South polar greenhouse insects (Arthropoda: Insecta: Coleoptera) from the mid-Cretaceous Tupuangi Formation,Chatham Islands,eastern Zealandia

Stilwell, J.D., Vitacca, J. & Mays, C., April 2016. South polar greenhouse insects (Arthropoda: Insecta: Coleoptera) from the mid-Cretaceous Tupuangi Formation, Chatham Islands, eastern Zealandia. Alcheringa 40, xxx–xxx. ISSN 0311-5518

Rare insect body fossils have been discovered for the first time after 175 years of research on the Chatham Islands, eastern ‘Zealandia’. The coleopteran (beetle) insects, dated to ca 95 Ma and extracted from fine-grained, upper delta plain facies in the lower Upper Cretaceous (Cenomanian–lowermost Turonian) Tupuangi Formation at Waihere Bay on the remote Pitt Island, represent the most southern, polar-latitude (ca 70–80°S) faunal assemblage from the Cretaceous recorded to date. Three species are represented in the insect fauna: a portion of a segmented abdomen of a probable carabid? ground beetle and two distinct coleopteran elytra, one preserved with a brilliantly iridescent carapace upon discovery, comparable with Cretaceous taxa within the Buprestidae (metallic wood borers), but identification with the Chrysomelidae (leaf beetles) or Tenebrionidae (darkling beetles) can not be discounted entirely. Another specimen has more weakly preserved greenish iridescence and has a morphology consistent with Carabidae; given the preservational deficiencies and rarity of material, the specimens are attributed to Buprestidae? genus et species indeterminate and Carabidae? genus et species indeterminate A and B, respectively. These coleopteran fossils represent the only recorded iridescence in Mesozoic invertebrates from Zealandia. Importantly, these mid-Cretaceous insects existed in South Polar forests near the height of the ‘hothouse’ phase of relatively warm, alternating intervals of full daylight in the summer months and total darkness during the winter, before eastern Zealandia diverged at ca 83 Ma from the Marie Byrd Land region of West Antarctica, as part of the final break-up of Gondwana.

Jeffrey D. Stilwell* [jeffrey.stilwell@monash.edu], Jesse V. Vitacca [jesse.vitacca@gmail.com] & Chris Mays [chris.mays@monash.edu], School of Earth, Atmosphere and Environment, Monash University, Clayton VIC 3800, Australia. *Also affiliated with the Australian Museum, 6 College Street, Sydney NSW 2000, Australia.     

A Devonian fish fauna from subsurface sediments in the eastern Officer Basin,South Australia

This paper reports the discovery of three of the most iconic New Caledonian endemic genera, Amphorogyne, Paracryphia and Phelline, as dispersed leaf cuticle fossils in the early Miocene of New Zealand. New Caledonia's endemic angiosperm families have given it a reputation as one of the most interesting botanical regions in the world, but unfortunately it has no known pre-Pleistocene Cenozoic plant fossil record. A once more widespread distribution of its key plants in the context of a cooling and drying Neogene world suggests the current vegetation of New Caledonia is the result of contraction, or even a migration, from more southerly landmasses. Thus, New Zealand may have been a source of at least some of New Caledonia's plants.     

Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand)

Conran, J.G., Bannister, J.M. & Lee, D.E., 2013. Fruits and leaves with cuticle of Laurelia otagoensis sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand). Alcheringa 37, 496–509. ISSN 0311-5518.

Laurelia otagoensis sp. nov. Conran, Bannister & D.E. Lee (Laurales: Atherospermataceae) is described from the earliest Miocene Foulden Maar diatomite deposit, Otago, New Zealand. The new species is represented by mummified fossil leaves with well-preserved cuticle and associated clusters of achenes bearing persistent, long plumose styles. This basal angiosperm family is of significance because of its classic southern disjunctions and ecological importance in extant Gondwana-type rainforests, but has a very sparse fossil record. The present study describes one of very few convincing leaf fossils for Atherospermataceae and the only definitive fossil fruits. The presence of fossil Laurelia in Oligo–Miocene New Zealand combined with fossil leaf impressions from the late Eocene, Miocene dispersed cuticle and pollen from the Oligocene to Holocene shows that the family has had a long history in Cenozoic New Zealand. These new fossils also support palaeoclimatic data suggesting warmer conditions in the earliest Miocene of New Zealand.

