A new fossil species, Luanpingia daohugouensis sp. nov., belonging to the family Sinoalidae is described from the Middle to Upper Jurassic Daohugou beds of Inner Mongolia, China, on the basis of two well-preserved complete specimens. The described species of Sinoalidae are reviewed and Jiania gracila is considered a junior synonym of Jiania crebra. The new discovery increases the palaeodiversity of sinoalids from the Daohugou beds. It also indicates stratigraphic correlation between the Daohugou beds, the Haifanggou Formation at Haifeng, Beipiao City, West Liaoning Province, and the Jiulongshan Formation at Zhouyingzi, Luanping County, Hebei Province. All of these units host the ‘early assemblage’ of the Yanliao biota.
Yanzhe Fu [yzfu41@163.
Extant tettigarctids are also known as hairy cicadas because they are covered by long and abundant hairs. This character had not been reported in fossil species of Tettigarctidae because previous examples were poorly preserved or lacked long hairs. Hirtaprosbole erromera gen. et sp. nov. (Tettigarctidae) with a hairy body, from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China, is described here. This new species provides evidence that tettigarctids with long dense hairs had appeared by the latest Middle Jurassic and lived at high altitudes.
Xiao-hui Liu [liuxh8917@163.
Campterophlebiidae is the most diverse family of fossil odonatans in China with ten genera recovered mostly from Middle Jurassic strata of Inner Mongolia. We describe a well-preserved campterophlebiid damsel-dragonfly from the Middle Jurassic Yanan Formation in Shanxi Province, northwestern China. This discovery adds to the diversity of Campterophlebiidae and identifies a new Middle Jurassic insect fossil locality in China. Within Campterophlebiidae, the new taxon most closely resembles Ctenogampsophlebia from the Middle Jurassic of Inner Mongolia but differs from other genera in having vein AA with four parallel posterior branches uncrossed in the anal triangle.
Daran Zheng [dranzheng@gmail.
A diminutive euphyllophyte, Douaphyton levigata gen. et sp. nov., is described from the upper Middle Devonian (Givetian) Hujiersite Formation of West Junggar, Xinjiang, China. The plant consists of more than three orders of axis branching, each axis being less than 2 mm wide. The second-order axes are short, laterally and alternately attached to the main axis. The third-order axes are paired and anisotomously divided, bearing the vegetative appendages or the fertile units. The fertile unit consists of a short recurved axis giving off up to four short pedicels along one side, each of which bears one to four pairs of terminal sporangia. Douaphyton has a three-dimensional branching system that has an intermediate form in the evolutionary context of euphyllophytes and lignophytes. It is also proposed that complex branching developed in multiple groups in the Middle Devonian.
*Hong-He Xu [hhxu@nigpas.
The smooth atrypoid brachiopod Thulatrypa gen. nov. incorporates two species, a younger (T. gregaria) from Norway, and an older (T. orientalis) from South China, which collectively span the middle Rhuddanian through Aeronian. In Baltica, the genus thrived just below the storm wave base in a tropical BA4 setting extending slightly into BA3 and BA5 respectively, whereas in South China, its representative occurs in a much shallower assemblage (BA2–3). Their palaeobiogeographical implications are carefully investigated. This study supports the arguments that Thulatrypa may have originated in South China in the middle Rhuddanian and extended its range to eastern Baltica in the late Rhuddanian. Larvae may have drifted along a channel from the east to the southwest of Baltica, which supports the reconstructions of palaeocurrents in the early Silurian in previous palaeogeographical studies.
Bing Huang [bhuang@nigpas.
New gazelle fossils are described from the Siwalik Group of Pakistan. The material includes horncores, maxilla and mandible fragments, and isolated teeth. The available samples are assigned to three Gazella species: Gazella sp. in the Lower Siwalik Subgroup (ca 14.2–11.2 Ma), and G. lydekkeri and G. superba in the Middle Siwalik Subgroup (ca 10.2–3.4 Ma). Based on a review of the Siwalik Group gazelles, G. padriensis is synonymized with G. lydekkeri. Gazella superba Pilgrim, 1939 sensu stricto is a large form and is a valid species of the genus in the Siwalik Group.
Muhammad Akbar Khan [akbaar111@yahoo.
Previously, Progiraffa exigua has been reported only from the Kamlial Formation (ca 18.3–14.2 Ma) of the Siwalik Group. We record Progiraffa exigua from the Lower Siwalik Subgroup at five localities: Jaba, Chinji Rest House, Rakh Wasnal, Dhok Bun Amir Khatoon and Ghungrila, Pakistan, thus extending the range of P. exigua to the Chinji Formation of the Siwalik Group (ca 14.2–11.2 Ma).
Kiran Aftab [kiranaftab2012@gmail.
