Early Darriwilian graptolites from central and western China

Taxonomic restudy of Early Darriwilian graptolites from the Yangtze region, Chiadam and Alxa, has enabled a precise correlation of the Undulograptus austrodentatus Zone (including two subzones) across the three regions, and with the Darriwilian GSSP at Huangnitang in the Jiangshan-Changshan-Yushan (JYC) area. Based on specimens from 17 localities in the Yangtze region, the Exigraptus clavus and Undulograptus austrodentatus zones are recognised as replacements respectively of the Undulograptus sinodentatus (Lower) and U. austrodentatus (Upper) zones of Mu et al. (1979). The U. sinicus Subzone, the upper subzone of the U. austrodentatus Zone, is also recognised from the Sichuan-Guizhou area of the Yangtze region. Two subzones of the U. austrodentatus Zone have also been identified from the Chiadam region with more diverse graptolite fauna. The U. austrodentatus Zone fauna also occurs in the Alxa region. A general pattern of changing graptolite diversity from platform through basinal slope to deep basin is proposed, based on early Darriwilian bio- and lithofacies changes from the Yangtze Platform through the Jiangnan belt to the Zhujiang basin. The graptolite diversity increased with water depth and more off-shore setting. Key graptolite species are illustrated.     

Early Cretaceous (Neocomian) microplankton from the Carpentaria Basin,northern Queensland

The Neocomian upper Helby Beds, basal Rolling Downs Group and Gilbert River Formation of the Carpentaria Basin contain a dinoflagellate sequence divisible into three zonal intervals; the zones represent the oldest continuous marine Cretaceous record for eastern Australia. Zone DK1 (dated latest Jurassic to Berriasian) commences with the first appearance of four species, zone DK2 (dated Valanginian) with that of four or five species, and zone DK3 (dated Hauterivian to Barremian) with that of three species.

Biostratigraphic aspects are discussed of species selected as possible zone marker fossils in so far as they (a) are relatively poorly known, or represent new species, and (b) have no previously published Neocomian record. New combinations are proposed of one acritarch species and six dinoflagellate species. One new acritarch genus and species, Pseudofromea collaris, and four new dinoflagellate species — Canningia crassicingulata, Cleistosphaeridium australe, Cyclonephelium asymmetricum, and Muderongia testudinaria — are proposed and described.     

Fusulinoideans from the early Midian (late Middle Permian) Metadoliolina dutkevitchi-Monodiexodina sutchanica Zone of the Senkina Shapka section,South Primorye,Far East Russia

Fusulinoideans from the Metadoliolina dutkevitchi-Monodiexodina sutchanica Zone of the lower part of the Chandalaz Formation in the Senkina Shapka section in South Primorye, Far East Russia, are described. The fusulinoidean zone is assigned to the early Midian (=Capitanian: late Middle Permian) based mainly on the morphologie and biostratigraphic characteristics of Metadoliolina dutkevitchi. Previously, a Midian age has been established for the Metadoliolina dutkevitchi-Monodiexodina sutchanica Zone by the coexistence of Lepidolina species. However, the occurrence of Lepidolina with the two zonal species in this area has not been verified by the illustration of Lepidolina specimens. We examined a fusulinoidean-bearing sample from the Metadoliolina dutkevitchi-Monodiexodina sutchanica Zone, and three fusulinoidean species, Monodiexodina sutchanica, Pseudofusulina sp. and Metadoliolina dutkevitchi, are de-scribed and illustrated.     

Palaeontology of the Cerros Bayos section,Roca Formation (Danian), La Pampa Province,Argentina

Palynofloras from an outlier of Tertiary sediments in the One Tree Hill area north of Adelaide are nonmarine and correlate with similar assemblages from Middle Eocene North Maslin Sands from Maslin Bay and Golden Grove. Although there are similarities with time-equivalent Lower Nothofagidites asperus Zone palynofloras in the Gippsland Basin, a number of species in the South Australian palynofloras do not extend below the Late Eocene in the Gippsland sequences. This indicates earlier appearances for these species in southcentral Australia. The sediments overlie a highly weathered bedrock palaeosurface, indicating that a phase of significant weathering occurred prior to the Middle Eocene. A number of selected taxa are illustrated and their distribution in the palynofloras is discussed. Two new species, Proteacidites mildenhallii and Proteacidites parrawirrensis, are described and one species, Rhoipites byfieldensis, is emended.     

Parotosuchus from the Early Triassic of Queensland and Western Australia

Three members of the labyrinthodont Family Capitosauridae are described. Two new species from the Arcadia Formation of the Rewan Group, Queensland, show affinities with more primitive capitosaurs and with pre-capitosaurid groups. A third skull from the Blina Shale of Western Australia is of indeterminate species. These are the first recorded capitosaurs from Queensland and Western Australia and the first Parotosuchus from a Lystrosaurus Zone fauna.     