John G. Conran [john.conran@adelaide.edu.au], Australian Centre for Evolutionary Biology & Biodiversity and Sprigg Geobiology Centre, School of Earth and Environmental Sciences, Benham Bldg DX 650 312, The University of Adelaide, SA 5005 Australia; Jennifer M. Bannister [jennifer.bannister@xtra.co.nz], Department of Botany, University of Otago, PO Box 56, Dunedin, New Zealand; Daphne E. Lee [daphne.lee@otago.ac.nz], Department of Geology, University of Otago, PO Box 56, Dunedin, New Zealand. Received 17.12.2012; revised 9.4.2013; accepted 15.4.2013.     

A new record of an aristonectine elasmosaurid (Sauropterygia,Plesiosauria) from the Upper Cretaceous of New Zealand: implications for the Mauisaurus haasti Hector, 1874 hypodigm

O’Gorman, J.P., Otero, R.A. & Hiller, N., 2014. A new record of an aristonectine elasmosaurid (Sauropterygia, Plesiosauria) from the Upper Cretaceous of New Zealand: implications for the Mauisaurus haasti Hector, 1874 Hector, J., 1874. On the fossil Reptilia of New Zealand. Transactions and Proceedings of the New Zealand Institute 6, 333–358. [Google Scholar] hypodigm. Alcheringa 38, 504–512. ISSN 0311-5518

An indeterminate aristonectine elasmosaurid is recorded from a lower Maastrichtian bed of the Conway Formation, Waipara River, South Island, New Zealand. The described specimen (CM Zfr 104), previously considered part of the hypodigm of Mauisaurus haasti, came from the upper part of the Alterbidinium acutulum biozone, the same zone from which the only well-known aristonectine from New Zealand, Kaiwhekea katiki, is recorded. The cervical vertebrae of CM Zfr 104 have the same distinctive features (i.e., with extremely broad rather than long centra) as those from previously recorded juvenile aristonectines from Argentina, Chile and Antarctica. This new record is congruent with the biogeographic relationships of Cretaceous marine amniotes from the Weddellian Palaeobiogeographic Province (i.e., Patagonia, western Antarctica, New Zealand and southeastern Australia). Therefore, this type of vertebra is regarded as a distinctive feature of the Weddellian aristonectine elasmosaurids.

José P. O’Gorman [joseogorman@fcnym.unlp.edu.ar], División Paleontología Vertebrados, Museo de La Plata, Universidad Nacional de La Plata, Paseo del Bosque s/n., B1900FWA, La Plata, Argentina; [CONICET (Consejo Nacional de Investigaciones Científicas y Técnicas)]; Rodrigo A. Otero [paracrioceras@gmail.com], Red Paleontológica U-Chile. Laboratorio de Ontogenia y Filogenia, Departamento de Biología, Facultad de Ciencias, Universidad de Chile, Av. Las Palmeras 3425, Santiago, Chile; Norton Hiller, [norton.hiller@canterbury.ac.nz], Department of Geological Sciences, University of Canterbury, PB 4800, Christchurch, 8001, New Zealand and Canterbury Museum, Rolleston Avenue, Christchurch, New Zealand, 8013.     

The record of Araucariaceae macrofossils in New Zealand

Pole, M., December, 2008. The record of Araucariaceae macrofossils in New Zealand. Alcheringa 32, 405–426. ISSN 0311-5518.

The Araucariaceae have a long record in New Zealand, extending back to the Jurassic at least, and Araucaria extends back to at least the Late Cretaceous. This paper reviews the macrofossil record of the family and presents new information based largely on the leaf cuticle record. Agathis, which is the only genus of the family currently growing in New Zealand, has no record before the Cenozoic. All specimens previously identified from pre-Cenozoic strata clearly belong to other taxa or do not show characteristic features of the genus. Araucariaceae macrofossils are virtually ubiquitous in the Cretaceous assemblages of New Zealand but are conspicuous by their absence or rarity in Palaeocene deposits. Their demise may be an expression of events at the Cretaceous–Palaeogene boundary.     

Bryozoans from the late Cretaceous Kahuitara Tuff of the Chatham Islands,New Zealand

Taylor, P.D., & Gordon, D.P., December, 2007. Bryozoans from the Late Cretaceous Kahuitara Tuff of the Chatham Islands, New Zealand. Alcheringa 31, 339-363. ISSN 0311-5518.

