Two new Middle Jurassic species of orthophlebiids (Insecta: Mecoptera) from Inner Mongolia,China

QIAO, X., SHIH, C.K. & REN, D., December 2012. Two new Middle Jurassic species of orthophlebiids (Insecta: Mecoptera) from Inner Mongolia, China. Alcheringa 36, 467–472. ISSN 0311-5518.

Two new species of the extinct family Orthophlebiidae, Orthophlebia nervulosa sp. nov. and Orthophlebia stigmosa sp. nov., are described and illustrated. These well-preserved specimens were collected from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Inner Mongolia, China. Morphological characters shown in these well-preserved specimens highlight the diversity of orthophlebiids during the Middle Jurassic and provide data for future phylogenetic studies of orthophlebiids.     

Two new species of Mesosciophilidae (Insecta: Diptera: Nematocera) from the Yanliao biota of Inner Mongolia,China

Wang, Q., Zhao, Y.Y. & Ren, D., December 2012. Two new species of Mesosciophilidae (Insecta: Diptera: Nematocera) from the Yanliao biota of Inner Mongolia, China. Alcheringa 36, 509–514. ISSN 0311-5518.

Two new species assigned to Mesosciophila Rohdendorf and Paramesosciophilodes Zhang in the Mesosciophilidae are described and illustrated: Mesosciophila sigmoidea sp. nov. and Paramesosciophilodes aequus sp. nov. They are established based on fossil specimens with bodies and complete wings. Both were collected from the Middle Jurassic, Jiulongshan Formation at Daohugou, Inner Mongolia, China.     

Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia,China

SHI, C.F., WANG, Y.J., YANG, Q. & REN, D., September 2012. Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China. Alcheringa 36, 311–320. ISSN 0311-5518.

A new grammolingiid genus, Chorilingia containing four new species (C. euryptera, C. parvica, C. translucida and C. peregrina) is described and illustrated from the Jiulongshan Formation at Daohugou, Inner Mongolia, China. The new genus is differentiated mainly on the first branch of vein Rs (Rs1) separating distal to the forks of both CuA and CuP. A key to species of the genus is provided.     

Poljanka strigosa,a new species of Protopsyllidiidae (Hemiptera,Sternorrhyncha) from the Middle Jurassic of China

Yang, G., Yao, Y.Z. & Ren, D., iFirst. Poljanka strigosa, a new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China. Alcheringa, 1–6. ISSN 0311-5518.

A new fossil species, Poljanka strigosa sp. nov., of the extinct family Protopsyllidiidae is described from the Middle Jurassic Jiulongshan Formation of Daohugou Village, Inner Mongolia, China. The new species is characterized by wings bearing long, stiff setae that are evident as stained impressions in the fine sedimentary rock. Comparison between Protopsyllidiidae and extant psylloids suggests that Protopsyllidiidae is probably closely related to extant psylloids.     

New fossil sawflies (Hymenoptera,Xyelidae) from the Middle Jurassic of northeastern China

Zheng, Y., Chen, J., Zhang, J. & Zhang, H., 6 August 2019. New fossil sawflies (Hymenoptera, Xyelidae) from the Middle Jurassic of northeastern China. Alcheringa 44, 115–120. ISSN 0311-5518.

Magnaxyela rara gen. et sp. nov. and Abrotoxyela curva sp. nov. (Xyelidae, Macroxyelinae, Gigantoxyelini) are described from the Middle Jurassic Jiulongshan Formation at Daohugou Village, Inner Mongolia, China. The two new species are confidently assigned to tribe Gigantoxyelini of subfamily Marcroxyelinae based on their wing venation and various other morphological characters. Magnaxyela rara is established based on the following forewing characters: pterostigma sclerotized completely, Sc two-branched and meeting R before origin of Rs, 1-Rs as long as 1-M, but shorter than Rs?+?M, R curved bewteen 1r-rs and 2r-rs and cell 2r longer than 1r. Abrotoxyela curva is differentiated from congeneric forms by the two-branched vein Sc of the forewing, Sc2 short and inclined towards the base of the wing, 1-Rs, 1-M and Rs?+?M nearly equal in length, crossvein 1m-cu longer than 1cu-a. The fossils described herein increase the diversity of the Mesozoic Xyelidae.

