New specimens of Mourasuchus (Alligatoroidea,Caimaninae) from the Miocene of Brazil and Bolivia and their taxonomic and morphological implications

Cidade, G.M., Souza-Filho, J.P., Hsiou, A.S., Brochu, C.A., & Riff, D., 18 March 2019. New specimens of Mourasuchus (Alligatoroidea, Caimaninae) from the Miocene of Brazil and Bolivia and their taxonomic and anatomic implications. Alcheringa 43, 261–278. ISSN 0311-5518.

Mourasuchus is one of the most peculiar crocodylians of all time, showing an unusual ‘duck-faced’ rostrum with thin, gracile mandibles. It includes four species restricted to the South American Miocene. Here, we describe ten late Miocene specimens of Mourasuchus, nine from the Solimões Formation of Brazil and one from Bolivia. All specimens are assigned to M. arendsi, but this assignment may change as the diversity and relationships within Mourasuchus are better understood. We also discuss several issues pertinent to the morphology of Mourasuchus: the presence of a braincase neomorph (the laterocaudal bridge), hypotheses about sexual dimorphism, the function of the squamosal ‘horns’ the presence of possible thermoregulatory functions in the genus. Additionally, the paleogeographic distribution of Mourasuchus in the Miocene of South America is also discussed.

Giovanne M. Cidade* [giovannecidade@hotmail.com], Universidade de São Paulo Campus de Ribeirao Preto, Biologia, Avenida Bandeirantes 3900, Ribeirao Preto, 14040-900, Brazil; Jonas P. Souza-filho [jpdesouzafilho@hotmail.com], Universidade Federal do Acre, Departamento de Ciências da Natureza, Campus Universitário, UFAC, BR 364, Km 4, Distrito industrial, CEP 69915-900, Rio Branco, Acre, Brazil; Annie Schmaltz Hsiou [anniehsiou@ffclrp.usp.br], Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Departamento de Biologia, Avenida dos Bandeirantes, 3900, Ribeirão Preto, 14040-901, Brazil; Christopher A. Brochu [chris-brochu@uiowa.edu], University of Iowa, Iowa City, IA 52245, USA; Douglas Riff [driff2@gmail.com], Universidade Federal de Uberlandia, Uberlandia, 38400902, Brazil.     

New fossil sawflies (Hymenoptera,Xyelidae) from the Middle Jurassic of northeastern China

Zheng, Y., Chen, J., Zhang, J. & Zhang, H., 6 August 2019. New fossil sawflies (Hymenoptera, Xyelidae) from the Middle Jurassic of northeastern China. Alcheringa 44, 115–120. ISSN 0311-5518.

Magnaxyela rara gen. et sp. nov. and Abrotoxyela curva sp. nov. (Xyelidae, Macroxyelinae, Gigantoxyelini) are described from the Middle Jurassic Jiulongshan Formation at Daohugou Village, Inner Mongolia, China. The two new species are confidently assigned to tribe Gigantoxyelini of subfamily Marcroxyelinae based on their wing venation and various other morphological characters. Magnaxyela rara is established based on the following forewing characters: pterostigma sclerotized completely, Sc two-branched and meeting R before origin of Rs, 1-Rs as long as 1-M, but shorter than Rs?+?M, R curved bewteen 1r-rs and 2r-rs and cell 2r longer than 1r. Abrotoxyela curva is differentiated from congeneric forms by the two-branched vein Sc of the forewing, Sc2 short and inclined towards the base of the wing, 1-Rs, 1-M and Rs?+?M nearly equal in length, crossvein 1m-cu longer than 1cu-a. The fossils described herein increase the diversity of the Mesozoic Xyelidae.

Yan Zheng* [zhengyan536@163.com], Jun Chen* [ubiscada@sina.com], and Junqiang Zhang [zhangjunqiang@lyu.edu.cn], Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi 276000, PR China; Haichun Zhang [hczhang@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, PR China. *Also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, East Beijing Road, Nanjing 210008, PR China.     

Late Silurian vertebrate microfossils from the Carnarvon Basin,Western Australia

Burrow, C.J., Turner, S., Trinajstic, K. &; Young, G.C., 27 February 2019. Late Silurian vertebrate microfossils from the Carnarvon Basin, Western Australia. Alcheringa 43, 204–219. ISSN 0311-5518.

