. Martin*? [sarah.martin@dmirs.wa.gov.au; martin.sarahk@gmail.com] Geological Survey of Western Australia, 100 Plain St, East Perth, Western Australia 6004, Australia; Anne-Marie P. Tosolini [a.tosolini@unimelb.edu.au] and Barbara E. Wagstaff [wagstaff@unimelb.edu.au] School of Earth Sciences, The University of Melbourne, Melbourne, Victoria 3010, Australia; Lynne B. Bean [lynne.bean@anu.edu.au] Research School of Earth Sciences, Australian National University, Acton, Canberra, Australian Capital Territory 2001, Australia; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden; Patricia Vickers-Rich§ [prich@swin.edu.au; pat.rich@monash.edu] Faculty of Science, Engineering and Technology, Swinburne University of Technology, John St, Hawthorn, Victoria 3122, Australia; Thomas H. Rich [trich@museum.vic.gov.au] Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia. *These authors contributed equally to this work. ?Also affiliated with: Australian Age of Dinosaurs Museum of Natural History, Lot 1 Dinosaur Drive, PO Box 408, Winton, Queensland 4735, Australia. ?Also affiliated with: Earth and Planetary Sciences, Western Australian Museum, Welshpool, Western Australia 6101, Australia. §Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne, Victoria 3800, Australia.     

A new Miocene carnivorous marsupial,Barinya kutjamarpensis (Dasyuromorphia), from central Australia

Binfield, P., Archer, M., Hand, S.J., Black, K.H., Myers, T.J., Gillespie, A.K. & Arena, D.A., June 2016. A new Miocene carnivorous marsupial, Barinya kutjamarpensis (Dasyuromorphia), from central Australia. Alcheringa 41, xx–xx. ISSN 0311-5518.

A new dasyuromorphian, Barinya kutjamarpensis sp. nov., is described on the basis of a partial dentary recovered from the Miocene Wipajiri Formation of northern South Australia. Although about the same size as the only other species of this genus, B. wangala from the Miocene faunal assemblages of the Riversleigh World Heritage Area, northwestern Queensland, it has significant differences in morphology including a very reduced talonid on M₄ and proportionately wider molars. Based on the structural differences and the more extensive wear on its teeth, the central Australian species might have consumed harder or more abrasive prey in a more silt-rich environment than its congener, which hunted in the wet early to middle Miocene forests of Riversleigh.

Pippa Binfield [pippa.binfield@outlook.com], Michael Archer [m.archer@unsw.edu.au], Suzanne J. Hand [s.hand@unsw.edu.au], Karen H. Black [k.black@unsw.edu.au], Troy J. Myers [t.myers@unsw.edu.au] Anna K. Gillespie [a.gillespie@unsw.edu.au] and Derrick A. Arena [rick@sitc.net.au], PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales 2052, Sydney, Australia.     

The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri

Rich, T.H., Hopson, J.A., Gill, P.G., Trusler, P., Rogers-Davidson, S., Morton, S., Cifelli, R.L., Pickering, D., Kool, L., Siu, K., Burgmann, F.A., Senden, T., Evans, A.R., Wagstaff, B.E., Seegets-Villiers, D., Corfe, I.J., Flannery, T.F., Walker, K., Musser, A.M., Archer, M., Pian, R. & Vickers-Rich, P., June 2016. The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri. Alcheringa 40, xx–xx. ISSN 0311-5518.

