New Human Remains of Archaic Homo sapiens and Lower Palaeolithic Industries from Visogliano (Duino Aurisina, Trieste, Italy)

New human remains have been found during recent excavations conducted in the lower section of Visogliano dolina (Trieste, Italy). They are represented by teeth and a fragment of particularly robust mandible that for their morphological and morphometrical characters can be generically attributed to archaic Homo sapiens. The lithic industry is formed by flakes, carinated points, scrapers, choppers and chopping tools and is characteristic of the Italian Pebble Culture. Sediments, pollen and faunal remains indicate that this part of the deposit was formed during interglacial climatic conditions in the early, but not initial phase of Middle Pleistocene (isotopic stadium 13 or 11).     

The lower paleolithic of the Near East

The Near East forms the geographic crossroads between Africa, Asia and Europe and was certainly a main route for the dispersal of Homo erectusinto Eurasia. The study of Lower Paleolithic sites in this region and in the neighboring Caucasus area sheds some light on several potential colonization events. Sites such as Ubeidiya (Jordan Valley) and Dmanisi (Georgia) suggest the early sorties took place around 1.4-1.0 Ma. Despite the lack of radiometric dates, sequences of raised beaches, marine deposits, river terraces, and paleolake formations have enabled various investigators to identify several series of major aggradation and erosion periods within the Pleistocene. Lithic assemblages derived from a few systematic excavations and collections from stratigraphically dated outcrops led to a threefold subdivision of the Acheulian sequence into the Lower, Middle, and Upper Acheulian. The study of nonbiface assemblages, however, has not resolved the question of whether these assemblages deserve inclusion as separate entities or should be viewed as sites within the Acheulian settlement pattern. While the typotechnological definitions of each major phase can be compared to what is known from other regions of the Old World, the Acheulo-Yabrudian (or the Mugharan Tradition) is seen as a local entity. Rare human remains and scarce data concerning subsistence activities do not warrant a comparative discussion with what is known from African and some European sites.     

A new Jungle-fowl from the Pleistocene of Europe

The domesticated Red Jungle-fowl G. gallus is believed to have been dispersed by man from India during the Holocene. The distal end of a radius from the Ipswichian Interglacial deposits at Crayford, Kent, was indistinguishable from that of the wild form of Red Jungle-folw. A coracoid from the early Middle Pleistocene of N Norfolk was also very similar to that species but showed differences comparable with those found between different species of Gallus. At least one species referable to Gallus is known from the Pliocene of South-eastern Europe. Extrapolating from zoogeographical speciation patterns it would be possible for a Gallus species to have evolved through Pleistocene speciation in the European region. The species might have become extinct during a glaciation or have been exterminated by early man. A new species Gallus europaeus is described with the coracoid as a holotype, and the radius from Kent is tentatively referred to it.     

Human consumption of tortoises at Level IV of Bolomor Cave (Valencia, Spain)

The acquisition and consumption of small prey in the pre-Upper Palaeolithic is a highly debated topic at present. For some authors, the systematic obtaining of these animals is only part of the subsistence strategies used by anatomically modern Humans. However, the consumption of small prey dates back to the Plio-Pleistocene chronologies in some sites. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of tortoise consumption is documented not as common for these periods. However, Level IV of Bolomor Cave has clear diagnostic elements to document the acquisition and use of tortoises (Testudo hermanni) for food in the form of: (1) cutmarks on limb bones and ventral surface of the carapace and plastron; (2) presence of burning on tortoise skeleton and shell; (3) elements of anthropogenic breakage on carapace and plastron: percussion pits, percussion notches and impact flakes; and (4) human toothmarks on limb bones. This paper tries to examine the possible patterns in the tortoise consumption sequence from Level IV of Bolomor Cave and improves data on the butchery process and tortoise consumption in the Late Middle Pleistocene.     