A core sample from the offshore Pendock 1A well, Carnarvon Basin, Western Australia yielded microvertebrate residues at an horizon in the lower part of the Hamelin Formation, dated as late Silurian, ? Ludlow, based on associated conodonts. The fish fauna comprises loganelliiform thelodont scales, the ? stem gnathostome Aberrosquama occidens nov. gen. et sp., the acanthodian Nostolepis sp. aff. N. alta, and the ? stem osteichthyan Andreolepis sp. aff. A. petri. Because of the paucity of the material, and some differences between the Pendock scales and those of established species, a precise age can not be confirmed; however, the composition of the fauna at generic level most closely resembles that of late Silurian (Ludlow) assemblages from northern Eurasia.

Carole J. Burrow* [carole.burrow@gmail.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Susan Turner [palaeoich@yahoo.com], Geosciences, Queensland Museum, Hendra QLD 4011, Australia; Kate Trinajstic [k.trinajstic@curtin.edu.au], School of Molecular and Life Sciences, Curtin University, Perth, WA 6102, Australia; Gavin C. Young [gavin.young@anu.edu.au], Research School of Physics and Engineering, Australian National University, Canberra, ACT 2000, Australia.     

A review of Australia’s Mesozoic fishes

Abstract

The Australian Mesozoic fish fauna is considered to be depauperate in comparison with fish faunas in the Northern Hemisphere. However, due to its geographical location as a potential radiation center in the Southern Hemisphere, Australia’s Mesozoic fish fauna is important for understanding fish radiations. Most of the modern fish groups originated during the Mesozoic, but the first records of a modern fish fauna (freshwater and marine) in Australia does not occur until the lower Paleogene. Here, we review all known fossil fish-bearing localities from the Mesozoic of Australia, to improve the understanding of the record. The apparent low Australian Mesozoic fish diversity is likely due to its understudied status of the constituent fossils rather than to a depauperate record. In addition, we review recent work with the aim of placing the Australian Mesozoic fish fauna in a global context. We review the taxonomy of Australian fossil fishes and conclude that the assignments of many actinopterygians need major revision within a modern phylogenetic context. The vast majority of chondrichthyans are yet to be formally described; to the contrary all of the known lungfish specimens have been described. This study considers the microscopic and fragmented remains of Mesozoic fish already found in Australia, allowing a more complete view of the diversity of the fishes that once inhabited this continent.

Rodney W. Berrell [r.berrell@postgrad.curtin.edu.au], School of Earth and Planetary Sciences, Curtin University, Kent Street, Bentley, Western Australia, 6102, Australia; Catherine Boisvert [Catherine.Boisvert@curtin.edu.au], School of Molecular and Life Sciences (MLS), Curtin University, Kent Street, Bentley, Western Australia, 6102, Australia; Kate Trinajstic [K.Trinajstic@curtin.edu.au], School of Molecular and Life Sciences (MLS), Curtin University, Kent Street, Bentley, Western Australia, 6102, Australia; Mikael Siversson# [mikael.siversson@museum.wa.gov.au], Department of Earth and Planetary Sciences, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia, 6106, Australia; Jesús Alvarado-Ortega [jalvarado@yahoo.com.mx], Instituto Geologia Cd universitaria, Circuito de la investigacion, Del. coyoacan, C.P. 04510, Ciudad de México, México; Lionel Cavin [lionel.cavin@ville-ge.ch], Section of Earth Sciences, Muséum d’Histoire naturelle de la Ville de Genèeve, CP 6434, 1211 Genève 6, Switzerland; Steven W. Salisbury [s.salisbury@uq.edu.au], School of Biological Sciences, The University of Queensland, Brisbane St Lucia, Queensland, 4072, Australia; Anne Kemp [annerkemp@gmail.com], 9 Hampton Grove, Norwood, Adelaide, South Australia 5067, Australia. #Also affiliated with: School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102, Australia.     