Early Silurian (Llandovery) trilobites from the Cotton Formation,near Forbes,New South Wales,Australia

A trilobite fauna from the upper part of the Cotton Formation (late Llandovery, Spirograptus turriculatus Zone) in central west New South Wales includes the new species Raphiophorus sandfordi, Odontopleura (Sinespinaspis) markhami and Aulacopleura pogsoni. Species of Odontopleura and Aulacopleura have not previously been documented in Australasia. The association of these genera is observed in offshore biofacies in the Llandovery/Wenlock on several palaeocontinents. Affinities of the species are with congeners from Bohemia and northwestern Canada (Odontopleura, Aulacopleura) and Tarim and South China (Aulacopleura, Raphiophorus).     

Trilobite faunas and palaeoenvironmental setting of the Silurian (early Ludlow) Melbourne Formation,central Victoria

Trilobites are common faunal elements in the Melbourne Formation, a unit of early Ludlow (upper nilssoni Biozone) age, which crops out extensively in the Darraweit Guim Province of the Melbourne Zone, central Victoria. New diagnoses are given for species previously described, including Maurotarion euryceps (McCoy, 1876; = Cyphaspis spryi Gregory, 1901), Raphiophorus jikaensis (Chapman, 1912; = Ampyx yarraensis Chapman, 1912), Cromus simpliciculus (Talent, 1964), Cromus spryi (Chapman, 1912), Sthenarocalymene kilmorensis (Gill, 1945; = Gravicalymene hetera Gill, 1945) and Trimerus harrisoni (McCoy, 1876). A new phacopid genus, Orygmatos is described, represented by the species O. yanyeani gen. et sp. nov. Other species newly described include Cromus melbournensis sp. nov., Arcticalymene australis sp. nov., “Ananaspis” woiwurrungi sp. nov. and Kettneraspis hollowayi sp. nov.

Species composition of the trilobite fauna varies spatially, and a number of distinct assemblages can be defined. Abundant trilobite moult configurations are conclusive for interpretation of the benthic fauna as autochthonous, inferring depth estimations based on benthic community distribution to be valid. A depth-related succession of communities is recognised and indicate the Melbourne Formation was deposited at relatively shallow depths on a broad, eastwardly deepening shelf, with deposition dominated by storm processes. The palaeoenvironment comprised a BA-1 community including the Arcticalymene australis trilobite assemblage, restricted to very shallow depths (～20 m) on the SW coastal margin of the shelf and preserved in proximal tempestite lithologies; and a BA-5 community group containing three distinct trilobite assemblages dominated by species of Cromus and a deeper water fauna, preserved in distal tempestite lithologies and ranging widely over the shelf at depths in the range of maximum storm wave base (～50 – 80 m).     

Conodont fauna and biostratigraphy of the Honghuayuan Formation (Early Ordovician) of Guizhou,South China

Serratognathus diversus An, Cornuodus longibasis (Lindström), Drepanodus arcuatus Pander, and eleven other less common conodonts, including Cornuodus? sp., Oistodus lanceolatus, Protopanderodus gradatus, Protoprioniodus simplicissimus, Juanognathus variabilis, Nasusgnathus dolonus, Paltodus? sp., Scolopodus houlianzhaiensis, Semiacontiodus apterus, Semiacontiodus sp. cf. S. cornuformis and Serratognathoides? sp., are described and illustrated from the Honghuayuan Formation in Guizhou, South China, concluding revision of the conodont fauna from this unit, which comprises 24 species in total. The most distinctive species in the fauna, S. diversus, consists of a trimembrate apparatus, including symmetrical Sa, asymmetrical Sb and strongly asymmetrical Sc elements. This species concept is supported by the absence of any other element types in a large collection represented by nearly 500 specimens of this species. The fauna indicates a late Tremadocian to mid-Floian age (Early Ordovician) for the Honghuayuan Formation, which was widely distributed on the Yangtze Platform in shallow water environments. Previously published biostratigraphic zonations for the Honghuayuan Formation are reviewed, and revised on the basis of our knowledge of the entire conodont fauna, supporting the establishment of three biozones, Triangulodus bifidus, Serratognathus diversus, and Prioniodus honghuayanensis biozones in ascending order. Species of Serratognathus enable correlation between Ordovician successions of South China, North China (North China Platform and Ordos Basin), Tarim Basin, and further afield into Malaysia and northwestern Australia.     