Yan Zheng* [zhengyan536@163.com], Jun Chen* [ubiscada@sina.com], and Junqiang Zhang [zhangjunqiang@lyu.edu.cn], Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi 276000, PR China; Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, PR China. *Also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, PR China.     

Two new species of Nevania (Hymenoptera: Evanioidea: Praeaulacidae: Nevaniinae) from the Middle Jurassic of China

Two new species of Nevania Zhang & Rasnitsyn 2007 Zhang, H.C. & Rasnitsyn, A.P., 2007. Nevaniinae subfam. n., a new fossil taxon (Insecta: Hymenoptera: Evanioidea: Praeaulacidae) from the Middle Jurassic of Daohugou in Inner Mongolia, China. Insect Systematics & Evolution 38, 149–166.[Crossref], [Web of Science ®] , [Google Scholar], Nevania perbella sp. nov. and Nevania aspectabilis sp. nov., are described and illustrated. They are assigned to Nevania based on their unique morphological characters: forewing venation complete; 2A complete; metasoma with first and second segments long and tube-like. A key to the known species of Nevania is provided. These fossils, collected from the Jiulongshan Formation (Middle Jurassic) of Daohugou in Inner Mongolia, China, broaden the known diversity of this genus in mid-Mesozoic ecosystems.     

Application of geometric morphometric analyses to confirm two new species of Karatavitidae (Hymenoptera: Karatavitoidea) from northeastern China

SHIH, M.J.H., LI, L.F. & REN, D., April 2017. Application of geometric morphometric analyses to confirm two new species of Karatavitidae (Hymenoptera:Karatavitoidea) from northeastern China. Alcheringa 41, 499–508. ISSN 0311-5518

Two new species, Praeratavites rasnitsyni and Karatavites ningchengensis in the wasp family Karatavitidae, are described and illustrated. These specimens were collected from the uppermost Middle Jurassic Jiulongshan Formation at Daohugou Village in Inner Mongolia, China. Key venation characters, as measured by relative positions or length ratios, are summarized for all documented species of Karatavites and Praeratavites to facilitate morphological comparisons and assess whether the studied specimens are referable to new species. In addition, geometric morphometric analyses were conducted to confirm the distinction of the new species, which add to the diversity of these genera in the Middle Jurassic.

Matthew J.H. Shih* [mpshih88@gmail.com] Union County Magnet High School, Scotch Plains, NJ 07076, USA; Longfeng Li [fenger4499@163.com] and Dong Ren [rendong@mail.cnu.edu.cn] College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, PR China. *Also affiliated with: Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA.     

A new species of Aboilus (Insecta,Orthoptera) from the Jurassic Daohugou beds of China,and discussion of forewing coloration in Aboilus

Wang, H., Li, S., Zhang, Q., Fang, Y., Wang, B. & Zhang, H., 13.02.2015. A new species of Aboilus (Insecta, Orthoptera) from the Jurassic Daohugou beds of China, and discussion of forewing coloration in Aboilus. Alcheringa 39, xxx–xxx. ISSN 0311-5518

He Wang* [wanghe0701@163.com], Sha Li* [lisha.paleo@gmail.com], Qi Zhang* [zhqi1105@126.com], Yan Fang [yanfang@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.*Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China. ?Also affiliated with Steinmann Institute, University of Bonn, Bonn 53115, Germany.

A new species of Aboilinae (Orthoptera: Prophalangopsidae), Aboilus perbellus, is described and illustrated based on three well-preserved forewings recovered from the Middle–Upper Jurassic Daohugou beds of Inner Mongolia, China. The new species differs from all congeneric forms in its special forewing coloration and features of its wing venation. To date, three types of forewing coloration have been found among different species of Aboilus at Daohugou, suggesting that these taxa inhabited different ecotopes.     

A new fossil sinoalid species from the Middle Jurassic Daohugou beds (Insecta: Hemiptera: Cercopoidea)

Fu, Y., Cai, C. & Huang, D., October 2017. A new fossil sinoalid species from the Middle Jurassic Daohugou beds (Insecta: Hemiptera: Cercopoidea). Alcheringa 42, 94–100. ISSN 0311-5518.