An extremely large saber-tooth cat skull from Uruguay (late Pleistocene–early Holocene,Dolores Formation): body size and paleobiological implications

Abstract

Among the three recognized species of Smilodon, S. populator is the largest in size and has the widest distribution across South America. The present contribution describes an almost complete skull assigned to the aforementioned felid. The material was recovered from sediments of the Dolores Formation (Lujanian Stage/Age) from the southern part of Uruguay. This specimen is remarkable for its unusual craniodental measurements, indicating that it is one of the largest known specimens of the genus. Estimates of body mass indicate that this individual weighed over 400?kg. In addition, maximum prey size estimation greatly surpasses 1 t and approach nearly 3 t. Based on this, aspects of the paleobiology and paleoecology of S. populator are discussed. Undoubtedly, this kind of carnivorous mammal was at the top of the food chain, with clear adaptations for feeding upon the largest available herbivores, which potentially includes several megafaunal mammal species found in South America during the late Pleistocene.

Aldo Manzuetti [aldomanzuetti@adinet.com.uy], Facultad de Ciencias (UdelaR), Iguá 4225, CP 11400, Montevideo, Uruguay; Daniel Perea [perea@fcien.edu.uy], Facultad de Ciencias (UdelaR), Iguá 4225, CP 11400, Montevideo, Uruguay; Washington Jones [wawijo@yahoo.com.ar], Museo Nacional de Historia Natural, 25 de Mayo 582, CP 11000, Montevideo, Uruguay; Martín Ubilla [ubilla@fcien.edu.uy], Facultad de Ciencias (UdelaR), Iguá 4225, CP 11400, Montevideo, Uruguay; Andrés Rinderknecht [apaleorinder@yahoo.com], Museo Nacional de Historia Natural, 25 de Mayo 582, CP 11000, Montevideo, Uruguay.     

Late Cambrian (Iverian,Jiangshanian) fossils from the Professor Range area,Western Tasmania

Abstract

Late Cambrian (Iverian, Jiangshanian) agnostoids and trilobites are described from two localities in the Professor Range area of western Tasmania. The fossils occur within a thick folded flysch succession of siltstone, mudstone, quartzwacke sandstone and siliceous conglomerate that forms the lower part of the Owen Group correlatives in this area. The flysch succession has a conformable and probable gradational contact with the underlying Mt Read Volcanics and is abruptly overlain by the siliceous pebble-cobble conglomerate correlated with the Middle Owen Conglomerate of the West Coast Range. The stratigraphically lower southern locality contains the agnostoids Pseudagnostus sp. and Agnostinae gen. et sp. indet. plus the polymerid trilobites Parabolina sp., Hedinaspis sp. cf. H. regalis and Eugonocare sp. The northern locality contains the agnostoids Rhaptagnostus sp. and Pseudagnostus sp. plus the polymerid trilobites Hedinaspis sp. cf. H. regalis, Ketyna? sp. 1, Ketyna? sp. 2, and Cermatops sp. It is possible that most specimens of the cosmopolitan genus Hedinaspis belong in the type species, H. regalis.

Christopher J. Bentley [bigfossil@bigpond.com], 30 Albert Street, Clare SA 5453; James B. Jago [jim.jago@unisa.edu.au], University of South Australia, School of Natural and Built Environment, Mawson Lakes, SA 5095; Keith D. Corbett [keith.corbett@bigpond.com], 35 Pillinger Drive, Fern Tree Tas 7054, Australia.     

The Triassic Mesophlebiidae,a little closer to the crown of the Odonata (Insecta) than other ‘triassolestids’

Abstract

Two new, subcomplete forewings belonging to the ‘triassolestid assemblage’, a group of Triassic stem-relatives of dragon- and damselflies (Odonata), are described. One, recovered from Australia (Aranbanga Volcanic Group), belongs to Mesophlebia antinodalis Tillyard, 1916, previously documented on the basis of two very incomplete wings. The other, recovered from South Africa (Molteno Formation), is assigned to a new species, Mesophlebia elegans sp. nov. The new data allow a reconsideration of the diagnosis of the genus Mesophlebia Tillyard, 1916 and a re-instatement of the family Mesophlebiidae Tillyard, 1916. Notably, the new specimens possess, near the wing base, a posterior lobe absent in most ‘triassolestid’ genera, but present in crown-Odonata and a number of their stem-relatives. Lobodonata tax. nov. is erected to accommodate odonates possessing this lobe. The nature of the ‘vein-like’ element anteriorly delimiting this lobe is discussed. We submit that it might have been initially composed of an invagination of the posterior wing-margin (‘fibula’), which was later captured by AA, imposing its course on CuP.