The discovery of a well-preserved dragonfly forewing in the Upper Jurassic Talbragar Fish Bed near Gulgong and attributed to Austroprotolindenia jurassica Beattie & Nel allows this taxon to be placed in Protolindeniidae. It extends the palaeogeographical distribution of this family, previously known only from the Jurassic of Europe, to Australia.
André Nel [anel@mnhn.
The oldest Paleogene strata on Chatham Islands, east of New Zealand, are the phosphatized conglomerates and sandstones of the Takatika Grit that crops out on the northeastern coast at Tioriori and unconformably overlies the Chatham Schist. An intact Cretaceous–Paleogene boundary transition is not preserved at this locality. New biostratigraphic analysis of dinoflagellate, diatom and radiolarian microfossil assemblages confirms that the Takatika Grit is of late early–middle Paleocene (New Zealand Teurian stage) age but contains reworked microfossils of early Campanian (Early Haumurian) age. Vertebrate fossils found in this unit are inferred to be a mixture of reworked Cretaceous and in situ Paleocene bones and teeth. The overlying Tutuiri Greensand is of middle–late Paleocene age in its lower part and also contains reworked Cretaceous microfossils.
Christopher J. Hollis [c.
Thanerocleridae is a small family of Cleroidea with no fossil representatives to date. Here we describe and figure the first fossil representative of Thanerocleridae, Cretozenodosus fossilis gen. et sp. nov., from the mid-Cretaceous amber of northern Myanmar. Cretozenodosus is referred to the extant subfamily Zenodosinae as evidenced by its open procoxal cavities and transverse procoxae. Cretozenodosus has close affinities with the North American Zenodosus Wolcott, suggesting that modern Zenodosinae is probably a relict group. Our discovery of a new thaneroclerid genus from Burmese amber suggests that Thanerocleridae originated no later than the mid-Cretaceous.
Chenyang Cai [cycai@nigpas.
A new dasyuromorphian, Barinya kutjamarpensis sp. nov., is described on the basis of a partial dentary recovered from the Miocene Wipajiri Formation of northern South Australia. Although about the same size as the only other species of this genus, B. wangala from the Miocene faunal assemblages of the Riversleigh World Heritage Area, northwestern Queensland, it has significant differences in morphology including a very reduced talonid on M4 and proportionately wider molars. Based on the structural differences and the more extensive wear on its teeth, the central Australian species might have consumed harder or more abrasive prey in a more silt-rich environment than its congener, which hunted in the wet early to middle Miocene forests of Riversleigh.
Pippa Binfield [pippa.
Six conodont and one fusuline zones are recognized on basis of a total of 25 conodont and 13 fusuline species (including seven unidentified species or species given with cf. or aff. in total) from the Bamchi Formation, Yeongwol, Korea. The conodont zones include the Streptognathodus bellus, S. isolatus, S. cristellaris, S. sigmoidalis, S. fusus and S. barskovi zones in ascending order, which can be correlated with the conodont zones spanning the uppermost Gzhelian to Asselian Age of the Permian globally. The fusuline zone is named the Rugosofusulina complicata–Pseudoschwagerina paraborealis zone. The co-occurrence of the conodont Streptognathodus isolatus (the Global Boundary Stratotype Section and Point index for the base of Permian) and Pseudoschwagerina (a Permian inflated fusuline) indicates that the Carboniferous–Permian boundary can be placed in the lower part of the Bamchi Formation in South Korea.
Qiulai Wang* [qlwang@nigpas.
Omma Newman is an extant ommatid genus currently endemic to Australia. A new Omma species, O. daxishanense sp. nov. is described and illustrated based on a compression fossil from the Upper Jurassic Tiaojishan Formation at Daxishan, a fossil locality well known for yielding mammals, feathered dinosaurs and diverse pterosaurs. Omma daxishanense is very similar morphologically to the extant O. sagitta, but differs from the latter by its broader body and prominent temples. The new discovery documents the first valid Omma species from the Mesozoic of China and highlights the antiquity and palaeodiversity of the extant Australian endemic genus.
Chenyang Cai [cycai@nigpas.
The problematic Small Shelly Fossil Cambroclavus absonus is described from the Xinji Formation in the Longxian area, which is located near the southwestern margin of the North China Platform. The Xinji Formation, the basal rock unit of the Cambrian in the studied area, yields an assemblage of skeletal fossils that share many common elements with contemporary faunas from South Australia. Sclerites of C. absonus reported herein represent the first occurrence of the species outside Australia, thus extending the palaeogeographic range of the taxon to northern China. To date, palaeogeographic occurrences of Cambroclavus sclerites are restricted to the Peri-Gondwana realm, including South China, Australia, Tarim, Kazakhstan, North China and Western Europe. These occurrences are divided into a Southern Group realm and Northern Group realm. Stratigraphically, Cambroclavus occurs mostly in Cambrian Stage 3 and has three occurrences in Stage 5, separated by Stage 4 in which Cambroclavus has not yet been found. The first appearance datum of Cambroclavus in Cambrian Stage 3 is of importance for regional and inter-regional correlations. In particular, the presence of Cambroclavus absonus in North China allows species-level correlation between North China and South Australia.
