Late Pleistocene protein residues from the Mahaffy cache,Colorado

This paper reports the analysis of protein residues from tools recovered in a cache within the city limits of Boulder, Colorado, USA. This cache included a total of 83 artifacts, all of which we subjected to cross-over immunoelectrophoresis (CIEP). Four of the 83 produced results, with residues from each of these reacting with antigens from a different taxon: one tool shows evidence of use on sheep, one on bear, one on horse, and one on camel. Varieties of sheep and bear have been present in Colorado throughout human history, but horses and camelids have been in the state either during the Pleistocene or the last 200 years. Several lines of evidence indicate that the cache cannot be recent, and our CIEP results therefore imply that the cache date to the late Pleistocene. Typological aspects of the artifacts in the cache are consistent with artifacts known to be Clovis, and the combination of CIEP and typological data thus indicate that the cache is Clovis as well. These data contribute to an increasing dataset documenting the broad range of animals other than elephants hunted by Clovis groups in North America.     

The Enisei River of Central Siberia in the Late Pleistocene

Recent Paleolithic work along the middle Enisei River of central Siberia has revealed a long history of occupation that almost certainly begins in the Middle Pleistocene. Although the evidence for the Lower Paleolithic is somewhat tentative, there is good reason to believe that hunter-gatherers had periodically occupied the middle Enisei before the last interglacial. The steppe environment of the region during the Upper Pleistocene was relatively bountiful; more than 200 Upper Paleolithic sites, both before and after the Last Glacial Maximum, have been located. The region appears to have been abandoned during the Last Glacial Maximum. Most of Soviet and Russian archaeological work has been guided by a cultural–historical orientation, but recently there has been increased interest in developing adaptationist and ecological research strategies. The middle Enisei and the wider central Siberian region are key to understanding early adaptations to the north and the dimensions of Paleolithic population movements.     

Mortality analysis of the Late Pleistocene bears from Grotta Lattaia,central Italy

Reported here are the results of a mortality analysis of an Ursus spelaeus death assemblage from Grotta Lattaia, southern Tuscany (central Italy), excavated in 1939. The Grotta Lattaia cave bears are among the latest representatives of the species. The large amount of cave bear remains indicates that the cavern was a lair for hibernating bears which repeatedly, but not necessarily yearly, occupied it. The mortality evidence indicates that deaths occurred primarily during hibernation from violent predation, and, therefore, that the bears had direct interaction with other carnivores. However, no sign of interrelation with humans could be found, even though Middle Paleolithic human remains and tools had been recovered associated with the bear material. Grotta Lattaia can be thus considered an example of cumulative, non-human-caused violent deaths in a hibernation context.     

Late Pleistocene mammalian extinctions in North America: Taxonomy,chronology, and explanations

Toward the end of the Pleistocene, North America lost some 35 genera of mammals. It has long been assumed that all or virtually all of the extinctions occurred between 12,000 and 10,000 years ago, but detailed analyses of the radiocarbon chronology provide little support for this assumption, which seems to have been widely accepted because of the kinds of explanations felt most likely to account for the extinctions in the first place. Approaches that attribute the losses to human predation depend almost entirely on the assumed synchroneity between the extinctions and the onset of large mammal hunting by North American peoples. The fact that only two of the extinct genera have been found in a convincing kill context presents an overwhelming problem for this approach. Climatic models, on the other hand, are becoming increasingly precise and account for a wide variety of apparently synchronous biogeographic events. While a role for human activities in the extinction of some taxa is fully possible, there can be little doubt that the underlying cause of the extinctions lies in massive climatic change.     

Cut marks on the Middle Pleistocene elephant carcass of Áridos 2 (Madrid,Spain)

Áridos 1 and Áridos 2 (Madrid, Spain) are two Middle Pleistocene sites belonging to the isotopic stages 9–11. Both places contain partial carcasses of Elephas (Paleoxodon) antiquus associated to Acheulian stone tools. In this work, the taphonomic study of the elephant remains of Áridos 2 is presented. This study has documented several cut marks on different bones, which indicate bulk flesh and viscerae extraction by Middle Pleistocene hominins. Several arguments are provided to support that at least some of the cut marks were made with handaxes, further suggesting that some of these artifacts were butchering tools in this stage of human evolution. Although cut marks on elephant carcasses have been documented at some Middle Pleistocene sites, very few have been published in detail to allow consideration of their status as hominin-imparted marks. By doing so, the present study provides more evidence of large carcass exploitation by hominins during this period.     

Late Pleistocene/Early Holocene seafaring in the Aegean: new obsidian hydration dates with the SIMS-SS method

Archaeological evidence regarding the presence of obsidian in levels that antedate the food production stage could have been the result of usage or intrusion of small obsidian artifacts from overlying Neolithic layers. The new obsidian hydration dates presented below employing the novel SIMS-SS method, offers new results of absolute dating concordant with the excavation data. Our contribution sheds new light on the Late Pleistocene/Early Holocene exploitation of obsidian sources on the island of Melos in the Cyclades reporting dates c. 13th millennium - end of 10th millennium B.P.     