Ayla Tierney [ayla.tierney@qm.qld.gov.au], Queensland Museum, PO Box 3300, South Brisbane BC, Queensland 4101, Australia; Isabelle Deregnaucourt [isabelle.deregnaucourt@edu.mnhn.fr], Sorbonne Université, MNHN, CNRS, Centre de recherche sur la paléontologie – Paris (CR2P), 57 rue Cuvier, CP38, F-75005 Paris, France; John M. Anderson [jmanderson.gondwana@gmail.com], Evolutionary Studies Institute, Witwatersrand University, 1 Jan Smuts Ave., Braamfontein, Johannesburg 2000, South Africa; Paul Tierney [paul.tierney@qm.qld.gov.au], Queensland Museum, PO Box 3300, South Brisbane BC, Queensland 4101, Australia; Torsten Wappler [torsten.wappler@hlmd.de], Hessisches Landesmuseum Darmstadt, Friedensplatz 1, 64283 Darmstadt, Germany; Olivier Béthoux [obethoux@mnhn.fr], Sorbonne Université, MNHN, CNRS, Centre de recherche sur la paléontologie – Paris (CR2P), 57 rue Cuvier, CP38, F-75005 Paris, France.     

Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough,Tasmania

Jago, J.B., Bentley, C.J., Laurie, J.R. &; Corbett, K.B., 26 June 2018. Some middle and late Cambrian trilobites and brachiopods from the Adamsfield Trough, Tasmania. Alcheringa 43, 1-17. ISSN 0311-5518.

Cambrian Series 3 and Furongian trilobites and brachiopods are described from the Adamsfield Trough in southwestern Tasmania. The oldest fossils are very poorly preserved trilobites, assigned to Asaphiscidae gen. et sp. indet. from within the Island Road Formation a short distance above the unconformity with the underlying Proterozoic Wedge River Beds. A trilobite species from within the isolated Boyd River Formation is referred to Lioparia sp. The Island Road Formation and the Boyd River Formation are stratigraphically equivalent to the Trial Ridge Beds which have previously been dated as belonging to the Lejopyge laevigata Zone. The Trial Ridge Beds are overlain unconformably by the Singing Creek Formation. In the Adamsfield, Clear Hill and Stepped Hills areas, stratigraphic equivalents of the Singing Creek Formation collectively contain the trilobites Pseudaphelaspis sp., Pseudaphelaspis? sp., Prochuangia sp., Mindycrusta sp., Nepeidae gen. et sp. indet., and Olenidae gen. et sp. indet. plus the brachiopods described herein as Billingsella sp. aff. costata, Billingsella sp. A, Billingsella sp. B and a possible member of the Billingselloidea. The Singing Creek Formation has been previously correlated with the Stigmatoa diloma Zone. The genus Lotosoides Shergold 1975 is placed in synonymy with Prochuangia Kobayashi 1935.

James B. Jago* [jim.jago@unisa.edu.au] University of South Australia, School of Natural and Built Environment, Mawson Lakes, SA 5095, Australia; Christopher J. Bentley [bigfossil@bigpond.com] 30 Albert Street, Clare, SA 5453, Australia; John R. Laurie [john.r.laurie@gmail.com] Department of Biological Sciences, Macquarie University, NSW 2109, Australia; Keith D. Corbett [keith.corbett@bigpond.com] 35 Pillinger Drive, Fern Tree, Tas 7054, Australia.     

First long-proboscid flies (Diptera: Zhangsolvidae) from the Lower Cretaceous of South Korea

Khramov, A.V., Nam, G.-S. & Vasilenko, D.V. 13 December 2019. First long-proboscid flies (Diptera: Zhangsolvidae) from the Lower Cretaceous of South Korea. Alcheringa 44, 160–168. ISSN 0311-5518

Buccinatormyia gangnami sp. nov. (Zhangsolvidae) is the first record of Diptera from the Lower Cretaceous Jinju Formation of the Korean Peninsula. B. gangnami is described based on six specimens, the largest number of fossils known for a single species of Zhangsolvidae. Four specimens of B. gangnami preserve a long proboscis; mouthparts of this species show a twofold variation in relative length, which may be partially explained by sexual dimorphism. B. gangnami is the third known species of Zhangsolvidae with extremely elongate antennal flagella. The preserved abdominal colouration pattern in these fossils suggests that B. gangnami was probably a wasp mimic.