Lagostomus maximus (Desmarest) (Rodentia,Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay

Ubilla, M. & Rinderknecht, A., April 2016. Lagostomus maximus (Desmarest) (Rodentia, Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The extant plains vizcacha, Lagostomus maximus, is described from the Late Pleistocene (Dolores Formation) of Uruguay based on an almost complete articulated skeleton. It is compared with the nominally extinct Pleistocene species of the genus. An AMS ¹⁴C taxon-age is determined for L. maximus at 11 879 ± 95 years BP (cal. BP 13 898–13 941). Lagostomus maximus is absent from modern mammal communities in Uruguay, and no Holocene evidence is available. Because L. maximus exhibits remarkable sexual and ontogenetic dimorphism, we examined a range of juvenile, sub-adult and adult male and female specimens. It is not possible to differentiate the articulated Pleistocene fossil from sub-adult specimens of L. maximus based on this sample. Moreover, the skull characteristics, including a broad vertical ramus of the zygomatic arch, semi-circular temporal crest and short robust sagittal crest, most closely resemble extant female individuals. Molar size is non-predictive for ontogenetic stage or body mass because it reaches stability during early adulthood. Other fragmentary skull remains are herein assigned to L. sp. cf. L. maximus. Lagostomus cavifrons from the Pleistocene of Argentina is considered synonymous with L. maximus. Its character states concur with the range of intraspecific variability and it has been established on a young adult or adult male individual. Likewise, the other Argentinean Pleistocene species based on incomplete mandibles, L. angustidens, L. striatus, L. heterogenidens, L. egenus, L. minimus and L. debilis, concur with ontogenetic morphs and are here assigned to Lagostomus sp. The ecological preferences of extant L. maximus infer open arid or semi-arid landscapes for the latest Pleistocene of southern Uruguay. This hypothesis is reinforced by the coeval presence of Microcavia, Galea and small camelids in the same strata. The Last Glacial Maximum likely promoted this environmental setting. Latest Pleistocene or early Holocene climatic change might have facilitated local extinctions and/or range shifts among this mammal fauna.

Martín Ubilla [martinubilla@gmail.com], Facultad de Ciencias, Universidad de la Republica, Iguá 4225, 11400, Montevideo, Uruguay; Andrés Rinderknecht [apaleorinder@yahoo.com], Museo Nacional de Historia Natural, Paleontología, 11000, Montevideo, Uruguay.     

Middle Pleistocene bird consumption at Level XI of Bolomor Cave (Valencia, Spain)

The consumption of small prey dates back to the Plio-Pleistocene chronologies in some African sites. However, the systematic acquisition and consumption of small prey in the pre-Upper Palaeolithic times is still a highly debated topic in Europe. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of bird consumption is not as common for these periods. Nevertheless, Level XI (MIS 6) of Bolomor Cave has clear diagnostic elements to document the acquisition and use of birds (Aythya sp.) for food in the form of: (1) cutmarks on bones of both the front and hind limb; (2) presence of burning patterns on the extremities of the bones (areas of the skeleton with less meat); and (3) human toothmarks on limb bones. The capture of birds is classified as quick-flying game in the archaeological sites. The acquiring of fast-running (mostly lagomorphs) and quick-flying small prey requires a sophisticated technology and involves obtaining and processing ways different from those used for large- and medium-sized animals. From this perspective, the aim of this paper is to examine possible patterns in the processing sequence of birds from Level XI of Bolomor Cave and to improve the data on their butchery and human consumption in the Middle Pleistocene of Iberian Peninsula.     

New record of Warendja wakefieldi (Vombatidae; Marsupialia) from Wombeyan Caves,New South Wales

Transverse sections of Chirodipterus australis and Griphognathus whitei are used to demonstrate the histological structures of the rostral and symphysial tubuli in Devonian lungfish. The walls of the tubuli are composed of bony tissue indistinguishable from the cancellar bone found in dermal bone. It has many spaces and perforations, and is contiguous in places with the cancellar bone of the external dermal bone. The walls of lateral-line canals have the same histological structure as the walls of the tubuli; they also intercommunicate with them. The tubuli and the lateral-line canals open to the surface through large perforations on the external surface. In Griphognathus the walls of the tubes under the external dermal bone have two layers: the outer one is dense bone, but the inner one has the appearance of calcified fibrous connective tissue of the kind associated with the tissue surrounding the lateral-line canals in living lungfish. The tubuli are closely related to the pore-canal system via canals that penetrate the dermal bone; this same relationship has been observed in Dipnorhynchus and Speonesydrion. The intimate connection between the tubuli and the lateral-line system suggests that they were formed by the sinking of neuromasts into and under the external dermal bone, with their walls surrounded by cancellar bone.     