High latitude,late Campanian (Cretaceous) Radiolaria from offshore south-eastern Tasmania

A well-preserved, low diversity (ten species), high latitude (palaeolatitude 70°S) radiolarian fauna is recorded from shallow-water late Campanian (Late Cretaceious) sediments recovered in cores taken from the continental slope of southeastern Tasmania. These are the first Radiolaria described from Cretaceous rocks of southeastern Australia. Most forms are previously described species but a new species of Petasiforma is described. Age control is provided by dinoflagellates but the radiolarian fauna is similar to late Campanian-Maastrichtian faunas described from Campbell Plateau. A late Campanian age can also be determined from the Radiolaria, which are correlated with the Patulibracchium dickinsoni Zone. The site has subsided some 3200 m since deposition, at an average rate of about 40–45 m/Ma, consistent with other indicators in the region. Radiolaria are similar to other coeval plankton in exhibiting high latitude characteristics.     

Artinskian (Early Permian) fusuline fauna from the Rongma area in northern Tibet: palaeoclimatic and palaeobiogeographic implications

Zhang, Y.C., Wang, Y., Zhang, Y.J. & Yuan, D.X., 2013. Artinskian (Early Permian) fusuline fauna from the Rongma area in northern Tibet: palaeoclimatic and palaeobiogeographic implications. Alcheringa 37, 529–546. ISSN 0311–5518.

A fusuline fauna consisting of ten species in five genera from the Qudi Formation of the Jiaomuri and Gangtangcuo sections in the Rongma area, northern Tibet, is described. This fauna contains a few typical Peri-Gondwanan fusuline species such as Pseudofusulina pamirensis, Neodutkevitchia insignis and N. sp. cf. N. tumidiscula. The fauna is dated as Artinskian based on the presence of both genera Chalaroschwagerina and Praeskinnerella. The Peri-Gondwanan fusuline fauna in the Qiangtang Block is grouped palaeobiogeographically into the Southern Transitional Zone, which is located in the southern part of the Peri-Gondwanan blocks during the late Sakmarian and Artinskian. Moreover, the occurrence of the Peri-Gondwanan fusuline fauna, the lithological transition from glacimarine deposits to carbonates, and the Peri-Gondwanan fusulines’ apparent southeastward migration from the Kalmard Block of central Iran during the Artinskian, are interpreted here to be the result of global warming after the peak of the Late Palaeozoic Ice Age.

Yi-chun Zhang [geozyc@yahoo.com], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China; Yue Wang [yuewang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China; Yu-jie Zhang [zebiac@163.com], Chengdu Center, China Geological Survey, 2 Renming Road North, Chengdu 610081, PR China; Dong-xun Yuan [yuanzi55@163.com], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, China; Department of Earth Sciences, Nanjing University, 22 Hankou Road, Nanjing 210008, PR China. Received 12.3.2013; revised 2.5.2013; accepted 12.5.2013.     

Coleoptera from Late Medieval Smoke-Blackened Thatch (SBT): their Archaeological Implications

Abstract

Distinctive assemblages of Coleoptera (beetles) recovered from seven samples of smoke-blackened thatch (SBT)from a range of late medieval roofs in the south of England are listed. The age, ecology and possible origins of this fauna are discussed. Although its precise origin is known, it is suggested that its occurrence is not limited to thatch, and so cannot be used as a 'finger print' for thatch per se. A find of Sitophilus granarius, the 'granary weevil', suggests that thatch may represent another route by which this species has been incorporated into the archaeological record.     

Late Ordovician linguliformean microbrachiopods from north-central Kazakhstan

An assemblage of well-preserved Devonian trilobites is described from limestones in Satun Province, southern Thailand. The biogeographic implications are briefly discussed. The fauna, which is of Early Devonian age, probably Emsian, includes Decoroproetus, Cornuproetus, Platyscutellum and two species of Reedops, one of which is new. No similar fauna has been described before from the Shan-Thai block. The fauna is a typical Hercynian assemblage with close specific comparisons with Turkey, Morocco and Bohemia.     

Writing the region: Jonathan Couch and the Cornish fauna

Abstract

The county of Cornwall was the subject of an early nineteenth century attempt to compile an exhaustive regional fauna, Jonathan Couch's three volume A Cornish Fauna (1838–44). Although the Fauna was part of a much broader contemporary impulse to document the natural history of Britain, it was a very early example of a zoological study organised by geography rather than faunal group. It therefore required the author to make an explicit defence of its geographical approach and focus. According to Couch the region was the best place from which to build a better understanding of nature and, ultimately, of nature's creator. In particular, careful observations of a naturalist's local fauna could contribute to mid-nineteenth century debates over the distribution and migration of animals, their extinction, and their value to human industry. According to Couch, the county of Cornwall was especially suited to this sort of investigation. Indeed, although little had been done on the Cornish fauna before Couch's own study, his work was the catalyst for a number of other Cornish naturalists over the latter half of the century.     