A new fossil species, Luanpingia daohugouensis sp. nov., belonging to the family Sinoalidae is described from the Middle to Upper Jurassic Daohugou beds of Inner Mongolia, China, on the basis of two well-preserved complete specimens. The described species of Sinoalidae are reviewed and Jiania gracila is considered a junior synonym of Jiania crebra. The new discovery increases the palaeodiversity of sinoalids from the Daohugou beds. It also indicates stratigraphic correlation between the Daohugou beds, the Haifanggou Formation at Haifeng, Beipiao City, West Liaoning Province, and the Jiulongshan Formation at Zhouyingzi, Luanping County, Hebei Province. All of these units host the ‘early assemblage’ of the Yanliao biota.

Yanzhe Fu [yzfu41@163.com], Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; University of Science and Technology of China, Hefei 230026, PR China; Chenyang Cai [cycai@nigpas.ac.cn], Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; Diying Huang* [dyhuang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.     

New sinoalids (Insecta: Hemiptera: Cercopoidea) from Middle to Upper Jurassic strata at Daohugou,Inner Mongolia,China

Fu, Y., & Huang, D., December 2018. New sinoalids (Insecta: Hemiptera: Cercopoidea) from Middle to Upper Jurassic strata at Daohugou, Inner Mongolia, China. Alcheringa 43, 246–256. ISSN 0311-5518.

A new fossil genus with three species of Sinoalidae, Parasinoala daohugouensis gen. et sp. nov., Parasinoala minuta gen. et sp. nov., Parasinoala magnus gen. et sp. nov., from the Middle to Upper Jurassic Haifanggou Formation at Daohugou, Inner Mongolia, northeastern China, is described and illustrated. The new genus differs from other sinoalids by the tegmen with RA having 2–3 branches, MP with 3–4 branches and MP of hind wings with two branches, which indicates the terminal branches of RA and MP are highly variable within Sinoalidae. The new discovery greatly increases the palaeodiversity of sinoalids in the early assemblage of the Yanliao biota from the Daohugou beds.

Yanzhe Fu [yzfu@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; University of Science and Technology of China, Hefei 230026, PR China; Diying Huang* [dyhuang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.     

A new saucrosmylid lacewing (Insecta,Neuroptera) from the Middle Jurassic of Daohugou,Inner Mongolia,China

Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A., 2014. A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa 38. ISSN 0311-5518.

A new genus and new species of Saucrosmylidae (Insecta, Neuroptera) are described (Daohugosmylus castus) based on a well-preserved hindwing from the Middle Jurassic of Daohugou, Inner Mongolia, China. Daohugosmylus gen. nov. is distinguished by a large and nearly semi-circular hindwing, relatively wide R1 space possessing several rows of cells, anteriorly bent Rs, dense crossveins over the entire wing, and smooth outer margin.

Qing Liu (corresponding author) [qingliu1029@gmail.com], Haichun Zhang [hczhang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], Yan Fang [yanfang@nigpas.ac.cn], Daran Zheng [dranzheng@gmail.com], Qi Zhang [zhqi1105@126.com] and Edmund A Jarzembowski [jarzembowski2@live.co.uk], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China; secondary address of Daran Zheng & Qi Zhang, University of Chinese Academy of Sciences, Beijing, 100049, PR China; and Ed Jarzembowski, Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. Received 13.11.2013; revised 20.1.2014; accepted 21.1.2014.     

Progonocimicids (Hemiptera,Coleorrhyncha) from the Middle Jurassic Haifanggou Formation,western Liaoning,northeast China support stratigraphic correlation with the Daohugou beds

Jiang, J.-Q., Cai, C.-Y. & Huang, D.-Y., October 2015. Progonocimicids from the Middle Jurassic Haifanggou Formation, western Liaoning, northeast China support stratigraphic correlation with the Daohugou beds. Alcheringa 40, XXX–XXX. ISSN 0311-5518.

The hemipteran suborder Coleorrhyncha includes only 37 extant species assigned to the family Peloridiidae. However, the suborder’s fossil record is diverse and abundant. The extinct family Progonocimicidae is very common in Middle Jurassic strata of northeastern China, especially in the Daohugou beds of Ningcheng County, Inner Mongolia, and the Haifanggou Formation in Beipiao, Liaoning Province. We re-studied the established progonocimicid species and examined 27 new specimens, indicating that the species from Daohugou are junior synonyms of those from Haifanggou. The progonocimicids from the Haifanggou Formation are assigned to two species of Cicadocoris: C. brunneus (=Mesoscytina brunnea, =Mesocimex lini) and C. sinensis (=Cicadocoris anisomeridis). Both species are common in the Daohugou beds and the Haifanggou Formation. This discovery is of significance for biostratigraphic correlation of these two lithostratigraphic units. It also contributes to our understanding of the geological age of the famous Daohugou biota, which has yielded the earliest known feathered dinosaurs and diverse early mammals.