The monotreme Teinolophos trusleri Rich, Vickers-Rich, Constantine, Flannery, Kool & van Klaveren, 1999 Rich, T.H., Vickers-Rich, P., Constantine, A., Flannery, T.F., Kool, L. & van Klaveren, N., 1999. Early Cretaceous mammals from Flat Rocks, Victoria, Australia. Records of the Queen Victoria Museum and Art Gallery 106, 1–34. [Google Scholar] from the Early Cretaceous of Australia is redescribed and reinterpreted here in light of additional specimens of that species and compared with the exquisitely preserved Early Cretaceous mammals from Liaoning Province, China. Together, this material indicates that although T. trusleri lacked a rod of postdentary bones contacting the dentary, as occurs in non-mammalian cynodonts and basal mammaliaforms, it did not share the condition present in all living mammals, including monotremes, of having the three auditory ossicles, which directly connect the tympanic membrane to the fenestra ovalis, being freely suspended within the middle ear cavity. Rather, T. trusleri appears to have had an intermediate condition, present in some Early Cretaceous mammals from Liaoning, in which the postdentary bones cum ear ossicles retained a connection to a persisting Meckel’s cartilage although not to the dentary. Teinolophos thus indicates that the condition of freely suspended auditory ossicles was acquired independently in monotremes and therian mammals. Much of the anterior region of the lower jaw of Teinolophos is now known, along with an isolated upper ultimate premolar. The previously unknown anterior region of the jaw is elongated and delicate as in extant monotremes, but differs in having at least seven antemolar teeth, which are separated by distinct diastemata. The dental formula of the lower jaw of Teinolophos trusleri as now known is i2 c1 p4 m5. Both the deep lower jaw and the long-rooted upper premolar indicate that Teinolophos, unlike undoubted ornithorhynchids (including the extinct Obdurodon), lacked a bill.

Thomas H. Rich [trich@museum.vic.gov.au], Sally Rogers-Davidson [srogers@museum.vic.gov.au], David Pickering [dpick@museum.vic.gov.au], Timothy F. Flannery [tim.flannery@textpublishing.com.au], Ken Walker [kwalker@museum.vic.gov.au], Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; James A. Hopson [jhopson@uchicago.edu], Department of Organismal Biology & Anatomy, University of Chicago,1025 East 57th Street, Chicago, IL 60637, USA; Pamela G. Gill [pam.gill@bristol.ac.uk], School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, U.K. and Earth Science Department, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Peter Trusler [peter@petertrusler.com.au], Lesley Kool [koollesley@gmail.com], Doris Seegets-Villiers [doris.seegets-villiers@monash.edu], Patricia Vickers-Rich [pat.rich@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Victoria 3800, Australia; Steve Morton [steve.morton@monash.edu], Karen Siu [karen.siu@monash.edu], School of Physics and Astronomy, Monash University, Victoria 3800, Australia; Richard L. Cifelli [rlc@ou.edu] Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, Norman, OK 73072, USA; Flame A. Burgmann [flame.burgmann@monash.edu], Monash Centre for Electron Microscopy, 10 Innovation Walk, Monash University, Clayton, Victoria 3800, Australia; Tim Senden [Tim.Senden@anu.edu.au], Department of Applied Mathematics, Research School of Physical Sciences and Engineering, The Australian National University, Canberra, Australian Capital Territory 0200, Australia; Alistair R. Evans [alistair.evans@monash.edu], School of Biological Sciences, Monash University, Victoria 3800, Australia; Barbara E. Wagstaff [wagstaff@unimelb.edu.au], School of Earth Sciences, The University of Melbourne, Victoria 3010, Australia; Ian J. Corfe [ian.corfe@helsinki.fi], Institute of Biotechnology, Viikinkaari 9, 00014, University of Helsinki, Finland; Anne M. Musser [anne.musser@austmus.gov.au], Australian Museum, 1 College Street, Sydney NSW 2010 Australia; Michael Archer [m.archer@unsw.edu.au], School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia; Rebecca Pian [rpian@amnh.org], Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA. Received 7.4.2016; accepted 14.4.2016.     

A preliminary report on new Ediacaran fossils from Iran

Vickers-Rich, P., Soleimani, S., Farjandi, F., Zand, M., Linnemann, U., Hofmann, M., Wilson, S.A., Cas, R. &; Rich, T.H. November, 2017. A preliminary report on new Ediacaran fossils from Iran. Alcheringa 42, 231–244. ISSN 0311-5518.