New Archaeocyatha from the Early Cambrian of the Mt. Wright area,New South Wales

An archaeocyathan fauna from a single limestone lens in the upper Cymbric Vale Formation at Mt. Wright shows species in common with the historic Ajax Mine fauna of the Flinders Ranges, South Australia. The Cymbric Vale fauna is considered equivalent to the lower Sanashtykgol ‘horizon’ of the Altay Sayan fold belt of Siberia.

New taxa described are the genera Hyptocyathus (Inessocyathacea, Hyptocyathidae fam. nov.), Aporosocyathus (Annulocyathidae), Wrighticyathus (Sigmocyathacea, Wrighticyathidae fam. nov.), Flexicyathus (Porocoscinidae) and Bractocyathus (Polycoscinidae), and the species Erugatocyathus cymbricensis and Veronicacyathus concavus.     

Wenlock brachiopods from Canberra,Australia

Twenty-four brachiopod species are described from a well-preserved shelly fauna in the Walker Volcanics, collected from the Molonglo valley west of Canberra. There are three inarticulates and twenty-one articulates, of which nine are new, three are closely compared with previously described species and twelve are under open nomenclature. New are Skenidioides thrinax, Salopina mediocostata, Aegiria (Epelidoaegiria subgen. nov.) chilidifera, Morinorhynchus oepiki, Australina (Australina) kausi, Coelospira cavata, Endospirifer anxius, Reticulariopsis silurica and Vadum (gen. nov.) coppinsense. The fauna is shown to be cosmopolitan in character and falls within the scope of Benthic Assemblages 3 to 5 of Boucot (1975). It is of Wenlock age.     

Outline of a Late Cretaceous dinoflagellate zonation of northwestern Australia

The Late Cretaceous and Danian dinoflagellate succession of northwestern Australia can be divided into nine zones, five subzones and one assemblage. In ascending order these are the Diconodinium multispinum Zone, the Palaeohystrichophora infusorioides Zone, the Isabelidinium acuminatum Zone, the Conosphaeridium striatoconus Zone, the Isabelidinium balmei Subzone, the Gillinia hymenophora Subzone, the Areosphaeridium suggestium Zone, the Areoligera coronata Zone, the Samlandia carnarvonensis Zone, the Deflandrea diebelii Zone, the Cladopyxidium foveolatum Subzone, the Alterbidinium acutulum Subzone, the Exochosphaeridium bifidum Subzone, the Alisocysta circumtabulata Zone and the Tectatodinium rugulatum Assemblage. Five new species are described; Areosphaeridium suggestium sp. nov., Cladopyxidium foveolatum sp. nov., Samlandia carnarvonensis sp. nov., Samlandia mayi sp. nov., Samlandia vermicularia sp. nov., and two new combinations proposed; Tectatodinium rugulatum (Hansen) comb. nov. and Membranilarnacia angustivela (Deflandre & Cookson) comb. nov.     

A new Early Permian bivalve fauna from the Río Genoa Formation,Patagonia, Chubut Province,Argentina

Fourteen Bivalvia species are described from the Lower Permian Río Genoa Formation, exposed in the southern Tepuel-Genoa Basin (Chubut Province, Argentina), of which one, Fletcheripecten genoensis, is new. The studied specimens are well preserved, retaining fine details of the ornament and shell morphology. Palaeotaxodonta is represented by two species of Nuculopsis and four species of Phestia; Pteriomorphia by five species in five genera; and Heteroconchia by three informal species. Most of the species recorded also occur in the northern part of Tepuel-Genoa Basin and allow demarcation of new and distinct marine intervals in the Río Genoa Formation, which was classically interpreted to represent continental deposits. The bivalve fauna confirms a Cisuralian (Early Permian) age for the Río Genoa Formation and offers potential for improved regional and global correlations.     

Palaeogeographic setting of a deep-water trilobite fauna from the Early Devonian (Pragian) of the Upper Yarra,central Victoria. Phacopidae of Victoria,Part 3

A Pragian (Early Devonian) trilobite fauna from the Norton Gully Formation in the Upper Yarra area of central Victoria consists exclusively of the phacopid Prokops moorei sp. nov., a species with a highly variable visual surface including greatly reduced and blind morphologies. Several trilobites are preserved as moult assemblages, but most occur as isolated tergites on bedding-planes crowded with dacryoconarids and small bivalves. The autecology and taphofacies of the fauna indicate a deep-water setting, with the biofacies associations closely resembling deep-water assemblages described from Devonian sequences elsewhere. The distributions of laterally equivalent late Pragian facies from eastern areas of the Melbourne Zone indicate an inclined shelf between Lilydale and the Upper Yarra area, deepening eastwards from shoreline to outer shelf settings. To the northwest and northeast of the Upper Yarra area, the shelf was bounded by tectonically active margins associated with the converging Benambra Terrain, and to the south by the Waratah Bay Platform.