Jia-Qian Jiang [jiaqianjiang@126.com], State Key Laboratory of Palaeobiology and Stratigraphy; Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; and Graduate School, University of Chinese Academy of Sciences, 19A Yuquanlu, Beijing 100049, PR China. Chen-Yang Cai [caichenyang1988@163.com], Di-Ying Huang [dyhuang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.     

The oldest root-eating beetle from the Middle Jurassic of China (Coleoptera,Monotomidae)

Cai, C.-Y., ?lipiński, A. & Huang, D.-Y., 31.3.2015. The oldest root-eating beetle from the Middle Jurassic of China (Coleoptera, Monotomidae). Alcheringa 39,488–493. ISSN 0311-5518.

Jurorhizophagus alienus gen. et sp. nov., a new fossil root-eating beetle, is described and figured based on an exceptionally well-preserved impression fossil from the Middle Jurassic Daohugou beds (ca 165 Ma), Inner Mongolia, northeastern China. It represents the earliest fossil Monotomidae known to date. Jurorhizophagus can not be assigned to either of two subfamilies Monotominae or Rhizophaginae based on the unique combination of many unusual characters, including an 11-segmented antenna with a 3-segmented club, the presence of a distinct frontoclypeal suture and transverse pronotum with a median longitudinal groove. The discovery of a new genus from the Middle Jurassic highlights the antiquity of Monotomidae and provides new information about the phylogenetic relationships between Monotomidae and its allied families.

Chen-Yang Cai [caichenyang1988@163.com], Di-Ying Huang [dyhuang@nigpas.ac.cn] (corresponding author), State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China; Adam ?lipiński [adam.slipinski@csiro.au], Australian National Insect Collection, CSIRO National Collections Australia, GPO Box 1700, Canberra ACT 2601, Australia.     

A hairy-bodied tettigarctid (Hemiptera: Cicadoidea) from the latest Middle Jurassic of northeast China

Liu, X.H., Li, Y., Yao, Y.Z. & Ren, D., April 2016. A hairy-bodied tettigarctid (Hemiptera: Cicadoidea) from the latest Middle Jurassic of northeast China. Alcheringa 40, xxx–xxx. ISSN 0311-5518

Extant tettigarctids are also known as hairy cicadas because they are covered by long and abundant hairs. This character had not been reported in fossil species of Tettigarctidae because previous examples were poorly preserved or lacked long hairs. Hirtaprosbole erromera gen. et sp. nov. (Tettigarctidae) with a hairy body, from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China, is described here. This new species provides evidence that tettigarctids with long dense hairs had appeared by the latest Middle Jurassic and lived at high altitudes.

Xiao-hui Liu [liuxh8917@163.com], Yi Li [liy0124@sina.com], Yun-zhi Yao*[Corresponding author: yaoyz100@gmail.com] and Dong Ren [rendong@mail.cnu.edu.cn], College of Life Sciences, Capital Normal University, Xisanhuanbeilu 105 Haidian District, Beijing, PR China 100048.     

New Archaeocyatha from the Early Cambrian of the Mt. Wright area,New South Wales

An archaeocyathan fauna from a single limestone lens in the upper Cymbric Vale Formation at Mt. Wright shows species in common with the historic Ajax Mine fauna of the Flinders Ranges, South Australia. The Cymbric Vale fauna is considered equivalent to the lower Sanashtykgol ‘horizon’ of the Altay Sayan fold belt of Siberia.

New taxa described are the genera Hyptocyathus (Inessocyathacea, Hyptocyathidae fam. nov.), Aporosocyathus (Annulocyathidae), Wrighticyathus (Sigmocyathacea, Wrighticyathidae fam. nov.), Flexicyathus (Porocoscinidae) and Bractocyathus (Polycoscinidae), and the species Erugatocyathus cymbricensis and Veronicacyathus concavus.     