Fourteen bryozoan species are described from the Campanian – Maastrichtian Kahuitara Tuff of Pitt Island, substantially increasing the known diversity in this deposit from the two species recorded previously and making it the most diverse bryozoan biota yet described from the Cretaceous of Australasia. Nine of the Kahuitara Tuff bryozoans are cyclostomes, four are cheilostomes, and one is a shell-boring ctenostome. Seven new species are described: Ceriocava hakepaensis sp. nov., Tholopora australis sp. nov., Crisidmonea lanauzeorum sp. nov., Cookobryozoon cretacea sp. nov., Chiplonkarina preeceorum sp. nov. Chiplonkarina bifoliata sp. nov. and Aechmella rangiauriensis sp. nov. The remaining species are left in open nomenclature because of preservational deficiencies or lack of taxon-diagnostic gonozooids. The ctenostome family Cookobryozoidae is subsumed in the Terebriporidae. The new family Chiplonkarinidae is proposed for anascan cheilostomes previously assigned to the paraphyletic Electridae and distinguished by having primarily erect colonies with long, tubular zooids reminiscent of stenolaemates. None of the Kahuitara Tuff bryozoan species is known elsewhere, but all apart from one genus occur in roughly coeval deposits. No families regarded as particularly characteristic of the austral post-Cretaceous are evident. The relatively large number (three) of co-occurring species of Chiplonkarina is notable, as is the dominance of cyclostomes and the first record of Tholopora in the Southern Hemisphere.

Paul D. Taylor [p.taylor@nhm.ac.uk], Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, UK; Dennis P. Gordon [d.gordon@niwa.co.nz], National Institute of Water and Atmospheric Research, Private Bag 14-901, Kilbirnie, Wellington, New Zealand; received 7.3.2006, revised 3.9.2006.     

A fossil sea turtle (Testudines: Pan-Cheloniidae) from the upper Oligocene Pomahaka Formation,New Zealand

Gard, H.J.L. & Fordyce, R.E., August 2016. A fossil sea turtle (Testudines: Pan-Cheloniidae) from the upper Oligocene Pomahaka Formation, New Zealand. Alcheringa 41, XX–XX. ISSN 0311-5518.

An isolated turtle xiphiplastron similar to that of Puppigerus sp. is described from the upper Oligocene (27.3–25.2 Ma) Pomahaka Formation near Tapanui, Otago, New Zealand. The bone is unlike any previously described turtle from the Cenozoic of New Zealand and is from a newly recognized estuarine vertebrate locality. It represents the first Oligocene cheloniid turtle bone described from the southwest Pacific.

Henry J. L. Gard [henrygard@hotmail.com] and R. Ewan Fordyce, [ewan.fordyce@otago.ac.nz], Department of Geology, University of Otago, PO Box 56, Dunedin, 9054, Otago, New Zealand.     

Protodammara reimatamoriori,a new species of conifer (Cupressaceae) from the Upper Cretaceous Tupuangi Formation,Chatham Islands,Zealandia

Mays, C. & Cantrill, D.J., January 2018. Protodammara reimatamoriori, a new species of conifer (Cupressaceae) from the Upper Cretaceous Tupuangi Formation, Chatham Islands, Zealandia. Alcheringa XXX, X–X. ISSN 0311-5518.

Isolated conifer female reproductive structures are common fossil elements from Cenomanian (ca 99–94 Ma) charcoal- and resin-rich beds of the Tupuangi Formation, Chatham Islands, southwest Pacific Ocean. Recent findings have proposed that these are the oldest fossil evidence of serotiny, a highly successful fire-adaptive reproductive strategy common among tree species living in fire-prone areas today. Herein, we systematically describe the external morphological and anatomical features of these fossils, by employing a combination of manual extraction and neutron tomography techniques. We propose a new species of conifer, Protodammara reimatamoriori, and a re-examination of fossil material of the Protodammara type species facilitated an emendation of the genus. Protodammara shares numerous features with extant Cunninghamia, Taiwania, Athrotaxis, and several extinct taxa of Cupressaceae, and is interpreted as an extinct lineage of the early-divergent ‘taxodioid Cupressaceae’ stem group.

Chris Mays [chris.mays@nrm.se] Department of Palaeobiology, Swedish Museum of Natural History, Frescativägen 40, Stockholm 114 18, Sweden; School of Earth, Atmosphere and Environment, Monash University, 9 Rainforest Walk, Clayton, VIC 3800, Australia; David J. Cantrill [david.cantrill@rbg.vic.gov.au] Royal Botanic Gardens Victoria, Private Bag 2000, South Yarra, VIC 3141, Australia; School of BioSciences, University of Melbourne, Parkville, VIC 3010, Australia.     