Recent exploratory field mapping of marine sedimentary sequences in the Koushk Mine locality of the Bafq region in Central Iran, and on the northern slopes of the Elborz Mountains south of the Caspian Sea, has yielded large complex body and trace fossils of Neoproterozoic–early Cambrian age. The recovered specimens resemble the previously documented Precambrian discoidal form Persimedusites, and a the tubular morphotype Corumbella, which is a novel occurrence for Iran and otherwise only recorded before from Brazil and the western USA. Additional enigmatic traces can not yet be interpreted unequivocally, but suggest that future work may uncover more unusual Ediacaran fossils from various localities in Central Iran.

Patricia Vickers-Rich* [prich@swin.edu.au, pat.rich@monash.edu], Faculty of Science, Swinburne University of Technology, Melbourne (Hawthorn), Victoria 3122, Australia; Sara Soleimani [sara_soleimani@yahoo.com], Palaeontology Department, Geological Survey of Iran, Tehran, Iran; Farnoosh Farjandi [farnooshfarjandi@gmail.com], Department of Geochemical Exploration, Geological Survey of Iran, Tehran, Iran; Mehdi Zand [zand.mehdi.geo@gmail.com], Geology Department, Bafq Mining Company, Koushk Mine, Yazd, Iran. Ulf Linnemann [ulf.linnemann@senckenberg.de], and Mandy Hofmann [mandy.hofmann@senckenberg.de], Senckenberg Naturhistorische Sammlungen, Dresden, Museum für Mineralogie und Geologie, Sektion Geochronologie, Koenigsbruecker Landstrasse 159, D-01109, Dresden, Germany; Siobhan A. Wilson [sasha.wilson@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Raymond Cas [ray.cas@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Thomas H. Rich? [trich@museum.vic.gov.au], Museum Victoria, Exhibition Gardens, PO Box 666, Melbourne, Victoria, 3001 Australia. *Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; School of Environmental Sciences, Deakin University, Melbourne (Burwood), Victoria, Australia 3125; Palaeontology Department, Museum Victoria, Carlton Gardens, PO Box 666, Melbourne, Victoria 3001, Australia. ?Also affiliated with: School of Earth, Atmosphere and Environment, Monash University, Melbourne (Clayton), Victoria 3800, Australia; Faculty of Science, Swinburne University of Science and Technology, Melbourne (Hawthorn), Victoria 3122, Australia.     

Editorial

Kear, Benjamin P., 11 November 2019. Editorial. Alcheringa 43, XX–XX. ISSN 0311-5518.

Benjamin P. Kear [benjamin.kear@em.uu.se], Museum of Evolution, Uppsala University, 752 36 Uppsala, Sweden.     

A new species of Aboilus (Insecta,Orthoptera) from the Jurassic Daohugou beds of China,and discussion of forewing coloration in Aboilus

Wang, H., Li, S., Zhang, Q., Fang, Y., Wang, B. & Zhang, H., 13.02.2015. A new species of Aboilus (Insecta, Orthoptera) from the Jurassic Daohugou beds of China, and discussion of forewing coloration in Aboilus. Alcheringa 39, xxx–xxx. ISSN 0311-5518

He Wang* [wanghe0701@163.com], Sha Li* [lisha.paleo@gmail.com], Qi Zhang* [zhqi1105@126.com], Yan Fang [yanfang@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.*Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China. ?Also affiliated with Steinmann Institute, University of Bonn, Bonn 53115, Germany.

A new species of Aboilinae (Orthoptera: Prophalangopsidae), Aboilus perbellus, is described and illustrated based on three well-preserved forewings recovered from the Middle–Upper Jurassic Daohugou beds of Inner Mongolia, China. The new species differs from all congeneric forms in its special forewing coloration and features of its wing venation. To date, three types of forewing coloration have been found among different species of Aboilus at Daohugou, suggesting that these taxa inhabited different ecotopes.     

A new saucrosmylid lacewing (Insecta,Neuroptera) from the Middle Jurassic of Daohugou,Inner Mongolia,China

Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A., 2014. A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa 38. ISSN 0311-5518.

A new genus and new species of Saucrosmylidae (Insecta, Neuroptera) are described (Daohugosmylus castus) based on a well-preserved hindwing from the Middle Jurassic of Daohugou, Inner Mongolia, China. Daohugosmylus gen. nov. is distinguished by a large and nearly semi-circular hindwing, relatively wide R1 space possessing several rows of cells, anteriorly bent Rs, dense crossveins over the entire wing, and smooth outer margin.