Alexander V. Khramov [a-hramov@yandex.ru], Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, 117997 Moscow, Russian Federation; Gi-Soo Nam [nks33@naver.com], Gongju National University of Education, 27, Ungjin-ro, Gongju-si, Chungcheongnam-do, 32553, South Korea; Dmitry V. Vasilenko* [damageplant@mail.ru], Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, 117997 Moscow, Russian Federation. *Also affiliated with: Cherepovets State University, Lunacharskogo 5, 162600 Cherepovets, Russian Federation.     

Exceptionally well-preserved dragonflies (Insecta: Odonata) in Mexican amber

Zheng, D., Nel, A., Jarzembowski, E.A., Chang, S.-C., Zhang, H. & Wang, B., May 2018. Exceptionally well-preserved dragonflies (Insecta: Odonata) in Mexican amber. Alcheringa xxx, xxx–xxx.

Dragonflies (odonatans) are comparatively rare as amber inclusions, and most are not well preserved on account of their size. Here, we report a single piece of Mexican amber with one complete dragonfly and two damselflies. The dragonfly is attributed to the extant gomphid Erpetogomphus Selys Longchamps, and the damselflies belong to the extant coenagrionid Argia Rambur. Both genera are nowadays distributed widely in Mexico. The new discovery dates the origins of these two genera to the Miocene at least.

Daran Zheng [dranzheng@gmail.com], Edmund A. Jarzembowski* [jarzembowski2@live.co.uk] Haichun Zhang [hczhang@nigpas.ac.cn] and Bo Wang? [bowang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; André Nel [anel@mnhn.fr] Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France; Daran Zheng, Su-Chin Chang [suchin@hku.hk] Department of Earth Sciences, The University of Hong Kong, Hong Kong Special Administrative Region, PR China. *Also affiliated with Department of Earth Sciences, The Natural History Museum, London SW7 5BD, UK. ?Also affiliated with Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, PR China. Received 23.1.2018; revised 6.3.2018; accepted 20.3.2018.     

Oldest record of the scale-leaved clade of Podocarpaceae,early Paleocene of Patagonia,Argentina

A new podocarpaceous conifer is described from the early Danian Salamanca Formation (southern Chubut, Patagonia, Argentina) based on compressions of leafy branches with cuticular remains. Kirketapel salamanquensis gen. et sp. nov. has amphistomatic, scale-like leaves with marginal frills distinguishable at the apex; stomata oriented randomly in relation to the major axis of the leaf with four to five subsidiary cells and extremely reduced Florin rings; and irregularly shaped epidermal cells. We compare K. salamanquensis with extant and extinct members of the imbricate-leaved podocarps, among which it closely resembles Florin’s Dacrydium group C genera (i.e., Lagarostrobos, Manoao, Lepidothamnus and Halocarpus). Among these genera, only Lepidothamnus has a living representative in South America, the Chilean L. fonkii, whose leaf macro- and micromorphological characters are described in detail for comparison. Overall, the Patagonian fossil species is most similar to the extant and extinct members of Lagarostrobos in its cuticular micromorphology; however, macromorphological characters, such as the leaf size, apex curvature and mode of flattening, clearly differentiate it from all four genera of Dacrydium group C. We include Kirketapel salamanquensis in a combined molecular and morphological phylogenetic analysis conducted under the maximum parsimony criterion. The new, early Paleocene fossil taxon is confidently recovered as part of the scale-leaved clade as defined herein, which also includes Halocarpus, Phyllocladus, Lepidothamnus, Parasitaxus, Lagarostrobos and Manoao, and it constitutes the oldest record known for the group by at least 17 million years as well as its first fossil occurrence outside Australasia, establishing a widespread Gondwanan history. Furthermore, based on its oldest locality of occurrence, K. salamanquensis shows that the divergence of the total group of the scale-leaved podocarps occurred by at least 65 million years ago, adding to the growing systematic knowledge of earliest Cenozoic macrofloras in the Southern Hemisphere.