The MSA sequence of Diepkloof and the history of southern African Late Pleistocene populations

This paper presents the first multidisciplinary synthesis of the Middle Stone Age sequence of Diepkloof Rock Shelter (South Africa). We explore the main cultural changes that characterized southern African hunter–gatherer societies from OIS 5 to the beginning of OIS 3. We discuss the tempo of these changes, test the current interpretative hypotheses and explore an empirical model to explain the early appearance of symbolic marking within the Pleistocene hunter–gather societies of southern Africa.     

First fossil fish (Teleostei,Siluriformes) from the Late Pleistocene of Santa Fe Province,Argentina

Vallone, E.R., Vezzosi, R.I. & Cione, A.L. February 2017. First fossil fish (Teleostei, Siluriformes) from the Late Pleistocene of Santa Fe Province, Argentina. Alcheringa 00, 000–000. ISSN 0311-5518.

The record of fossil fish from the Pleistocene of Argentina is poor. Here we describe the first ichthyofauna from Late Pleistocene riverbank beds in the Salado River of Santa Fe Province, Argentina. The material consists of isolated pectoral and dorsal fin spines, together with skull fragments. Four species-level taxa referable to three families can be identified: Pterodoras granulosus (Doradidae), Pimelodus cf. maculatus and Pimelodus cf. albicans (Pimelodidae) and cf. Hypostomus sp. (Loricariidae). Specimens attributed to Pterodoras granulosus and Pimelodus maculatus represent a minimum age for origin of these taxa. The Salado River assemblage includes the richest record of Pleistocene catfishes yet documented from southern South America.

Evelyn Romina Vallone [evelynvallone@conicet.gov.ar] and Raúl Ignacio Vezzosi [vezzosiraul@gmail.com], Laboratorio de Paleontología de Vertebrados, CICYTTP-CONICET, Materi y España, (3105) Diamante, Entre Ríos, Argentina; Alberto Luis Cione [acione@museo.fcnym.unlp.edu.ar], División Paleontología de Vertebrados. Museo de La Plata. Paseo del Bosque s/n, (1900) La Plata, Argentina.     

Standing at the gates of Europe: Human behavior and biogeography in the Southern Carpathians during the Late Pleistocene

This study presents a behavioral analysis of Middle and Upper Paleolithic lithic assemblages from 14 sites located in the southern Carpathian Mountains. Using a whole assemblage behavioral indicator, we show that the hominins that manufactured those stone tools do not appear to have differed in terms of the flexibility of the mobility strategies they employed to exploit their landscapes. Rather than biological change, we argue that large-scale climate changes are likely more important drivers of behavioral changes during the Late Pleistocene of the region, including during the Middle–Upper Paleolithic transition. These results agree well with the results of studies having employed this methodology in other regions, suggesting that this is a generalized feature of the transition across Eurasia. Recasting the transition as a mainly ecological rather than purely biocultural process allows us to generate new perspectives from which to approach the question of behavioral change during the Late Pleistocene, and ultimately suggests that the process referred to as the ‘Middle–Upper Paleolithic transition’ is essentially a brief segment of a much more extensive process driven by prehistoric human–environment interactions that would culminate in the highly logistical mobility strategies documented throughout the continent at the Last Glacial Maximum.     

Middle Pleistocene bird consumption at Level XI of Bolomor Cave (Valencia, Spain)

The consumption of small prey dates back to the Plio-Pleistocene chronologies in some African sites. However, the systematic acquisition and consumption of small prey in the pre-Upper Palaeolithic times is still a highly debated topic in Europe. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of bird consumption is not as common for these periods. Nevertheless, Level XI (MIS 6) of Bolomor Cave has clear diagnostic elements to document the acquisition and use of birds (Aythya sp.) for food in the form of: (1) cutmarks on bones of both the front and hind limb; (2) presence of burning patterns on the extremities of the bones (areas of the skeleton with less meat); and (3) human toothmarks on limb bones. The capture of birds is classified as quick-flying game in the archaeological sites. The acquiring of fast-running (mostly lagomorphs) and quick-flying small prey requires a sophisticated technology and involves obtaining and processing ways different from those used for large- and medium-sized animals. From this perspective, the aim of this paper is to examine possible patterns in the processing sequence of birds from Level XI of Bolomor Cave and to improve the data on their butchery and human consumption in the Middle Pleistocene of Iberian Peninsula.     