Qing Liu (corresponding author) [qingliu1029@gmail.com], Haichun Zhang [hczhang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], Yan Fang [yanfang@nigpas.ac.cn], Daran Zheng [dranzheng@gmail.com], Qi Zhang [zhqi1105@126.com] and Edmund A Jarzembowski [jarzembowski2@live.co.uk], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China; secondary address of Daran Zheng & Qi Zhang, University of Chinese Academy of Sciences, Beijing, 100049, PR China; and Ed Jarzembowski, Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. Received 13.11.2013; revised 20.1.2014; accepted 21.1.2014.     

Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae,Caimaninae) in the Yecua Formation (late Miocene) of Bolivia

Tineo, D.E., Bona, P., Pérez, L.M., Vergani, G.D., González, G., Poiré, D.G., Gasparini, Z.N. & Legarreta, P., 1.10.2014. Palaeoenvironmental implications of the giant crocodylian Mourasuchus (Alligatoridae, Caimaninae) in the Yecua Formation (late Miocene) of Bolivia. Alcheringa 39, xxx–xxx. ISSN 0311-5518

Outcrops of the Yecua Formation (late Miocene) are exposed for approximately 230 m along the La Angostura section of the Piraí River (50 km southwest of Santa Cruz de la Sierra). These reveal massive (argillic palaeosols) and laminated (quiet-water lacustrine and marsh settings) mudstones interbedded with thin sandstones containing microfossils, molluscs and vertebrate remains. Significantly, the succession hosts a giant crocodylian, Mourasuchus (Alligatoridae, Caimaninae), which is represented by both skull and postcranial fragments found in association with freshwater turtles and fishes. Mourasuchus was distributed widely from the middle Miocene of Colombia to upper Miocene of Venezuela, Brazil and Argentina, suggesting connections between major fluvial systems and an active mechanism for dispersal of South American freshwater vertebrates during the Miocene.

David Eric Tineo [tineo.d.e@gmail.com] and Daniel Gustavo Poiré [dgpoire@yahoo.com.ar], CONICET—Centro de Investigaciones Geológicas, Universidad Nacional de La Plata. Calle 1 (644), B1900FWA, La Plata, Argentina; Paula Bona [paulabona26@gmail.com] and Zulma Gasparini [zgaspari@fcnym.unlp.edu.ar], CONICET—División Paleontología Vertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Leandro Martín Pérez [pilosaperez@gmail.com] CONICET—División Paleozoología Invertebrados, Museo de La Plata. Paseo del Bosque s/n, B1900FWA, La Plata, Argentina; Gustavo Dardo Vergani [gvergani@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina; Gloria González Rigas [ggonzalez@pluspetrol.net]—Pluspetrol Bolivia Corporation SA, Av. Grigotá esq. Las Palmas, Santa Cruz de la Sierra, Bolivia; Pablo Legarreta [plegarreta@pluspetrol.net]—Pluspetrol S.A. Lima (339), C1073AAG, Ciudad Autónoma de Buenos Aires, Argentina.     

Ammonite dating of latest Cretaceous mosasaurid reptiles (Squamata,Mosasauroidea) from Jordan—preliminary observations

Jagt, J.W.M., Jagt-Yazykova, E.A., Kaddumi, H.F. & Lindgren, J., April 2017. Ammonite dating of latest Cretaceous mosasaurid reptiles (Squamata, Mosasauroidea) from Jordan—preliminary observations. Alcheringa 42, 587-596. ISSN 0311-5518