Ana Andruchow-Colombo [aandruchow@mef.org.ar] and Ignacio Escapa [iescapa@mef.org.ar] CONICET, Museo Paleontológico Egidio Feruglio Av. Fontana 140, Trelew 9100, Chubut, Argentina; Raymond J. Carpenter* [raymond.carpenter@adelaide.edu.au] School of Biological Sciences, University of Tasmania, Private Bag 55, Hobart, Tas. 7001, Australia; Robert S. Hill [bob.hill@adelaide.edu.au] School of Biological Sciences and Environment Institute, University of Adelaide, Adelaide, SA 5005, Australia; Ari Iglesias [ari_iglesias@yahoo.com.ar] CONICET, Instituto de Investigaciones en Biodiversidad y Medioambiente INIBIOMA-CONICET, Universidad Nacional del COMAHUE, Quintral 1250, San Carlos de Bariloche 8400, Río Negro, Argentina; Ana Abarzua [anaabarzua@uach.cl] Instituto de Ciencias de la Tierra, Facultad de Ciencias, Universidad Austral de Chile, Av. Rector Eduardo Morales Miranda 23, Valdivia 5090000, Región de los Ríos, Chile; Peter Wilf [pwilf@psu.edu] Department of Geosciences, Pennsylvania State University, University Park, PA 16802, USA. *Also affiliated with: School of Biological Sciences, University of Adelaide, Adelaide, SA 5005, Australia. Received 28.2.2018; revised 22.8.2018; accepted 24.8.2018.     

Pliocene corals from the Togopi Formation of the Dent Peninsula,Sabah, northeastern Borneo,Malaysia

Saw, J.V.M., Hunter, A.W., Johnson, K.G. & Abdul Rahman, A.H.B., November 2018. Pliocene corals from the Togopi Formation of the Dent Peninsula, Sabah, northeastern Borneo, Malaysia. Alcheringa 43, 291–319. ISSN 0311-5518

The palaeobiology of the Malay Archipelago region remains poorly documented, despite its present-day significance as a modern global marine biodiversity hotspot. The Togopi Formation of the Dent Peninsula, situated in Borneo on the western Sulu Sea and eastern coast of Sabah, Malaysia, preserves Plio-Pleistocene sedimentary sequences interpreted to represent localized transgressive episodes, and which have a relatively high coral diversity. Fossil corals were sampled from three available quarries on the Dent Peninsula, the sediments of which have been previously dated as 4.5–3.4?Ma in age based on foraminiferal data and radiometric analyses. These Pliocene corals are identified here based on their macromorphology, micromorphology and microstructural characteristics. In total, this study describes 28 fossil coral taxa, with 16 genera recognized and 22 taxa identified to species level, 21 of which can be confidently assigned to extant species. These new data have resulted in revised stratigraphic ranges for eight of these species. As the most comprehensive systematic survey of corals from the Pliocene of the Indo-Pacific to date, this study indicates a high diversity of corals on the margin of the Sabah Sea, Borneo, at this time, including taxa found today, thus casting doubt on the local impact of the Plio-Pleistocene extinction previously reported from faunal analyses of the central Indo-Pacific.

Jasmin V.M. Saw [varnmay@yahoo.com] Department of Petroleum Geosciences, Universiti Teknologi PETRONAS, 32610 Seri Iskandar, Perak, Malaysia; Aaron W. Hunter* [awh31@cam.ac.uk] Department of Earth Sciences, University of Cambridge, Downing Street, Cambridge CB2 3EQ, UK; Kenneth G. Johnson [k.johnson@nhm.ac.uk] Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK; Abdul Hadi B Abdul Rahman [hadi_rahman@utp.edu.my] Department of Petroleum Geosciences, Universiti Teknologi PETRONAS, 32610 Seri Iskandar, Perak, Malaysia. *Also affiliated with: School of Earth Sciences, The University of Western Australia, 35 Stirling Highway, Crawley, WA 6009, Australia.     

First record of the ichthyodectiform fish Xiphactinus (Teleostei) from Patagonia,Argentina

Abstract

We report the first record of the genus Xiphactinus from southern South America. The recovered fossil material consists of an associated maxilla and abdominal vertebra, probably derived from latest Maastrichtian marine deposits of the Salamanca Formation in Chubut Province, Argentina. Xiphactinus has been widely reported from Late Cretaceous strata throughout the Northern Hemisphere, although to date, equivalent Southern Hemisphere occurrences include only a single specimen from Venezuela. Our new discovery thus indicates that Xiphactinus had a much more cosmopolitan distribution, encompassing the southern paleolatitudinal extremities of South America during the terminal Cretaceous.