Luoyang Li [lly@stumail.
Numerous small specimens of hypercalcified sponges of the genus Plectroninia (Jurassic to Recent) are recorded from deep water in the Atlantic, Indian and Pacific oceans, where they are attached to diverse hard substrata, mostly scleractinian skeletons. Being represented as skeletons of linked calcareous tetractines with an incomplete free spicule complement, the specimens could not be identified at the species level. These observations show that Plectroninia spp. have a wide distribution in the bathyal zone of the Recent World Ocean, where they may be the most common calcareous sponges.
Jean Vacelet* [jean.
Palynological analysis was carried out on Middle to Upper Triassic strata from Tulong, Nyalam County, southern Xizang (Tibet), China. Well-preserved miospore (pollen and spore) assemblages and sparse acritarch occurrences were identified. We recognized four formal and one informal biozones based on stratigraphically important taxa and compositional changes through the succession, in ascending order: the Triplexisporites Interval Zone (Anisian), the Staurosaccites quadrifidus Taxon-range Zone (upper Anisian to lower Norian), the Striatella Interval Zone (lower Norian), the Craterisporites rotundus Taxon-range Zone (middle to upper Norian) and the informal ‘Dictyophyllidites harrisii zone’ (Rhaetian). The zonation was supported by marine fossils (e.g., ammonoids and conodonts), and compositional similarity between the zones was examined using non-metric multidimensional scaling (NMDS). Correlation with other representative palynological sequences across Gondwana was also conducted. The presence of miospore taxa not previously recovered from the Late Triassic North and South China palynofloral provinces (e.g., Ashmoripollis reducta, Craterisporites rotundus, Enzonalasporites vigens, Minutosaccus crenulatus, Samaropollenites speciosus and Staurosaccites quadrifidus) calls for a new province in southwestern China, i.e., the Southern Xizang Province. It is proposed here that the modern expression of the northern boundary runs along the Yarlung Zangbo Suture, the remnant of the Tethys that separated the Indian Plate (southern Xizang) and the Lhasa Block during the Late Triassic. This new palynofloral province comprises typical elements of the Onslow Microflora, indicating the need for an extension of this microflora in southern Xizang, China.
Jungang Peng [jgpeng@nigpas.
An isolated turtle xiphiplastron similar to that of Puppigerus sp. is described from the upper Oligocene (27.3–25.2 Ma) Pomahaka Formation near Tapanui, Otago, New Zealand. The bone is unlike any previously described turtle from the Cenozoic of New Zealand and is from a newly recognized estuarine vertebrate locality. It represents the first Oligocene cheloniid turtle bone described from the southwest Pacific.
Henry J. L. Gard [henrygard@hotmail.
A new insect species, Cixius discretus (Hemiptera, Fulgoromorpha), from the Lower Miocene Garang Formation of Zeku County, Qinghai Province (northeastern Tibetan Plateau) is described. This species can be assigned to Cixiidae and represents the first fossil representative of this family from Qinghai Province. Based on the recent single-origin hypothesis and the distribution of tectonic plates in the Cretaceous, we consider that ancient Cixius had dispersed globally prior to the Cretaceous. Through analysis of the habitats of extant Cixius, the palaeoclimate and fossil flora of the Zeku area during the Miocene, we interpret the climate of Zeku in the early Miocene to have been warm-temperate and mildly arid. The new species constitutes evidence of wooded and shrubby habitats in Zeku during the Miocene.
Yi Li [liy0124@sina.
Two new species of Itaphlebia Sukatsheva, 1985, Itaphlebia longiovata and I. amoena (Nannochoristidae Tillyard, 1917), are described and illustrated from the Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. Previously described Middle Jurassic nannochoristid genera, Chrysopanorpa Ren in Ren et al., 1995 and Protochoristella Sun, Ren & Shih, 2007b, together with Stylopanorpodes and Netropanorpodes Sun, Ren & Shih, 2007a (originally assigned to Mesopanorpodidae) are revised and considered to be synonyms of Itaphlebia. The following tentative species synonymies are proposed: Protochoristella formosa and Stylopanorpodes eurypterus = Itaphlebia ruderalis (Ren in Ren et al., 1995), comb. nov.; Netropanorpodes sentosus = I. jeniseica Novokshonov, 1997a, syn. nov.; and Protochoristella polyneura = I. multa Novokshonov, 1997a, syn. nov. Netropanorpodes decorosus is transferred to Itaphlebia. These new species, new material and the new combinations broaden the diversity of the Itaphlebia in mid-Mesozoic ecosystems and provide new characters enabling amendment of the generic diagnosis.
YiZi Cao [easycaoyinzi@aliyun.