Newly collected ammonoid material from the uppermost Cretaceous portion of the Muwaqqar Chalk Marl Formation exposed some 30 km southeast of the Qasr Al’Harrana area (east-central Jordan) includes medium-sized baculitids (Baculites ovatus auctorum, non Say), the sphenodiscid Libycoceras acutodorsatus (Noetling) and the pachydiscids Menuites fresvillensis (Seunes) and Pachydiscus (Pachydiscus) dossantosi (Maury). Of the two last named taxa, the former is a good marker species for the upper Maastrichtian, with records from Europe, central Chile, South India, Baluchistan (Pakistan), Australia, Madagascar and South Africa. The latter is known from the United Arab Emirates/Oman border area, from strata of (late) early to early late Maastrichtian age, as well as from more poorly constrained Maastrichtian levels in Brazil and Nigeria. A comparison with ammonoid assemblages from the Maastrichtian type area (southeast Netherlands/northeast Belgium) suggests correlation of the Muwaqqar Chalk Marl Formation with the middle/upper Maastricht Formation (Emael and Nekum members, ca 66.5–66.1 Ma) and the upper part of the coeval Kunrade Formation. However, associated ‘tegulated’ inoceramids of the Tenuipteria argentea group from the Muwaqqar Chalk Marl Formation favour equivalence with a higher level of the Maastrichtian type area, i.e., the Meerssen Member. From the upper Maastricht Formation and the equivalent upper part of the Kunrade Formation, the following mosasaur genera are currently known: Mosasaurus, Prognathodon, Plioplatecarpus and Carinodens. Interestingly, coeval strata of the Muwaqqar Chalk Marl Formation in east-central Jordan have yielded remains of a largely comparable suite comprising Prognathodon, Mosasaurus, Carinodens and an unnamed, highly derived plioplatecarpine.

John W.M. Jagt* [john.jagt@maastricht.nl], Natuurhistorisch Museum Maastricht, de Bosquetplein 6–7, 6211 kJ Maastricht, the Netherlands; Elena A. Jagt-Yazykova [eyazykova@uni.opole.pl], Opole University, Department of Biosystematics, Oleska 22, 45-052 Opole, Poland; Hani F. Kaddumi [blossom_trail@yahoo.com], Eternal River Museum of Natural History, Maroof Al’Rusafi Street, PO Box 11395, Amman 11123, Jordan; Johan Lindgren [johan.lindgren@geol.lu.se], Lund University, Department of Geology, Sölvegatan 12, 223 62 Lund, Sweden.     

First record of Homoctenus (Tentaculitoidea,Homoctenida) from the Late Devonian of northwest Peninsular Malaysia

Meor, H.A.H., Yong, A.M., Zakaria, M.Z.Z. & Ghani, A.A., 2.6.2015. First record of Homoctenus (Tentaculitoidea, Homoctenida) from the Late Devonian of northwest Peninsular Malaysia. Alcheringa 39, 550–558. ISSN 0311-5518.

The homoctenid tentaculitoid genus Homoctenus is reported for the first time from Peninsular Malaysia. The fossils derive from the Upper Devonian Sanai Limestone, exposed in the state of Perlis, northwest Peninsular Malaysia. The fossils are closely related to Homoctenus tenuicinctus tenuicinctus and are described as Homoctenus sp. cf. H. tenuicinctus. The homoctenids were recovered from an interval containing a rich conodont assemblage, indicating a Frasnian age (Palmatolepis linguiformis Zone).

Meor Hakif Amir Hassan [meorhakif@um.edu.my], Yong Adilah Mustafa [yongadilah@gmail.com], Mohamad Z.Z. Zakaria [zulhafiszariq@siswa.um.edu.my], Azman A. Ghani [azmangeo@um.edu.my], Geology Department, University of Malaya, 50603 Kuala Lumpur, Malaysia. Received 12.4.2015; revised 27.5.2015; accepted 2.6.2015.     

Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan,southern Guangxi,South China

Yang, T.L., He, W.H., Zhang, K.X., Wu, S.B., Zhang, Y., Yue, M.L., Wu, H.T. & Xiao, Y.F., November 2015. Palaeoecological insights into the Changhsingian–Induan (latest Permian–earliest Triassic) bivalve fauna at Dongpan, southern Guangxi, South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

The Talung Formation (latest Permian) and basal part of Luolou Formation (earliest Triassic) of the Dongpan section have yielded 30 bivalve species in 17 genera. Eight genera incorporating 11 species are systematically described herein, including three new species: Nuculopsis guangxiensis, Parallelodon changhsingensis and Palaeolima fangi. Two assemblages are recognized, i.e., the Hunanopecten exilis–Euchondria fusuiensis assemblage from the Talung Formation and the Claraia dieneri–Claraia griesbachi assemblage from the Luolou Formation. The former is characterized by abundant Euchondria fusuiensis, an endemic species, associated with other common genera, such as Hunanopecten, which make it unique from coeval assemblages of South China. A palaeoecological analysis indicates that the Changhsingian bivalve assemblage at Dongpan is diverse and represented by various life habits characteristic of a complex ecosystem. This also suggests that redox conditions were oxic to suboxic in deep marine environments of the southernmost Yangtze Basin during the late Changhsingian, although several episodes of anoxic perturbations and declines in palaeoproductivity saw deterioratation of local habitats and altered the taxonomic composition or population size of the bivalve fauna.

Tinglu Yang [yang@geology.so], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Weihong He* [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Shunbao Wu [shbwu@cug.edu.cn], Yang Zhang [zhangy05@163.com], Mingliang Yue [812182779@qq.com], Huiting Wu [ht_wu415@163.com] and Yifan Xiao [shadowyi@sohu.com], School of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

The age of the Takatika Grit,Chatham Islands,New Zealand

Hollis, C.J, Stickley, C.E., Bijl, P.K., Schiøler, P., Clowes, C.D., Li, X, Campbell, H. March 2017. The age of the Takatika Grit, Chatham Islands, New Zealand. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

The oldest Paleogene strata on Chatham Islands, east of New Zealand, are the phosphatized conglomerates and sandstones of the Takatika Grit that crops out on the northeastern coast at Tioriori and unconformably overlies the Chatham Schist. An intact Cretaceous–Paleogene boundary transition is not preserved at this locality. New biostratigraphic analysis of dinoflagellate, diatom and radiolarian microfossil assemblages confirms that the Takatika Grit is of late early–middle Paleocene (New Zealand Teurian stage) age but contains reworked microfossils of early Campanian (Early Haumurian) age. Vertebrate fossils found in this unit are inferred to be a mixture of reworked Cretaceous and in situ Paleocene bones and teeth. The overlying Tutuiri Greensand is of middle–late Paleocene age in its lower part and also contains reworked Cretaceous microfossils.

Christopher J. Hollis [c.hollis@gns.cri.nz], Chris Clowes [c.clowes@gns.cri.nz], Xun Li [x.li@gns.cri.nz], Hamish Campbell [h.campbell@gns.cri.nz], GNS Science, PO Box 30-368, Lower Hutt 5040, New Zealand; Catherine Stickley, Evolution Applied Limited, 50 Mitchell Way, Upper Rissington, Cheltenham GL54 2PL, UK [catherine.stickley@gmail.com]; Peter Bijl [p.k.bijl@uu.nl], Marine Palynology and Paleoceanography, Department of Earth Sciences, Utrecht University, 3584 CS Utrecht, the Netherlands; Poul Schiøler [poul.schioler@mgpalaeo.com.au], Morgan Goodall Palaeo, Unit 1/5 Arvida St, Malaga, WA 6090, Australia.     

The first palaeontinid from the Late Jurassic of Australia (Hemiptera,Cicadomorpha, Palaeontinidae)

Chen, J., Beattie, R., Wang, B., Jiang, H., Zheng, Y. & Zhang, H., 12 April 2019. The first palaeontinid from the Late Jurassic of Australia (Hemiptera, Cicadomorpha, Palaeontinidae). Alcheringa 43, 449–454. ISSN 0311-5518.

Palaeontinidae, an extinct group of large arboreal insects, has the most diverse record among the Mesozoic Hemiptera, but only a few taxa have been reported from the Southern Hemisphere. Herein, Talbragarocossus jurassicus Chen, Beattie & Wang gen. et sp. nov., one of the earliest representatives of ‘late’ Palaeontinidae, is described and illustrated from the Upper Jurassic Talbragar Fossil Fish Bed in New South Wales, Australia. This new taxon constitutes the first representative of Palaeontinidae in Australia and the first Jurassic example in Gondwanaland, providing significant distributional and stratigraphic extensions to the family.

