A new trilobite species from the upper Rhuddanian (lower Silurian) of South China,and its biogeographical implications

Anacaenaspis yanpingensis sp. nov., from the lower Niuchang Formation (upper Rhuddanian, lower Llandovery, lower Silurian), is the first record of this genus from South China. The biogeographical distribution of Anacaenaspis and some other trilobite genera from South China (e.g., Gaotania, Hyrokybe, Aulacopleura and Raphiophorus) in the Llandovery evidences faunal exchanges between Avalonia–Baltica, Laurentia, Australia and South China. We attribute these dispersals to prevailing ocean currents, and especially equatorial countercurrents, which would have propagated dispersals during the Rhuddanian, shortly after the end-Ordovician mass extinction.     

The radiation of early Silurian spiriferide brachiopods,with new taxa from the Llandovery of Iran

Popov, L.E. & Cocks, L.R.M., 2013. The radiation of early Silurian spiriferide brachiopods, with new taxa from the Llandovery of Iran. Alcheringa 38, 560–566. ISSN 0311–5518.

Although there were Late Ordovician spiriferides in the superfamily Cyrtioidea, namely Eospirifer and Odakella, only the former genus survived the terminal Ordovician extinction, and only Eospirifer is known from the earliest Llandovery (Rhuddanian). However, in the succeeding mid-Llandovery (Aeronian), the spiriferides radiated to include not only more species within Eospirifer and Striispirifer in the Eospiriferidae, but also the new genus Iranospirifer described here, which is the earliest representative of the other family within the superfamily, the Hedeinopsidae. The Ordovician species were confined to the South China continent and the Boshchekul volcanic island arc in Kazakhstan, but by the Aeronian the superfamily had spread westwards to various other continents, including the main Gondwanan superterrane (which included Iran) in the early Aeronian, and Avalonia-Baltica and Laurentia in the late Aeronian. The new species Eospirifer ghobadiae and Iranospirifer qarabilensis are both described from the lower Aeronian of Iran, and there is a note on the Aeronian rhynchonellide Stegocornu, which is endemic to Iran and nearby areas.

Leonid E. Popov [lepbarry@yahoo.co.uk; leonid.popov@museumwales.ac.uk] Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, UK. L. Robin M. Cocks [r.cocks@nhm.ac.uk], Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK. Received 29.5.2013, revised 12.7.2013, accepted 21.7.2013.     

First appearance datum of the Silurian graptolite Oktavites spiralis,and its evolution on the northern margin of South China

Wang, X., Wang, J. & Zhang, J., May 2016. First appearance datum of the Silurian graptolite Oktavites spiralis, and its evolution on the northern margin of South China. Alcheringa 41, xxx–xxx. ISSN 0311-5518.

Xin Wang [wx200315046@163.com], Early Life Institute and State Key Laboratory of Continental Dynamics, Department of Geology, Northwest University, Xian 710069, PR China & Research Center for Orogenic Geology, Xian Geological Survey Center, China Geological Survey, Xian 710054, PR China; Jian Wang [332891824@qq.com], Ju Zhang [710726657@qq.com], Research Center for Orogenic Geology, Xian Geological Survey Center, China Geological Survey, Xian 710054, PR China.

Graptolites represent the standard tools for biozonation and correlation of Ordovician and Silurian strata. Oktavites spiralis, one of the index graptolite species of the Telychian Age (late Llandovery, Silurian), is of great significance in biostratigraphy. However, the broad definition of the species and the lack of data on its evolutionary changes have led to controversies regarding its stratigraphic range and, consequently, to difficulties in regional correlations. Recent investigations in multiple Telychian profiles in the Ziyang–Langao area, along the northern margin of South China, reveal that the first appearance datum of Oktavites spiralis is located at the base of the Spirograptus turriculatus Zone, and its last appearance datum lies at the top of the Cyrtograptus lapworthi Zone (Telychian stage). The thecae vary markedly up-section and such change is consistent across all studied sections. This morphological change can be used as the basis for accurate stratigraphic subdivision and, hence, for effective regional correlation. In addition, this species has a wide range in population size and population density relative to other coexisting graptolite species, and can be better distinguished by using a more precise set of criteria.     