On Microlithic Quartz Industries at the End of the Pleistocene in Central Africa: The Evidence from Shum Laka (NW Cameroon)

The rock shelter of Shum Laka, situated in the Grassfields of northwestern Cameroon, was occupied during the later Pleistocene and Holocene. Plant remains reveal that there were no drastic changes in the immediate environment of the shelter between 30,000 and 10,000 bp. The industry is microlithic during this entire period. Quartz was deliberately chosen as the preferred raw material and there were no changes in technology or raw material usage over time. A comparison with similar sites in the northeastern Democratic Republic of Congo reveals that the microlithic quartz tradition was apparently appropriate for the exploitation of a wide variety of environmental settings. This flexibility may represent an adequate technological response to the environmental changes in Central Africa at the end of the Pleistocene.L'abri sous roche de Shum Laka, situé dans les Grassfields au nord-ouest du Cameroun, a été occupé depuis la fin du Pléistocène. Les restes de plantes montrent qu'il n'y a pas eu de changements importants dans l'environnement immédiat de l'abri entre 30,000 et 10,000 bp. L'industrie est microlithique pendant toute cette période. Le choix du quartz comme matière première préferentielle était délibéré par ailleurs, ni la technologie, ni l'utilisation des matières premières n'ont subi de modifications conséquentes au cours de cette période. Une comparaison avec des sites similaires du nord-est de la République Démocratique du Congo nous montre que la tradition microlithique sur quartz aurait été appropriée dans l'exploitation d'une grande variété d'écosystèmes. Cette flexibilité pourrait traduire une réponse technologique adéquate face aux changements environnementaux qui ont eu lieu en Afrique Centrale à la fin du Pléistocène.     

Patterning and Its Causation in the Pre-Neolithic Colonization of the Mediterranean Islands (Late Pleistocene to Early Holocene)

In 1981 one of us (Cherry) first attempted to identify spatial and temporal patterning in the human colonization of the Mediterranean islands. Since the 1980s, slowly accumulating evidence has suggested that the Mediterranean islands were sporadically inhabited by hunter-gatherers during the Late Pleistocene and Early Holocene. Here we seek to establish whether or not these data exhibit regularity. We suggest that evidence for Upper Palaeolithic to Mesolithic activity, tending to cluster on larger or less remote islands, indicates that while humans were clearly capable of reaching the Mediterranean islands prior to the Neolithic, their general reluctance to do so can be explained in terms of the variable environmental attractiveness of the insular Mediterranean. Tending to be relatively small, dry, and biologically depauperate, the Mediterranean islands were largely inhospitable to mobile groups preferring extensive territories with diverse and robust biotas. Sedentism only became a widely viable strategy in the insular Mediterranean with the development in the Neolithic of what we might regard as “terraforming”—that is, the introduction of cereals, pulses, and ovicaprids, all tolerant of xeric environments.     

Late Pleistocene Crocuta crocuta spelaea (Goldfuss, 1823) clans as prezewalski horse hunters and woolly rhinoceros scavengers at the open air commuting den and contemporary Neanderthal camp site Westeregeln (central Germany)

Late Pleistocene Ice Age Crocuta crocuta spelaea ( Goldfuss, 1823) hyenas from the open-air gypsum karst site Westeregeln (Saxony-Anhalt, central Germany) is dated into the early to middle Late Pleistocene. Hyena clans apparently used the karst for food storage and as “commuting den”, where typical high amounts (15% of the NISP) of hyena remains appear, also faecal pellets in concentrations for den marking purposes. Additionally small carnivores Meles, Vulpes and Mustela appear to have used some cavities as dens. Several hundreds of lowland “mammoth steppe fauna” bones (NISP = 572) must have been accumulated primarily by hyenas, and not by Neanderthals at the contemporary hyena/human camp site. Abundant caballoid horse remains of “E. germanicus Nehring, (1884)” are revised by the holotype and original material to the small E. c. przewalskii horse. Woolly rhinoceros Coelodonta antiquitatis remains are also abundant, and were left in several cases with typical hyena scavenging damages. Rangifer tarandus (11%) is mainly represented by numerous fragments of shed female antlers that were apparently gathered by humans, and antler bases from male animals that were collected and chewed in few cases (only large male antlers) by hyenas. The large quantities of small reindeer antlers must have been the result of collection by humans; their stratigraphic context is unclear but such large quantities most probably resulted from schamanic activities. The hyena site overlaps with a Middle Palaeolithic Neanderthal camp, as well as possibly with a later human Magdalénian site.     

Interpreting pachyderm single carcass sites in the African Lower and Early Middle Pleistocene record: A multidisciplinary approach to the site of Nadung’a 4 (Kenya)

Nadung’a 4 is one of the single carcass pachyderm sites recorded in East Africa during the Lower and Early Middle Pleistocene. The site has yielded an abundant lithic assemblage in close association with the partial carcass of an elephant. Conjoined pedological, geoarchaeological, spatial, technological, and taphonomical analyses have been carried out to address the relationship between hominids and elephant. The resulting data are consistent with a non-fortuitous association between both categories of remains. The lithic artefacts do not match a classical Acheulean tool-kit, as would be expected for the time period ascribed to the site, and the functional patterns inferred from their analysis make this site radically different from other purported butchery sites. The implications of these original features are discussed.     