Small and common: the oldest tropical Chrysomelidae (Insecta: Coleoptera) from the lower Eocene Cambay amber of India

Nadein, K.S. & Perkovsky, E.E. 2 July 2019. Small and common: the oldest tropical Chrysomelidae (Insecta: Coleoptera) from the lower Eocene Cambay amber of India. Alcheringa XXX, X–X. ISSN 0311-5518.

Three new genera and species of flea beetles (Chrysomelidae: Alticini) are described from the lowermost Eocene Cambay amber: Cambaltica paleoindica Nadein, gen. et sp. nov., Protorthaltica setosella Nadein, gen. et sp. nov., and Davidaltica cambayensis Nadein, gen. et sp. nov. These taxa share a zoogeographic affinity with extant Oriental and Afrotropical flea beetle faunas, with similarities to Afrotropical elements interpreted to be a result of Neogene migrations from Laurasia to Africa. The flea beetles within the Cambay amber are characterized by their small body size (1.2–1.9?mm), and the absence or rarity of larger flea beetles in the Cambay amber forest is assumed to be evidence for a progressive increase in the average body size of tropical flea beetles beginning in the early Eocene.

Konstantin S. Nadein [k.nadein@gmail.com], University of Kiel, Zoological Institute, Functional Morphology and Biomechanics, Am Botanischen Garten 1–9, Kiel, Germany; Evgeny E. Perkovsky [perkovsk@gmail.com], Schmalhausen Institute of Zoology, National Academy of Science of Ukraine, Bogdana Khmelnitskogo Str. 15, Kiev, Ukraine, Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya Str. 123, Moscow, 117997, Russia.     

The first damsel-dragonfly (Odonata: Isophlebioidea: Campterophlebiidae) from the Middle Jurassic of Shaanxi Province,northwestern China

ZHENG, D., DONG, C., WANG, H., YE, Y., WANG, B., CHANG S-C. & ZHANG, H., May 2017. The first damsel-dragonfly (Odonata: Isophlebioidea:Campterophlebiidae) from the Middle Jurassic of Shaanxi Province, northwestern China. Alcheringa 41, 509–513. ISSN 0311-5518.

Campterophlebiidae is the most diverse family of fossil odonatans in China with ten genera recovered mostly from Middle Jurassic strata of Inner Mongolia. We describe a well-preserved campterophlebiid damsel-dragonfly from the Middle Jurassic Yanan Formation in Shanxi Province, northwestern China. This discovery adds to the diversity of Campterophlebiidae and identifies a new Middle Jurassic insect fossil locality in China. Within Campterophlebiidae, the new taxon most closely resembles Ctenogampsophlebia from the Middle Jurassic of Inner Mongolia but differs from other genera in having vein AA with four parallel posterior branches uncrossed in the anal triangle.

Daran Zheng [dranzheng@gmail.com] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China; Chong Dong [cdong@nigpas.ac.cn], He Wang [hwang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Yifei Ye [459353251@qq.com] Shannxi Non-ferrous Yulin Coal Co., Ltd, Yulin, PR China; Bo Wang [bowang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China; Su-Chin Chang [suchin@hku.hk] Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China.     

Significance of new species of Cryptorhynchia (Brachiopoda) from the Middle Jurassic of Kutch,India

Cryptorhynchia is a brachiopod genus, until recently known only from the late Bathonian. Two new species C. karuna and C. jhooraensis are here described from the middle Bathonian at Kutch, western India and record the earliest known occurrences of the genus. They constitute ancestor-descendant lineages with the two existing younger species C. pulcherrima and C. rugosa respectively. Evolution in both cases shows ‘parallel’ trends and the descendants are scaled-down versions of their respective progenitors. Statistical analyses reveal that this evolutionary miniaturization involves allometry-induced heterochrony, especially progenesis, and is marked by a rapid speciation event compatible with the punctuational model. Evolution appears to be anagenetic and may be attributed to the unstable nature of environment where onshore innovation produced smaller descendants that adopted r-strategy. Facies distribution and functional morphology suggest that species of Cryptorhynchia were well adapted to shallow, unstable carbonate shelf. However, they disappeared suddenly at the Bathonian - Callovian boundary which was marked by a global transgression. The possible causal factors of their extinction may be the consequent changes in bathymetry and substrate condition.