Jun Chen*? [rubiscada@sina.com] and Yan Zheng? [zhengyan536@163.com], Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi 276000, China. Bo Wang? [bowang@nigpas.ac.cn], Hui Jiang [huijiang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. Robert Beattie [rgbeattie@bigpond.com], Australian Museum, 1 William St., Sydney, NSW 2010, Australia. ?Also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. ?Also affiliated with: Shandong Provincial Key Laboratory of Depositional Mineralization & Sedimentary Minerals, Shandong University of Science and Technology, Qingdao, Shandong 266590, China.     

A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida

Zhang, Y., He, W.H., Shi, G.R., Zhang, K.X. & Wu, H.T., 26.2.2015. A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida. Alcheringa 39, xxx–xxx. ISSN 0311-5518.

As the third and last part of a systematic palaeontological study of the brachiopod fauna from the Permian–Triassic boundary section at Zhongzhai in Guizhou Province (South China), this paper reports 15 species (including three new species: Tethyochonetes minor sp. nov., Neochonetes (Zhongyingia) transversa sp. nov., Paryphella acutula sp. nov.) in Order Productida. In addition, the morphological features and definitions of several key Changhsingian brachiopod taxa (e.g., Paryphella and Oldhamina interrupta) are clarified and revised.

Yang Zhang* [zhangy05@163.com] and G.R. Shi [guang.shi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; Weihong He [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Huiting Wu [ht_wu415@163.com], Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China. *Also affiliated with: Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

Gazella (Bovidae,Ruminantia) remains from the Siwalik Group of Pakistan

Khan, M.A., Babar, M.A., Akhtar, M., Iliopoulos, G., Rakha, A. & Noor, T., November 2015. Gazella (Bovidae, Ruminantia) remains from the Siwalik Group of Pakistan. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

New gazelle fossils are described from the Siwalik Group of Pakistan. The material includes horncores, maxilla and mandible fragments, and isolated teeth. The available samples are assigned to three Gazella species: Gazella sp. in the Lower Siwalik Subgroup (ca 14.2–11.2 Ma), and G. lydekkeri and G. superba in the Middle Siwalik Subgroup (ca 10.2–3.4 Ma). Based on a review of the Siwalik Group gazelles, G. padriensis is synonymized with G. lydekkeri. Gazella superba Pilgrim, 1939 sensu stricto is a large form and is a valid species of the genus in the Siwalik Group.

Muhammad Akbar Khan [akbaar111@yahoo.ca], Muhammad Adeeb Babar [babar_fcc2005@yahoo.com], Muhammad Akhtar [drakhtarfdrc@hotmail.com], Allah Rakha [babar.441@googlemail.com], Tuba Noor [akbar.zool@pu.edu.pk], Abu Bakr Fossil Display & Research Centre, Department of Zoology, Quid-e-Azam Campus, Punjab University (54590), Lahore, Pakistan; George Iliopoulos [borelakias@gmail.com], Geology Department of the University of Patras, Patras, Greece.     

New damselflies (Odonata: Zygoptera: Hemiphlebiidae,Dysagrionidae) from mid-Cretaceous Burmese amber

Zheng, D., Zhang, Q., Nel, A., Jarzembowski, E.A., Zhou, Z., Chang, S.-C. &; Wang, B., May 2016. New damselflies (Odonata: Zygoptera: Hemiphlebiidae, Dysagrionidae) from mid-Cretaceous Burmese amber. Alcheringa XX, xxx–xxx. ISSN 0311-5518

Two damselflies, Burmahemiphlebia zhangi gen. et sp. nov. and Palaeodysagrion cretacicus gen. et sp. nov., are described from the mid-Cretaceous Burmese amber. Burmahemiphlebia zhangi is the first record of Hemiphlebiidae from this amber, although the family was cosmopolitan during the Mesozoic. It can be readily distinguished from all other members of Hemiphlebiidae in having very short MP and CuA veins, and in its rectangular discoidal cell. The new fossils support the view that hemiphlebiid damselflies were one of the dominant groups of Zygoptera during the Mesozoic. Palaeodysagrion cretacicus is the first dysagrionid damselfly from Burmese amber and the second Mesozoic representative of this predominantly Paleogene group. It differs from other members of Dysagrionidae in having a unique elongate discoidal cell. These new finds increase the diversity of damselflies in mid-Cretaceous Burmese amber.