Getting “Out of Africa”: Sea Crossings,Land Crossings and Culture in the Hominin Migrations

Palaeoanthropologists and archaeologists have advanced a wide range of explanatory narratives for the various movements of Homo erectus/Homo ergaster, and the first modern Homo sapiens, “Out of Africa”—or even back again. The application of Occam's razor—a parsimonious approach to causes—gives a more cautious approach. There is nothing in the available evidence that would require the ability for a human water crossing from Africa before the later Pleistocene, whether across the Strait of Gibraltar, the Sicilian Channel or the southern Red Sea (Bab el-Mandab). A parsimonious narrative is consistent with movements across the Sinai peninsula. The continuous arid zone from northern Africa to western Asia allowed both occupation and transit during wet phases of the Pleistocene; there is no requirement for a “sponge” model of absorption followed by expulsion of human groups. The Nile Valley as a possible transit route from East Africa has a geological chronology that could fit well much current evidence for the timing of human migration. The limited spatial and temporal opportunities for movements “Out of Africa,” or back again, also puts particular difficulties in the way of the gene flow required for the multiregional hypothesis of the development of modern Homo sapiens.     

ESR dose response of Al center measured in quartz samples from the Yellow River (China): Implications for the dating of Upper Pleistocene sediment

The ESR dating method requires to describe the evolution of the ESR signal intensities vs. increasing gamma doses, then to extrapolate the equivalent dose of radiation received by the sample since its deposition using mathematical fitting. The function classically used to describe the growth curves of ESR aluminium signal in quartz was recently discussed and challenged for Lower Pleistocene sediments. In the present work, some alluvial sediments sampled in Upper Pleistocene fluvial terraces of the Yellow River system (China) permit us to test the application of another extrapolation function (linear + exponential) recently proposed for Lower Pleistocene sediments. The equivalent doses obtained here for the recent deposits of the Yellow River system and the corresponding ages are promising and indicate the potential of ESR to date quartz deposits from Upper Pleistocene times.     

Further Ektopodontidae (Phalangeroidea,Mammalia) from southwestern Victoria

The youngest members of the Ektopodontidae, an enigmatic family of Phalangeroidea, are reported from southwestern Victoria. Four isolated teeth are described as Ektopodon paucicristata sp. nov. (from the ?mid-?late Pliocene, and early Pliocene) and two isolated Pleistocene teeth are assigned to Darcius duggani Rich, 1986. Previously, the youngest known ektopodontid was early Pliocene.     

Reappraisal of ‘chronospecies’ and the use of Arvicola (Rodentia,Mammalia) for biochronology

The water vole, genus Arvicola, is characterised by a broad geographic distribution throughout Europe and is widespread during the late Middle and Upper Pleistocene. This genus is used as a major biostratigraphic tool within the Quaternary. Specific determinations using the Schmelzband-Differenzierung-Quotient or SDQ have identified many chronospecies within the fossil species Arvicola cantiana (Hinton, 1910). As SDQ calculation remains limited, this study reappraises the Arvicola genus in terms of morphodiversity and morphospace using outline analysis which takes into account the tooth as a whole. Outline analysis suggests that one single species of Arvicola, A. cantiana, was present during the Pleistocene. This species shows great variability with no trends or patterns in morphospace. Thus, these results call into question the reliability of SDQ for specific determinations and throw doubt on the biochronological framework based on Arvicola.     