Deciduous Nothofagus leaves from the Miocene of Cornish Head,New Zealand

Pole, M.S., 1994:03:28. Deciduous Nothofagus leaves from the Miocene of Cornish Head, New Zealand. Alcheringa 18, 79–83. ISSN 0311-5518.

Nothofagus leaves with plicate vernation, indicating a deciduous habit, are recorded from the Late Miocene of New Zealand. This is the first unequivocal record in New Zealand of deciduous Nothofagus, deciduous species now being restricted to Tasmania and South America.     

Fossil arachnids from the earliest Miocene Foulden Maar Fossil-Lagerstätte,New Zealand

Selden, P.A. & Kaulfuss, U., May 2018. Fossil arachnids from the earliest Miocene Foulden Maar Fossil-Lagerstätte, New Zealand. Alcheringa XX, XX–XX. ISSN 0311-5518.

Fossil Arachnida from New Zealand are extremely rare and represented by some unidentifiable amber inclusions only. The first fossil arachnids from New Zealand to be described in detail are presented here, based on four compression fossils from the earliest Miocene Fossil-Lagerstätte at Foulden Maar, South Island. Two specimens are referred to Arachnida incertae sedis and Araneomorpha incertae sedis, respectively, whereas two specimens are mygalomorph spiders. One of these is placed in the Rastelloidina clade of Mygalomorphae, probably in the Idiopidae, which is represented in New Zealand by the extant Cantuaria.

P.A. Selden* [selden@ku.edu] Paleontological Institute and Department of Geology, University of Kansas, Lindley Hall, 1475 Jayhawk Boulevard, Lawrence, KS 66045, USA; U. Kaulfuss [uwe.kaulfuss@otago.ac.nz] Department of Geology, University of Otago, PO Box 56, Dunedin, New Zealand. *Also affiliated with: Natural History Museum, London, UK.     

Conodonts and tabulate corals from the Upper Ordovician Angullong Formation of central New South Wales,Australia

Zhen, Y.Y., Wang, G.X. &; Percival, I.G., August 2016. Conodonts and tabulate corals from the Upper Ordovician Angullong Formation of central New South Wales, Australia. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

The Angullong Formation is the youngest Ordovician unit exposed in the Cliefden Caves area of central New South Wales. Its maximum age is constrained by a Styracograptus uncinatus graptolite Biozone fauna at the very top of the underlying Malongulli Formation, but the few fossils previously reported from higher in the Angullong Formation are either long-ranging or poorly known. From allochthonous limestone clasts in the middle part of the formation, we document a conodont fauna comprising Aphelognathus grandis, A. solidum, Aphelognathus sp., Aphelognathus? sp., Belodina confluens, Drepanoistodus suberectus, Panderodus gracilis, Panderodus sp., Phragmodus undatus, Pseudobelodina inclinata and Pseudobelodina? sp. aff. P. obtusa, which supports correlation with the Aphelognathus grandis Biozone (late Katian) of the North American Midcontinent succession. The species concepts of Aphelognathus and Pseudobelodina are reviewed in detail. Associated corals are exclusively tabulates, dominated by agetolitids, including Agetolites angullongensis sp. nov., Heliolites orientalis, Hemiagetolites breviseptatus, Hemiagetolites sp. cf. H. spinimarginatus, Navoites sp. cf. N. circumflexa, Plasmoporella bacilliforma, P. marginata, Quepora sp. cf. Q. calamus and Sarcinula sp. Affinities of the coral fauna from the Angullong Formation are closer to faunas from northern NSW and northern Queensland than to the locally recognized Fauna III of late Eastonian age in central NSW. We propose a subdivision of Fauna III to account for this difference, with the late Katian Fauna IIIB characterized by the incoming of agetolitid corals. The currently known distribution of representatives of this group with adequate age constraints suggests that agetolitids possibly originated in North China, subsequently migrating to Tarim, South China and adjacent peri-Gondwanan terranes while also spreading eastward to northern Gondwana, where they progressively moved through eastern Australia to reach the central NSW region by the early Bolindian.

Yong Yi Zhen* (yong-yi.zhen@industry.nsw.gov.au) and Ian G. Percival (ian.percival@industry.nsw.gov.au), Geological Survey of New South Wales, W.B. Clarke Geoscience Centre, 947–953 Londonderry Road, Londonderry, NSW 2753, Australia; Guangxu Wang (gxwang@nigpas.ac.cn), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road Nanjing 210008 PR China.     