Single carcass of Mammuthus primigenius with lithic artifacts in the Upper Pleistocene of northern Italy

This paper presents the find of a Mammuthus primigenius carcass and associated Mousterian implements from the Last Glacial site of Asolo, in north-eastern Italy. We review the exploitation of proboscidean carcasses at Lower and Middle Paleolithic sites of Africa, Europe and the Levant, including evidence of elephant killing, and summarize recent research on hafting and use of Mousterian points as spearheads already before the end of the Middle Pleistocene. The bones and implements from Asolo are described in detail; we provide information on other Italian sites with mammoth remains and on the Mousterian record of the north-eastern quadrant of the peninsula. A possible impact fracture has been detected on a Levallois point from Asolo; its interpretation is based on comparisons to similar scars found on spear points of verified function from archaeological sites of later age, and on experimental material. We conclude that the evidence of Asolo is consistent with information from a number of Western European sites supporting a picture of Neanderthals as capable hunters of large game, such as woolly mammoth.     

Isotopic analysis of the ecology of herbivores and carnivores from the Middle Pleistocene deposits of the Sierra De Atapuerca,northern Spain

Carbon and oxygen isotope values reveal resource partitioning among the large mammal fauna from three contemporaneous Middle Pleistocene hominid-bearing localities within the Sierra de Atapuerca (northern Spain). Carbon isotope values sampled from the tooth enamel of fauna present during Atapuerca Faunal Unit 6 show that a C₃-dominated ecosystem surrounded the area where fossils were preserved during this time. For the herbivores, Fallow deer isotope values are significantly different from Red deer and horses and show that this species did not forage in open environments at this locality. Red deer and horses show similar feeding strategies with less negative carbon values implying use of more open environments for these taxa. For the carnivores, carbon isotope values for Ursus deningeri are significantly different from either lions (Panthera leo) or foxes (Vulpes vulpes) and support the contention that this species is herbivorous. Special metabolic mechanisms involved in hibernation in U. deningeri might also have influenced its isotope values. The carbon isotope values of remaining carnivores were similar and suggest that each was typically a generalist carnivore, eating a wide variety of prey items. While the isotopic results generally correspond to ecology indicated by previous techniques, this study shows that isotope analyses can provide further insights that alternate techniques do not provide. Isotope analyses can help elucidate the ancient ecology of taxa present in the Sierra de Atapuerca during the Middle Pleistocene allowing for an accurate portrayal of the setting in which humans lived.     

Palaeopathology in late Pleistocene and early Holocene Central Plains bison: dental enamel hypoplasia,fluoride toxicosis and the archaeological record

Multiple formative agents are cited for dental enamel hypoplasia (DEH) in modern and fossil mammals. Here, four late Pleistocene-early Holocene bison assemblages from the Central Great Plains are analyzed for DEH and other bone pathologies to investigate the possible influence of chronic fluoride toxicosis, a condition that may have presented a long-term source of duress for prehistoric bison and devalued them as nutritional resources to human hunters. Although DEH is present in all the analyzed assemblages, the lack of corroborating dental and post-cranial anomalies consistent with chronic fluoride toxicosis speaks against it as a primary factor of DEH formation or general ill health in these bison. Rather, maternal nutritional stress, weaning, and seasonal forage condition are invoked as probable promoters of DEH formation. Other identified abnormalities are discussed in reference to their contribution to reduced feeding efficiency and lameness, as are their roles in developing palaeoecological models applicable to archaeological questions about ancient human lifeways.     

Archaeological survey,paleogeography, and the search for Late Pleistocene Paleocoastal peoples of Santa Rosa Island,California

Abstract

The northern Pacific Coast is an important area for understanding human colonization of the Americas, but Late Pleistocene coastal sites are rare and interglacial sea level rise has inundated the continental shelf and the primary areas where Paleocoastal archaeological sites are likely to occur. Here we outline a terrestrial archaeological survey project designed to identify Paleocoastal sites on Santa Rosa Island, California. Using reconstructions of ancient shorelines and paleogeography, we predicted that sites might be found where lithic resources, freshwater springs, caves or rockshelters, and strategic vistas drew Paleocoastal peoples into the island interior. We identified nine new Paleocoastal sites, including four radiocarbon dated to >11,000 cal b.p. that are among the oldest sites on North America’s Pacific coast. Our targeted survey demonstrates an important technique for investigating island and coastal settings where sea level rise remains a significant challenge for locating early sites.