Evolutionary morphological changes in the new genus Duoforisa: implication for classification and habitat of the unilocular Foraminifera

Qianyu Li & Brian Mcgowran, 1994:03:28. Evolutionary morphological changes in the new genus Duoforisa: implication for classification and habit of the unilocular Foraminifera. Alcheringa 18, 121–134. ISSN 0311-5518.

Unlike other unilocular foraminifera, the new genus Duoforisa from the Late Oligocene to Early Miocene possesses a kidney-shaped test with two apertures on its distal ends. It contains two distinct and successional species, both new, and differentiated by their apertural details. In Duoforisa rima the apertures are slit-like, and become radial in the descendant D. diducta. Intermediate forms have transitional apertural configurations between the slit type and radial type, accompanied also by a change of the test outline from subtriangular to compactly U-shaped. The evolution of the lineage occurred during a period of enhanced upwelling in the Early Miocene and it was terminated just before the global warming at the Early-Middle Miocene boundary. This example suggests evolution of the unilocular foraminifera through successional morphological changes in test shape and in the aperture. Unilocular taxa have tended to flourish or speciate in cool or upwelling environments. Their contraction in the latest Early Miocene to early Middle Miocene was probably due to global warming and well oxygenated conditions which were widespread in the neritic domains of southern Australia.     

Two middle Cambrian trilobite genera,Cyclolorenzella Kobayashi, 1960 and Jiulongshania gen. nov., from Korea and China

As is widely recognised, fossil florules are difficult to correlate because they appear to have been more controlled by environmental and ecological factors than uniform changes which reflect the passage of geological time. Here I present a worked example of a generally applicable approach to this problem in dating and mapping fossil plant associations.

Floral associations are best described and named by the classical methods of phytosociology. The three dimensional shape of an association in rocks and its palaeoecology can then be critically assessed. For example, during Middle Triassic time the following plant associations would have been encountered on a traverse from the coast to several hundred kilometres inland from the Pacific margin of Gondwanaland: Pachydermophylletum (mangrove scrub), Linguifolietum (coastal swamp woodland), Dicroidietum odontopteroidium (floodplain forest), Phoenicopsetum (levee bank scrub), Dicroidietum odontopteroidium xylopterosum (xerophytic woodland) and Johnstonietum (mallee-like woodland).

The fourth dimension of fossil plant associations (time) is best assessed from the evolution of a prominent group of plants. In conjunction with the ranges of other plant megafossils, evolutionary changes can be used to define Oppel-zones. For the Late Permian to Early Jurassic of eastern Australasia, I have used the evolution of Dicroidium and leaves of related pteridosperm plants together with other species of restricted stratigraphic range or prominent in the definition of the fossil plant associations. From this evidence four Oppel-zones can be recognised. ‘Thinnfeldia’ callipteroides Oppel-zone (Chhidruan to Smithian, 250–230 × 10⁶ years), Dicroidium zuberi Oppel-zone (Smithian to Middle Anisian, 230–220 × 10⁶ years), D. odontopteroides Oppel-zone (Late Anisian to Ladinian, 220–210 × 10⁶ years) and Yabeiella Oppel-zone (Carnian to Rhaetian, 210–200 × 10⁶ years.

In a complete revision of the pteridosperm form-genera Dicroidium, Johnstonia and Xylopteris, a large number of new combinations and the following new taxa are proposed; Dicroidium gouldii sp. nov., D. odontopteroides var. argenteum var. nov., D. odontopteroides var. moltenense var. nov., D. radiatum sp. nov., D. townrovii sp. nov. and Johnstonia coriacea var. obesa var. nov.     

A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida

Zhang, Y., He, W.H., Shi, G.R., Zhang, K.X. & Wu, H.T., 26.2.2015. A new Changhsingian (Late Permian) brachiopod fauna from the Zhongzhai section (South China) Part 3: Productida. Alcheringa 39, xxx–xxx. ISSN 0311-5518.

As the third and last part of a systematic palaeontological study of the brachiopod fauna from the Permian–Triassic boundary section at Zhongzhai in Guizhou Province (South China), this paper reports 15 species (including three new species: Tethyochonetes minor sp. nov., Neochonetes (Zhongyingia) transversa sp. nov., Paryphella acutula sp. nov.) in Order Productida. In addition, the morphological features and definitions of several key Changhsingian brachiopod taxa (e.g., Paryphella and Oldhamina interrupta) are clarified and revised.

Yang Zhang* [zhangy05@163.com] and G.R. Shi [guang.shi@deakin.edu.au], School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; Weihong He [whzhang@cug.edu.cn] and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China; Huiting Wu [ht_wu415@163.com], Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China. *Also affiliated with: Faculty of Earth Sciences, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China.     

A new diminutive euphyllophyte from the Middle Devonian of West Junggar,Xinjiang, China and its evolutionary implications

Xu, H.-H., Wang, Y., Tang, P. & Wang, Y., May 2017. A new diminutive euphyllophyte from the Middle Devonian of West Junggar, Xinjiang, China and its evolutionary implications. Alcheringa 41, 524–531. ISSN 0311-5518.

A diminutive euphyllophyte, Douaphyton levigata gen. et sp. nov., is described from the upper Middle Devonian (Givetian) Hujiersite Formation of West Junggar, Xinjiang, China. The plant consists of more than three orders of axis branching, each axis being less than 2 mm wide. The second-order axes are short, laterally and alternately attached to the main axis. The third-order axes are paired and anisotomously divided, bearing the vegetative appendages or the fertile units. The fertile unit consists of a short recurved axis giving off up to four short pedicels along one side, each of which bears one to four pairs of terminal sporangia. Douaphyton has a three-dimensional branching system that has an intermediate form in the evolutionary context of euphyllophytes and lignophytes. It is also proposed that complex branching developed in multiple groups in the Middle Devonian.

*Hong-He Xu [hhxu@nigpas.ac.cn], Yao Wang [yaowang@nigpas.ac.cn], Peng Tang [pengtang@nigpas.ac.cn], Yi Wang [yiwang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210,008, PR China. Yao Wang [yaowang@nigpas.ac.cn] University of Science and Technology of China. 96 Jinzhai Road, Hefei, Anhui Province, 230,026, PR China.     

Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America

Camilleri, T.A., Warne, M.T., Holloway, D.J. & Weldon, E.A., 10 May 2019. Revision of the ostracod genus Velibeyrichia Henningsmoen, 1954 from the Silurian and Lower Devonian of North America. Alcheringa XXX, X–X. ISSN 0311-5518.

Known occurrences of the ostracod genus Velibeyrichia are restricted to a number of Silurian to Lower Devonian geological strata in North America: the McKenzie Member of the Mifflintown Formation of Maryland and West Virginia; the Tonoloway Limestone of Maryland, West Virginia, Virginia and Pennsylvania; the Bloomsburg Formation of Maryland, Virginia and Pennsylvania; the Manlius Limestone of New York; and the Decker Limestone of New Jersey and New York. The genus includes six species: V. moodeyi (type species), V. mesleri, V. paucigranulosa, V. reticulosaccula, V. tonolowayensis and V. tricornia. The diagnostic combination of characters for this genus are: distinct deflection of the velum where it crosses the crumina in heteromorphs (adult female specimens), dorsal nodes on lobes L1 and L3, sexual dimorphism of the velum, and in tecnomorph specimens, either a shallow sulcus on lobe L3 or a zygal ridge (in adult tecnomorph specimens) extending from lobe L2 to lobe L3. The presence of one or the other of the latter two characters defines two distinct species groups.

Tamara T.A. Camilleri* [tamara.camilleri@deakin.edu.au], Mark T. Warne* [mark.warne@deakin.edu.au] and Elizabeth A. Weldon [l.weldon@deakin.edu.au], Deakin University, Geelong, School of Life and Environmental Sciences & Centre for Integrative Ecology (Melbourne Campus), 221 Burwood Highway, Burwood, Victoria 3125, Australia; David J. Holloway [dhollow@museum.vic.gov.au], Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia. *Also affiliated with: Museums Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia.     

Pragian and lower Emsian (Lower Devonian) conodonts from Liujing,Guangxi, South China

LU, J.-F., QIE, W.-Q. & CHEN, X.-Q., July 2016. Pragian and lower Emsian (Lower Devonian) conodonts from Liujing, Guangxi, South China. Alcheringa 40, xxx–xxx. ISSN 0311-5518.

Lower Devonian (Pragian and Emsian) conodonts are described from the Liujing section in Guangxi, South China, highlighting the phylogeny of early polygnathids and establishing the basis for faunal correlations with deposits throughout the world. Records of Eognathodus kuangi sp. nov. and E. nagaolingensis Xiong increase the biodiversity of the Pragian eognathodids in South China and represent intermediate stages and probably the latest eognathodids in the evolutionary succession from Eognathodus to Polygnathus. During this evolutionary succession, the degeneration of the sulcus (or the flattening of the upper platform surface) is the most important morphological change, especially in the eognathodid lineage. Polygnathus trilinearis, P. pireneae and P. sokolovi are recognized together for the first time in South China. The contemporaneous occurrences of the kitabiformis and sokoloviformis morphs of P. pireneae with P. sokolovi and their respective similarities to P. kitabicus and P. sokolovi suggest that the latter two species are phylogenetically linked with P. pireneae. Evidence from the Liujing section also favours Polygnathus probably having evolved from Eognathodus in a tropical or subtropical area.

Jian-feng Lu* [lujfivan@sina.cn] and Xiu-qin Chen [xqchen@nigpas.ac.cn], Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Wen-Kun Qie [wkqie@nigpas.ac.cn], Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing 210008, PR China. *Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China.     

A new genus and species of Steninae from the late Eocene of France (Coleoptera,Staphylinidae)

Cai, C., Clarke, D.J., Huang, D. & Nel, A., 2014. A new genus and species of Steninae from the late Eocene of France (Coleoptera, Staphylinidae). Alcheringa 38, 557–562. ISSN 0311-5518.

A remarkable new genus and species of rove beetle, Eocenostenus fossilis gen. et sp. nov., is described and figured based on one well-preserved specimen from the late Eocene of Monteils (near Alès, Gard, France). Eocenostenus is definitively placed in the extant subfamily Steninae, based on the combination of dense and coarse body punctation, globular and protruding eyes, exposed and closely spaced antennal insertions on the vertex, and six visible abdominal terga. Eocenostenus differs from the two extant stenine genera Stenus and Dianous most notably in the structure of the prothorax, which is strongly transverse and with unusual anterolateral projections, and in the anteriorly placed antennal insertions. This new discovery highlights the palaeodiversity of a genus-poor subfamily and suggests that the early diversification of Steninae is probably complicated.

Chenyang Cai [caichenyang1988@163.com] and Diying Huang [dyhuang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Rd., Nanjing 210008, PR China; Dave J Clarke [dclarke@fieldmuseum.org], Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA; and André Nel [anel@mnhn.fr], Muséum National d’Histoire Naturelle, Institut de Systématique, Evolution, Biodiversité, ISYEB, UMR 7205 CNRS UPMC EPHE, CP50, 45 rue Buffon, F-75005 Paris, France. Received 23.1.2014; revised 2.5.2014; accepted 12.5.2014.     

A new Cenozoic cyclostome bryozoan genus from Argentina and New Zealand: strengthening the biogeographical links between South America and Australasia

Taylor, P.D. & Brezina, S., February 2018. A new Cenozoic cyclostome bryozoan genus from Argentina and New Zealand: strengthening the biogeographical links between South America and Australasia. Alcheringa XX, xx–xx. ISSN 0311-5518.

Uniserial encrusting cyclostome bryozoans (‘stomatoporiforms’) are especially challenging to study taxonomically because of the paucity of their morphological characters. Here we introduce Axilosoecia gen. nov. for two previously undescribed species characterized by gonozooidal brood chambers located in the axils of branch bifurcations. The type species, Axilosoecia giselae sp. nov., comes from the upper, Danian part of the Roca Formation of La Pampa, Argentina; the second species, Axilosoecia mediorubiensis sp. nov., is from the lower Miocene of Southland, New Zealand. On account of its basal gonozooids, Axilosoecia is assigned to the family Oncousoeciidae despite similarities in colony form with Stomatoporidae. The two known occurrences of this new genus support previously suggested biogeographical links between southern South America and Australasia.

Paul D. Taylor, Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK [p.taylor@nhm.ac.uk]; Soledad Brezina, Instituto de Investigación en Paleobiología y Geología, Unidad de doble dependencia, Universidad Nacional de Río Negro—Consejo Nacional de Investigaciones Científicas y Técnicas, Av. Roca 1242, R8332FDJ, General Roca, Río Negro, Argentina [sbrezina@unrn.edu.ar].     

A new species of Aboilus (Insecta,Orthoptera) from the Jurassic Daohugou beds of China,and discussion of forewing coloration in Aboilus

Wang, H., Li, S., Zhang, Q., Fang, Y., Wang, B. & Zhang, H., 13.02.2015. A new species of Aboilus (Insecta, Orthoptera) from the Jurassic Daohugou beds of China, and discussion of forewing coloration in Aboilus. Alcheringa 39, xxx–xxx. ISSN 0311-5518

He Wang* [wanghe0701@163.com], Sha Li* [lisha.paleo@gmail.com], Qi Zhang* [zhqi1105@126.com], Yan Fang [yanfang@nigpas.ac.cn], Bo Wang? [bowang@nigpas.ac.cn] and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China.*Also affiliated with University of Chinese Academy of Sciences, Beijing 100049, PR China. ?Also affiliated with Steinmann Institute, University of Bonn, Bonn 53115, Germany.

A new species of Aboilinae (Orthoptera: Prophalangopsidae), Aboilus perbellus, is described and illustrated based on three well-preserved forewings recovered from the Middle–Upper Jurassic Daohugou beds of Inner Mongolia, China. The new species differs from all congeneric forms in its special forewing coloration and features of its wing venation. To date, three types of forewing coloration have been found among different species of Aboilus at Daohugou, suggesting that these taxa inhabited different ecotopes.     

A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin,northwestern China

Zheng, D., Wang, H., Nel, A., Dou, L., Dai, Z., Wang, B. & Zhang, H. 27 June 2019. A new damsel-dragonfly (Odonata: Anisozygoptera: Campterophlebiidae) from the earliest Jurassic of the Junggar Basin, northwestern China. Alcheringa XX, X–X. ISSN 0311-5518.

A new genus and species of campterophlebiid damsel-dragonfly, Jurassophlebia xinjiangensis gen. et sp. nov., is described from the Lower Jurassic Badaowan Formation in the Junggar Basin, northwestern China. Jurassophlebia differs from all other campterophlebiid genera in having PsA in the same orientation as the distal branch of AA, and in its uniquely open subdiscoidal cell with very acute apical angle in the hind wing. The new discovery adds to the Asian diversity of damsel-dragonflies in the earliest Jurassic.

Daran Zheng* [dranzheng@gmail.com], He Wang [hwang@nigpas.ac.cn], Bo Wang [bowang@nigpas.ac.cn], and Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Longhui Dou [345317122@qq.com], Comprehensive Geology Exploration Team, Xinjiang Coalfield Geology Bureau, West Mountain Road, Ürümqi 830000, PR China; Zhenlong Dai [286321388@qq.com], No.9 Geological Team, Xinjiang Bureau of Geology and Mineral Resources, Ürümqi 830011, PR China; Daran Zheng also affiliated with Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China.