Editorial

Similarities in late Middle Cambrian trilobite, inarticulate brachiopod and molluscan faunas are apparent between Bornholm (Denmark) and New Zealand. Tuarangia gravgaerdensis sp. nov. is described from the late Middle Cambrian Andrarum Limestone of Bornholm, Denmark. The Andrarum Limestone is stratigraphically correlated with the Tasman Formation, New Zealand, from where the genus Tuarangia was first described. A single specimen, referred to Tuarangia, has been found in an erratic boulder from NW Poland. The boulder is estimated to be of early Late Cambrian age. The initial taxonomic assignment of Tuarangia to the Bivalvia, Subclass Pteriomorphia, is upheld. The distance between Bornholm and New Zealand, the position at opposite hemispheres, and the palaeogeography in general clearly indicate isocommunities with restricted possibility of exchange of the gene pool at species level.     

Early Carboniferous brachiopod faunas from the Baoshan block,west Yunnan,southwest China

38 brachiopod species in 27 genera and subgenera are described from the Yudong Formation in the Shidian-Baoshan area, west Yunnan, southwest China. New taxa include two new subgenera: Unispirifer (Septimispirifer) and Brachythyrina (Longathyrina), and seven new species: Eomarginifera yunnanensis, Marginatia cylindrica, Unispirifer (Unispirifer) xiangshanensis, Unispirifer (Septimispirifer) wafangjieensis, Brachythyrina (Brachythyrina) transversa, Brachythyrina (Longathyrina) baoshanensis, and Girtyella wafangjieensis. Based on the described material and constraints from associated coral and conodont faunas, the age of the brachiopod fauna from the Yudon Formation is considered late Tournaisian (Early Carboniferous), with a possibility extending into earlyViséan.     

Thelodont fish and conodonts from the Early Devonian of Reefton,New Zealand

Thelodont scales and associated fragments of conodont elements identified as Turinia sp. cf. Turinia australiensis and Ozarkodina sp. cf. Ozarkodina buchanensis respectively are described for the first time from the Early Devonian Murray Creek Formation of Reefton, New Zealand. The conodont and thelodont components of the fauna suggest a late Pragian to early Emsian age for the assemblage.     

A new early Paleocene dinoflagellate cyst species,Trithyrodinium partridgei: its biostratigraphic significance and palaeoecology

A palynological investigation of lower Paleocene (lower Teurian) marine sediments of the Abbotsford Formation, exposed in the Fairfield Quarry, Dunedin, New Zealand, reveals a well-preserved association of dinoflagellate cysts and miospores. Thirty-six dinoflagellate species representing 26 genera were recorded, including one new peridinioid dinoflagellate species, Trithyrodinium partridgei characterized by a three-layered wall and a distinct brown, foveolate mesophragm. Trithyrodinium partridgei has a narrow range and at Fairfield Quarry co-occurs with typical New Zealand late Danian to Selandian index species such as Glaphyrocysta perforata and Alterbidinium pentaradiatum; hence it may prove an additional useful stratigraphic marker. The terrestrial palynomorph assemblage from samples within the range of Trithyrodinium partridgei includes 36 spore-pollen species dominated by the podocarp pollen Phyllocladidites mawsonii. The presence of Nothofagidites waipawaensis and Tricolpites phillipsii confirms a Paleocene age for the assemblage. The abundance of terrestrial palynomorphs and the composition of the dinocyst assemblages suggest a marine nearshore depositional environment.     

A new dinoflagellate species from the Upper Cretaceous of New Zealand - a morphological intermediate between three genera

The new dinoflagellate Isabelidinium marshallii sp. nov. was encountered in the lower to middle Campanian Satyrodinium haumuriense Interval Zone and in the middle to upper Campanian Isabelidinium korojonense Interval Zone, in southern Marlborough, South Island, New Zealand. The new taxon is attributed to Isabelidinium but it also closely resembles species of Alterbidinium and Satyrodinium. Despite its close morphological affiliation to three dinoflagellate genera, I. marshallii represents a discrete population of peridinioid cysts that has a stratigraphically useful range in New Zealand.     

Tertiary megafossil flora of Maslin Bay,South Australia: a preliminary report

The Maslin Bay flora of South Australia is of lower Middle Eocene age and contains diverse, well preserved angiosperm assemblages. It has yielded 2700 specimens belonging to approximately 200 leaf taxa.

Physiognomic analysis (sensu Webb, 1959; Wolfe 1970) indicates greatest similarity to extant Simple Mesophyll Vine Forest and Complex Notophyll Vine Forest found currently in Queensland.