Fossil fruit identification by SEM investigations, a tool for palaeoenvironmental reconstruction of Dmanisi site, Georgia

Archaeological excavations and multidisciplinary studies at the site of Dmanisi (Lesser Caucasus, Georgia) have provided an assemblage of lithic artefacts and numerous palaeontological and human remains in a volcano-sedimentary context dating to the early Lower Pleistocene. The sieving of archaeological sediments from Dmanisi also yielded mineralised botanical macro-remains that were analysed according to a particular protocol. The use of scanning electron microscopy (SEM) allowed the comparison between fossil fruit remains and modern specimens, leading to a precise identification of the earlier. Seed remains recognised by this method belonging to several taxa from the Boraginaceae family as well as to the genus Celtis L., of the Ulmaceae family. The ecological requirements of these taxa suggest the presence of xeric conditions, a result that is in accordance with data obtained by faunal and stratigraphic studies.     

Lithostratigraphy of Border Cave, KwaZulu, South Africa: a Middle Stone Age sequence beginning c. 195,000 b.p.

Border Cave is well-known for its Middle Stone Age (MSA) sequence and associated hominids, as well as for the earliest demonstrable Later Stone Age (LSA) (c. 38,000 b.p.) strata in southern Africa. Detailed lithostratigraphic and sedimentological study permits identification of 8 Pleistocene sedimentary cycles, including 6 major cold phases and 2 intervening weathering horizons. The 2 youngest cold phases are associated with the LSA and have 8 ¹⁴C dates 38,600-13,300 b.p. By gauging sedimentation rates in finer and coarser sediments, duration of sedimentary breaks, and allowing for differential compaction, the excellent radiocarbon framework provided by 28 available ¹⁴C dates can be extrapolated to the 6 cold intervals and 2 palaeosols that are older than 50,000 b.p. These clearly span oxygenisotope stages 4, 5 and 6, placing the base of the MSA deposits at c. 195,000 b.p., Homo sapiens sapiens at c. 90,000–115,000 b.p. and the sophisticated, microlithic “Howieson's Poort” industry at 95,000 b.p. These results require radical reassessment of the age and nature of the MSA complex and of the earliest evolution of anatomically-modern people.     

On the occurrence of the scimitar-toothed cat, Homotherium latidens (Carnivora; Felidae), at Kents Cavern,England

Teeth of Homotherium latidens recovered from late Pleistocene sediments, Kents Cavern, England have long been the source of controversy. H. latidens is conspicuously absent from other late Pleistocene cave deposits in Britain, and is widely thought to have been extirpated from the region during the isotope stage 10 glacial period. Here we present high spatial resolution analyses of fluorine and uranium uptake profiles in teeth of three species from the same cave. The H. latidens tooth is clearly distinguished from the unambiguously provenanced Late Pleistocene hyaena and Middle Pleistocene cave bear teeth. These results are consistent with the theory that the H. latidens teeth originated at an exogenous location, were probably transported to Kents Cavern as Palaeolithic trade goods, and were buried in Kents Cavern in Palaeolithic times.     

The genera Cryptaulax and Procerithium (Procerithiidae,Caenogastropoda) in the Early Jurassic of Patagonia,Argentina

Ferrari, S.M., September 2012. The genera Cryptaulax and Procerithium (Procerithiidae, Caenogastropoda) in the Early Jurassic of Patagonia, Argentina. Alcheringa 36, 325–339. ISSN 0311-5518.

New species of Cryptaulax and Procerithium (Procerithiidae, Caenogastropoda) are reported from the Lower Jurassic (Pliensbachian–Toarcian) marine deposits of west-central Chubut province, Argentinean Patagonia. Three new species are described: Cryptaulax redelii, Procerithium (Rhabdocolpus) patagoniensis and Procerithium (Infacerithum) nodosum; and the diagnoses of Cryptaulax damboreneae Ferrari and Procerithium nulloi (Ferrari) are emended. The new fossils derive from the Mulanguiñeu and Osta Arena formations and expand the known diversity of the Procerithiidae, extending its palaeobioeographical distribution into the South American Jurassic.     