Daran Zheng* [dranzheng@gmail.com], Su-Chin Chang [suchin@hku.hk], Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Qingqing Zhang [qqzhang@nigpas.ac.cn], Edmund A. Jarzembowski? [edj@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Zhicheng Zhou [172692308@qq.com], The PLA Information Engineering University, 62 Kexue Ave, Gaoxin District, Zhengzhou 450002, Henan, PR China. *Also affiliated with State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China. ?Also affiliated with Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. ?Also affiliated with Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China.     

Late Ordovician cephalopods,tentaculitides, machaeridians and echinoderms from Kuh-e Faraghan,High Zagros,Iran

Ghavidel-Syooki, M., Evans, D.H., Ghobadi Pour, M., Popov, L.E., Álvaro, J.J., Rakhmonov, U., Klishevich, I.A. & Ehsani, M.D., 15.5.2015. Late Ordovician cephalopods, tentaculitides, machaeridians and echinoderm columnals from Kuh-e Faraghun, High Zagros, Iran. Alcheringa 39, 530–549. ISSN 0311-5518.

Late Ordovician (Katian, uppermost Acanthochitina barbata to Armoricochitina nigerica chitinozoan zones) cephalopods, tentaculitides, machaeridians and echinoderms are documented for the first time from the southern Zagros Ranges. A low-diversity cephalopod fauna includes Geisonocerina dargazense sp. nov., Isorthoceras sp. cf. I. bisignatum (Barrande) and other undetermined orthoceratides. The presence of Late Ordovician tentaculitides in the high- to mid-latitude margins of Gondwana has been documented previously, but no examples have been described in detail. Thus, Costatulites kimi sp. nov., which currently occurs associated with brachiopods characteristic of the Svobodaina havliceki (brachiopod) Association, represents the earliest undoubted record of tentaculitides in Gondwana. Machaeridians constitute a relatively common component of the Late Ordovician benthic faunas from the Mediterranean margin of Gondwana, but no previous records on the Gondwanan Iranian-Arabian segment have been reported. Three echinoderm taxa based on dissociated columnals are documented from the Armoricochitina nigerica chitinozoan Zone, including Sumsaricystis radiatus Stukalina, Ristnacrinus sp. and Rosulicrinus rosulus Stukalina.

Mohammad Ghavidel-Syooki [m_ghavidelsyooki@yahoo.com] Institute of Petroleum Engineering, Technical Faculty of Tehran University, PO Box 11365-4563, Tehran, Iran; David H. Evans [david.evans@naturalengland.org.uk], Natural England, Suite D, Unex House, Bourges Boulevard, Peterborough PE1 1NG, UK; Mansoureh Ghobadi Pour* [mghobadipour@yahoo.co.uk], Department of Geology, Faculty of Sciences, Golestan University, Gorgan, Iran; Leonid E. Popov [leonid.popov@museumwales.ac.uk], Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK; J. Javier Álvaro [jj.alvaro@csic.es], Instituto de Geociencias (CSIC-UCM), c/ José Antonio Novais 2, 28040 Madrid, Spain; Utkyr Rakhmonov, Kitab State Geological Reserve, 9 Ipak Yuli Street, Sakhrisabz, Uzbekistan; Inna A. Klishevich, Department of Historical Geology, Geological Faculty, St Petersburg State University, Universitetskaya Naberezhnaya 7/9, 199034 St Petersburg, Russia [inna@IK4848.spb.edu]; Mohammad H. Ehsani [mh_ehsani@hotmail.com], Institute of Petroleum Engineering, Technical Faculty of Tehran University, PO Box 11365-4563, Tehran, Iran. *Also affiliated with Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK.