The record of fossil fish from the Pleistocene of Argentina is poor. Here we describe the first ichthyofauna from Late Pleistocene riverbank beds in the Salado River of Santa Fe Province, Argentina. The material consists of isolated pectoral and dorsal fin spines, together with skull fragments. Four species-level taxa referable to three families can be identified: Pterodoras granulosus (Doradidae), Pimelodus cf. maculatus and Pimelodus cf. albicans (Pimelodidae) and cf. Hypostomus sp. (Loricariidae). Specimens attributed to Pterodoras granulosus and Pimelodus maculatus represent a minimum age for origin of these taxa. The Salado River assemblage includes the richest record of Pleistocene catfishes yet documented from southern South America.
Evelyn Romina Vallone [evelynvallone@conicet.
Selenariidae Busk 1854 (Bryozoa) is considered endemic to Australia and New Zealand. Here we describe a new species of Selenaria Busk 1854 from the lower Miocene Monte León Formation (Patagonia, Argentina). Selenaria lyrulata sp. nov. is characterized by autozooids with a lyrula-like, anvil-shaped cryptocystal denticle, opesiular indentations and lateral condyles, as well as avicularia with a shield of fused costae. This is the first record of a selenariid bryozoan in South America.
Juan López-Gappa [lgappa@macn.
Typical glacial–postglacial sequences associated with the Late Palaeozoic Ice Age (LPIA) are recognized in the Calingasta-Uspallata Basin, central-western Argentina, particularly in the Hoyada Verde and El Paso formations (late Serpukhovian–Bashkirian) at Barreal Hill (San Juan province). Brachiopods and bivalves accompanied by gastropods, conulariids, nautiloids, corals and ostracods constitute the marine assemblages of the El Paso Formation. They are assigned to the Aseptella–Tuberculatella/Rhipidomella–Micraphelia (AT/RM) fauna, characterized by two fossil assemblages: Aseptella–Tuberculatella, identified in the lower fossiliferous interval, and Rhipidomella–Micraphelia in the upper. The development of the different invertebrate assemblages within the El Paso Formation, and their relationship with coeval suite in the Hoyada Verde Formation, can be explained by a complex array of abiotic factors (substrate stability, turbidity, nutrient availability, variation in oxygen levels, poor circulation and salinity variations in the water column) that were directly related to glacial retreat dynamics and coastal configuration. A restricted palaeofjord setting is proposed for the depositional environment of the El Paso Formation in contrast to an exposed open marine coast with a gently sloping shelf for the Hoyada Verde Formation. The study of the postglacial fauna of the El Paso Formation and its relationship with the Levipustula fauna in the Calingasta-Uspallata Basin, help determine the main controls on the distribution of the postglacial faunas in other late Palaeozoic South American basins, such as the Tepuel Genoa Basin in Patagonia and the Tarija Basin in Bolivia.
Gabriela A. Cisterna [gabrielacisterna@conicet.
Water fern foliage is described from the Paleogene Redbank Plains Formation at Dinmore in southeast Queensland. The material, which is based upon leaf impressions, records early sporophyte growth stages. The specimens occur at discrete levels in clay pits at Dinmore, and the different leaf stages present suggest that they represent colonies of young submerged plants, mats of floating leaves, or a mixed assemblage of both. The leaf material closely matches the range of variation evident in young sporophytes of Ceratopteris Brongn., but in the complete absence of Cenozoic fossils of the spore genus Magnastriatites Germeraad, Hopping & Muller emend. Dettmann & Clifford from mainland Australia, which are the fossil spores of this genus, it is referred to a new genus, Tecaropteris. The record of ceratopterid-like ferns adds significantly to our limited knowledge of Cenozoic freshwater plants from Australia. The geoheritage significance of sites, such as Dinmore, is discussed briefly.
Andrew C. Rozefelds [andrew.
Extant tettigarctids are also known as hairy cicadas because they are covered by long and abundant hairs. This character had not been reported in fossil species of Tettigarctidae because previous examples were poorly preserved or lacked long hairs. Hirtaprosbole erromera gen. et sp. nov. (Tettigarctidae) with a hairy body, from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China, is described here. This new species provides evidence that tettigarctids with long dense hairs had appeared by the latest Middle Jurassic and lived at high altitudes.
Xiao-hui Liu [liuxh8917@163.
New gazelle fossils are described from the Siwalik Group of Pakistan. The material includes horncores, maxilla and mandible fragments, and isolated teeth. The available samples are assigned to three Gazella species: Gazella sp. in the Lower Siwalik Subgroup (ca 14.2–11.2 Ma), and G. lydekkeri and G. superba in the Middle Siwalik Subgroup (ca 10.2–3.4 Ma). Based on a review of the Siwalik Group gazelles, G. padriensis is synonymized with G. lydekkeri. Gazella superba Pilgrim, 1939 sensu stricto is a large form and is a valid species of the genus in the Siwalik Group.
Muhammad Akbar Khan [akbaar111@yahoo.
A new insect species, Cixius discretus (Hemiptera, Fulgoromorpha), from the Lower Miocene Garang Formation of Zeku County, Qinghai Province (northeastern Tibetan Plateau) is described. This species can be assigned to Cixiidae and represents the first fossil representative of this family from Qinghai Province. Based on the recent single-origin hypothesis and the distribution of tectonic plates in the Cretaceous, we consider that ancient Cixius had dispersed globally prior to the Cretaceous. Through analysis of the habitats of extant Cixius, the palaeoclimate and fossil flora of the Zeku area during the Miocene, we interpret the climate of Zeku in the early Miocene to have been warm-temperate and mildly arid. The new species constitutes evidence of wooded and shrubby habitats in Zeku during the Miocene.
Yi Li [liy0124@sina.
A new bandicoot species, Kutjamarcoot brevirostrum gen. et sp. nov. (Peramelemorphia), is described here from the Leaf Locality, Kutjamarpu Local Fauna (LF), Wipajiri Formation (South Australia). The age of the fossil deposit is interpreted as early Miocene on the basis of biocorrelation between multiple species in the Kutjamarpu LF and local faunas from the Riversleigh World Heritage Area (WHA). Kutjamarcoot brevirostrum is represented by isolated teeth and three partial dentaries and appears to have been short-snouted with an estimated mass of 920 g. Phylogenetic analyses place K. brevirostrum in a clade with extant Australian bandicoots and the extinct Madju, but potentially exclude the extant New Guinean bandicoots. Morphometric analysis infers close similarity between K. brevirostrum and species of Galadi in both size and rostral length. They, thus, potentially occupied compatible ecological niches with competitive exclusion perhaps explaining geographical segregation between these broadly coeval lineages.
Philippa M. Chamberlain [philippa.
The discovery of a well-preserved dragonfly forewing in the Upper Jurassic Talbragar Fish Bed near Gulgong and attributed to Austroprotolindenia jurassica Beattie & Nel allows this taxon to be placed in Protolindeniidae. It extends the palaeogeographical distribution of this family, previously known only from the Jurassic of Europe, to Australia.
André Nel [anel@mnhn.
A diminutive euphyllophyte, Douaphyton levigata gen. et sp. nov., is described from the upper Middle Devonian (Givetian) Hujiersite Formation of West Junggar, Xinjiang, China. The plant consists of more than three orders of axis branching, each axis being less than 2 mm wide. The second-order axes are short, laterally and alternately attached to the main axis. The third-order axes are paired and anisotomously divided, bearing the vegetative appendages or the fertile units. The fertile unit consists of a short recurved axis giving off up to four short pedicels along one side, each of which bears one to four pairs of terminal sporangia. Douaphyton has a three-dimensional branching system that has an intermediate form in the evolutionary context of euphyllophytes and lignophytes. It is also proposed that complex branching developed in multiple groups in the Middle Devonian.
*Hong-He Xu [hhxu@nigpas.
The smooth atrypoid brachiopod Thulatrypa gen. nov. incorporates two species, a younger (T. gregaria) from Norway, and an older (T. orientalis) from South China, which collectively span the middle Rhuddanian through Aeronian. In Baltica, the genus thrived just below the storm wave base in a tropical BA4 setting extending slightly into BA3 and BA5 respectively, whereas in South China, its representative occurs in a much shallower assemblage (BA2–3). Their palaeobiogeographical implications are carefully investigated. This study supports the arguments that Thulatrypa may have originated in South China in the middle Rhuddanian and extended its range to eastern Baltica in the late Rhuddanian. Larvae may have drifted along a channel from the east to the southwest of Baltica, which supports the reconstructions of palaeocurrents in the early Silurian in previous palaeogeographical studies.
Bing Huang [bhuang@nigpas.
Anatomical variation in the sacrum is useful for differentiating sauropod lineages. Morphological variation in the sacrum has been underestimated mainly because of its anatomical complexity and uncertain homologies with presacral and postsacral elements. We describe a titanosaurian sauropod pelvis (MLP 46-VIII-21-2) from Plottier (Neuquén Province, Argentina) collected in the 1940s from Coniacian–Santonian (Upper Cretaceous) strata. The degree of bone fusion in the specimen (fused vertebral centra forming a single rod, with neural arches fused to the corresponding centrum and to adjacent neural arches) indicates a late ontogenetic age. The presence of a space between the first vertebra and the anterior margin of the illium, together with a scar located on the anterior part of the preactabular process, suggest the possible presence of a sixth sacral vertebra (putative dorsosacral additional characters present in the ilium point towards affinities with Titanosauria. The completeness and good preservation of the specimen allowed us to track features along the sacral series and to compare characters with other sauropods. Within a phylogenetic context, and based on the pattern present in basal sauropodomorphs, the presence of three sacral elements attached to the rim of the acetabulum in eusauropods opens the possibility for considering such elements as primordial sacrals.
Florencia S. Filippini [florencia304@live.
Campterophlebiidae is the most diverse family of fossil odonatans in China with ten genera recovered mostly from Middle Jurassic strata of Inner Mongolia. We describe a well-preserved campterophlebiid damsel-dragonfly from the Middle Jurassic Yanan Formation in Shanxi Province, northwestern China. This discovery adds to the diversity of Campterophlebiidae and identifies a new Middle Jurassic insect fossil locality in China. Within Campterophlebiidae, the new taxon most closely resembles Ctenogampsophlebia from the Middle Jurassic of Inner Mongolia but differs from other genera in having vein AA with four parallel posterior branches uncrossed in the anal triangle.
Daran Zheng [dranzheng@gmail.
A new form of dendroid multicellular thallophyte is documented in the Ediacaran Doushantuo phosphorite at Weng’an, Guizhou Province, southern China. The dendroid thallophytes have variable forms, possibly owing to heteromorphic variation. Many lateral branches extend from the upper portions of the main axes; the lateral branches bear terminal vegetative vesicles, reproductive vesicles, monosporangium-like discoidal vesicles and urn-shaped pseudoparenchymatous structures. The vegetative vesicles give rise to clavate pseudoparenchymatous structures, characterized by differentiation of the thallus medulla/cortex, which might represent an early stage of thallus development. An oogamous conceptacle arising from one carpogonial vesicle forms a highly specialized goblet-shaped conceptacle. The discovery of the new dendroid multicellular thallophytes provides not only the first fossil-based evidence of the morphological complexity and tissue differentiation in the Precambrian organisms but also insights into the life cycle of the Precambrian red algae.
Wei Du* [duwei@ea.
The problematic Small Shelly Fossil Cambroclavus absonus is described from the Xinji Formation in the Longxian area, which is located near the southwestern margin of the North China Platform. The Xinji Formation, the basal rock unit of the Cambrian in the studied area, yields an assemblage of skeletal fossils that share many common elements with contemporary faunas from South Australia. Sclerites of C. absonus reported herein represent the first occurrence of the species outside Australia, thus extending the palaeogeographic range of the taxon to northern China. To date, palaeogeographic occurrences of Cambroclavus sclerites are restricted to the Peri-Gondwana realm, including South China, Australia, Tarim, Kazakhstan, North China and Western Europe. These occurrences are divided into a Southern Group realm and Northern Group realm. Stratigraphically, Cambroclavus occurs mostly in Cambrian Stage 3 and has three occurrences in Stage 5, separated by Stage 4 in which Cambroclavus has not yet been found. The first appearance datum of Cambroclavus in Cambrian Stage 3 is of importance for regional and inter-regional correlations. In particular, the presence of Cambroclavus absonus in North China allows species-level correlation between North China and South Australia.
Luoyang Li [lly@stumail.
Six conodont and one fusuline zones are recognized on basis of a total of 25 conodont and 13 fusuline species (including seven unidentified species or species given with cf. or aff. in total) from the Bamchi Formation, Yeongwol, Korea. The conodont zones include the Streptognathodus bellus, S. isolatus, S. cristellaris, S. sigmoidalis, S. fusus and S. barskovi zones in ascending order, which can be correlated with the conodont zones spanning the uppermost Gzhelian to Asselian Age of the Permian globally. The fusuline zone is named the Rugosofusulina complicata–Pseudoschwagerina paraborealis zone. The co-occurrence of the conodont Streptognathodus isolatus (the Global Boundary Stratotype Section and Point index for the base of Permian) and Pseudoschwagerina (a Permian inflated fusuline) indicates that the Carboniferous–Permian boundary can be placed in the lower part of the Bamchi Formation in South Korea.
Qiulai Wang* [qlwang@nigpas.
The ‘tuning-fork’ didymograptid previously referred to as Didymograpt(ell)us protobifidus is common in Victoria where it is confined to the Chewtonian (mid-Floian). Biometric differences indicate that the mid-Floian form is not conspecific with the holotype of the Darriwilian Didymograptus protobifidus Elles, 1933 and the Floian form is thus renamed Didymograptellus kremastus n. sp. Study of the Valhallfonna Formation faunas on Spitsbergen indicated that the Floian form of D. ‘protobifidus’ differs from Didymograptellus bifidus (Hall) in both its morphology and stratigraphic distribution but a later study of the Cow Head Group on Newfoundland concluded that they are one species. My study, of more than 50 specimens of Didymograptellus from the Floian of Victoria, Australia, shows that the two are different and that similar differences exist in the Cow Head Group populations of Didymograptellus. The Chewtonian (Ch1) Didymograptellus protobifidus Biozone is renamed D. kremastus Biozone.
Alfons H.M. Vandenberg, [lanceolatus@hotmail.
A new grylloblattid (Permoshurabia argentina sp. nov.: Geinitziidae) is described and illustrated from the Upper Triassic of Argentina. The material represents the first record of this family from Argentina and expands the geographic distribution of this group during the Triassic.
María Belén Lara [lara.