Paleodemographic analysis of a fossil porcupine (Hystrix refossa Gervais, 1852) population from the Upper Pleistocene site of Geula Cave (Mount Carmel,Israel)

For decades zooarchaeological studies mainly focused on teeth age profiles to derive central hypotheses concerning human subsistence during the Pleistocene. Behavior was often related to hunting and/or scavenging strategies according to two models: attritional versus catastrophic. However, few studies have estimated basic demographic parameters using standard methods of population ecology, which are necessary for a more confident interpretation of animal paleopopulation structures. Thanks to a remarkably rich fossil bone assemblage from the Upper Pleistocene cave of Geula (Mount Carmel, Israel), in this paper we perform a paleodemographic analysis of Hystrix refossa, Gervais 1852 (Mammalia, Rodentia). We first consider phylogenetic and morphometric dental criteria for the comparison of Pleistocene porcupine species, followed by an examination of taphonomic conditions of the bone assemblage. We focus strongly on the age structure of the fossil population, presenting a new methodological approach that combines life tables and Leslie matrix models – one of the best-known methods in population ecology for assessing population growth and age distribution. We detail demographic parameters derived from cross-sectional life tables and past trends in the porcupine population at Geula Cave using simple matrix projection model. Finally, we were able to perform an elasticity analysis to identify which demographic component was potentially the most critical for influencing growth. We show that the population of H. refossa did not decline or shrink but was stable. Our results permit us to characterize Geula Cave as a natural shelter for porcupines, with limited evidence to scavenging and hunting by either hyenas or humans.     

The first record of Galea Meyen, 1832 (Rodentia,Hystricognathi, Caviidae) in the late Pleistocene of southern Brazil and its palaeobiogeographic implications

The first caviine rodent referable to Galea Meyen, 1832 is described from the late Pleistocene of southern Brazil based on a left dentary with the p4–m3 series. The specimen derives from the Ponte Velha I locality in the Touro Passo Creek (Touro Passo Formation, upper Pleistocene), western Rio Grande do Sul State. The main characters used to assign this specimen to Galea are: anterior area of horizontal crest at the level of prism I of p4; deep anterior area of masseteric fossa; incisor alveolus on the medial face of the dentary extended up to the level of prism II of m2; and presence of cement in the hypoflexid. Currently, the genus has a disjunct distribution, with a group in Argentina, Bolivia and Peru, and another in northern and northeastern Brazil. The presence of this taxon in Pleistocene deposits of Rio Grande do Sul State, Uruguay and the Argentine Mesopotamian, where there are no extant representatives of the genus, indicates its wider distribution during the late Pleistocene.     

MOLLUSC SHELL SIZES IN ARCHAEOLOGICAL CONTEXTS IN NORTHERN SPAIN (13 200 TO 2600 CAL BC): NEW DATA FROM LA GARMA A AND LOS GITANOS (CANTABRIA)

Shellfish metrical data are a source of information about the exploitation of marine resources in the past. In this study, we propose a methodological approach based on the size structures of different rocky intertidal gastropod species. Three limpet species (Patella vulgata, Patella intermedia and Patella ulyssiponensis) and the toothed topshell Osilinus lineatus are studied from two sites in Cantabrian Spain: La Garma A and Los Gitanos caves over a period of 10 000 years, covering the Pleistocene–Holocene transition. Data are also supplied about a further sea snail species, the periwinkle Littorina littorea (Upper Magdalenian). A reduction in size can be seen, between the upper Magdalenian and the late Neolithic, in the case of the first four species. The explanation for this decline is probably related to the climate change that occurred in the transition between the Late Pleistocene and the Holocene, but it is possible that human impact might also have influenced shell sizes in the Mesolithic and Neolithic.     

Taphonomic analysis of remains of mammals eaten by pumas (Puma concolor Carnivora, Felidae) in central Argentina

Prey remains recovered from puma (Puma concolor) scat were analyzed in order to identify taphonomic features produced on them by this predator. Mammal remains are abundant in puma scat, but recognition of bone remains decreases as prey size increases. Modifications produced by digestion are strong. However, micromammal bones show a gradation in preservation that includes well preserved specimens. Data presented here and their interpretation may possibly be extrapolated to zooarchaeological or paleontological assemblage.