Preliminary taxonomic studies have confirmed the presence of taxa closely resembling Podocarpus (Podocarpaceae), Agathis (Araucariaceae), Fatsia (Araliaceae) and Banksia (Proteaceae). Microfloral analysis of the deposit confirms the presence of the latter three families, but suggests far greater occurrence of Proteaceae than the number of leaf specimens of this family indicates. Nothofagus pollen is the dominant type, although leaves of this genus are absent from collections. Based on sedimentation, physiognomy and systematic studies of leaves and fungi (Lange, 1969) it is concluded that the Maslin Bay region supported tropical to subtropical rain forest during the lower Middle Eocene.     

Probable oribatid mite (Acari: Oribatida) tunnels and faecal pellets in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation,central-western Queensland,Australia

Fletcher, T.L. & Salisbury, S.W., XX.XX. 2014. Probable oribatid mite (Acari: Oribatida) tunnels and faecal pellets in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, central-western Queensland, Australia. Alcheringa 38, 541–545. ISSN 0311-5518.

Tunnels and faecal pellets likely made by oribatid mites have been found in silicified conifer wood from the Upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, central-western Queensland, Australia. Although this is the first identified and described record of oribatid mites in the Mesozoic of Australia, other published, but unassigned material may also be referable to Oribatida. Current understanding of the climatic significance of mite distribution is limited, but the presence of oribatids and absence of xylophagus insects in the upper portion of the Winton Formation are consistent with indications that the environment in which this unit was deposited was relatively warm and wet for its palaeolatitude. Such traces may provide useful and durable proxy evidence of palaeoclimate, but more detailed investigation of modern taxa and their relationship to climate is still needed.

Tamara L. Fletcher [t.fletcher1@uq.edu.au] and Steven. W. Salisbury, [s.salisbury@uq.edu.au] School of Biological Sciences, The University of Queensland, Australia, 4072. Received 28.1.2014; revised 1.4.2014; accepted 3.4.2014.     

A palynological investigation of plesiosaur-bearing rocks from the Upper Cretaceous Tahora Formation,Mangahouanga, New Zealand

High-palaeolatitude plesiosaur, mosasaur and, more rarely, dinosaur fossils are well known from the Maungataniwha Sandstone Member of the Tahora Formation in Mangahouanga Stream, Hawkes Bay, New Zealand. A palynological investigation of strata exposed along Mangahouanga Stream and of transported boulders hosting vertebrate fossils reveals well-preserved assemblages dominated by terrestrial pollen and spores but also including marine dinoflagellate cysts in some samples. The palynofacies are strongly dominated by wood fragments including charcoal; one outcrop sample and the sample taken from a boulder hosting plesiosaur vertebrae contain entirely terrestrially derived palynoassemblages, suggesting a freshwater habitat for at least some of the plesiosaurs. The host unit spans the Santonian to lowermost Maastrichtian, while the key pollen taxa Nothofagidites senectus and Tricolpites lilliei, together with the dinocyst Isabelidinium pellucidum and the megaspore Grapnelispora evansii, indicate a late Campanian to early Maastrichtian age for the fossiliferous boulders. The palynoflora indicates a mixed local vegetation dominated by podocarp conifers and angiosperms with a significant tree-fern subcanopy. The presence of taxa with modern temperate distributions, such as Nothofagus (southern beech), Proteaceae and Cyatheaceae (tree-ferns), indicates a mild-temperate climate and lack of severe winter freezing during the latest Cretaceous, providing an ecosystem that most probably made it possible for polar dinosaurs to overwinter in this part of the world.     

Kāi Tahu me te Hopu Tītī ki Rakiura:

This article examines te hopu tītī ki Rakiura — the customary harvesting of tītī or ‘muttonbirds’ (sooty shearwaters/puffinus griseus) from islands adjacent to Rakiura (Stewart Island), by members of Kāi Tahu — the iwi (tribe) that has traditional authority over the majority of Te Wāhi Pounamu (the South Island of New Zealand). The article illustrates the pre and post-contact importance of te hopu tītī to Kāi Tahu and argues that, because the harvest is now and has always been the sole domain of the iwi, this sets it apart from prevailing settler society narratives whereby indigenous people usually lose out. The article also shows how the harvest did and continues to contribute to Kāi Tahu tribal identity. The author is both Kāi Tahu and an active participant in the tītī harvest as well as a post-graduate history student. This enables him to offer both a unique reading of the archives relating to the harvest as well as access to oral histories and early photographs associated with it.