Julieta J. De Pasqua [julietadepasqua@gmail.com], Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Ángel Gallardo, 470, C1405DJR, Buenos Aires, Argentina;

Federico L. Agnolin [fedeagnolin@yahoo.com.ar], Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Ángel Gallardo, 470, C1405DJR, Buenos Aires, Argentina. CONICET; Fundación de Historia Natural ‘Félix de Azara’, Departamento de Ciencias Naturales y Antropología, Universidad Maimónides; Hidalgo 775 piso 7, C1405BDB, Buenos Aires, Argentina;

Sergio Bogan [sergiobogan@yahoo.com.ar], Fundación de Historia Natural ‘Félix de Azara’, Departamento de Ciencias Naturales y Antropología, Universidad Maimónides; Hidalgo 775 piso 7, C1405BDB, Buenos Aires, Argentina.     

The first Chinese representative of the Jurassic damsel-dragonfly genus Hypsothemis (Odonata: Isophlebioidea: Campterophlebiidae)

HUANG, D., FU, Y. & NEL, A. 4 October 2019. The first Chinese representative of the Jurassic damseldragonfly genus Hypsothemis (Odonata: Isophlebioidea: Campterophlebiidae). Alcheringa 44, 99–103. ISSN 0311–5518

A new campterophlebiid damsel-dragonfly, Hypsothemis sinensis sp. nov., is described from the lowermost Upper Jurassic Haifanggou Formation at the Daohugou locality in the Ningcheng Basin, China. This is the first Chinese representative of this genus, previously known only from the coeval upper Karabastau Formation in Kazakhstan, reflecting strong palaeobiogeographic links between these two entomofaunas.

Diying Huang [dyhuang@nigpas.ac.cn] and Yanzhe Fu [yzfu@nigpas.ac.cn], State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, PR China; André Nel [anel@mnhn.fr], Institut de Systématique, Evolution, Biodiversité, ISYEB, UMR 7205, CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, Université des Antilles, 57, rue Cuvier, CP 50, Entomologie 75005 Paris, France.     

Morphology,palaeoecology and taphonomy of the Devonian mitrate Placocystella langei from Paraná Basin,Brazil

Sandro Marcelo Scheffler, Rodrigo Scalise, Horodyski & Elvio Pinto, Bosetti, 2018. Morphology, palaeoecology and taphonomy of the Devonian mitrate Placocystella langei from Paraná Basin, Brazil. Alcheringa 43, 228–240. ISSN 0311-5518.

The original holotype and paratypes of the stylophoran mitrate Placocystella langei (Echinodermata) are definitively lost. A neotype is designated here, based on new material collected in the marine Devonian strata of the Apucarana Sub-basin, Paraná, Brazil. Placocystella langei was the first species of Stylophora to be identified in Devonian basins from Brazil. Its range extends from the late Emsian to early Givetian. Taphonomic data indicate that P. langei specimens were preserved in a storm-influenced, transitional offshore environment, and that skeletal remains were few exposed at the water–sediment surface. Considering that the specimens do not exhibit evidence of reworking, it is possible that their final burial was rapid and definitive, which facilitated their almost complete preservation. Finally, taphonomic, diagenetic and sedimentologic data all suggest a shallow infaunal habit for Placocystella langei.

Sandro Marcelo Scheffler [schefflersm@mn.ufrj.br] Departamento de Geologia e Paleontologia, Laborat orio de Paleoinvertebrados (LAPIN), Museu Nacional—UFRJ, Parque Quinta da Boa Vista, s/n, São Cristóvão 20940040, Rio de Janeiro, RJ, Brasil; Rodrigo Scalise Horodyski* [rhorodyski@unisinos.br] Graduate Program in Geology, Vale do Rio dos Sinos University, Av. Unisinos, 950, Cristo Rei, 93022-000 São Leopoldo, RS, Brasil; Elvio Pinto Bosetti [elviobosetti@gmail.com] Departamento de Geociências, Universidade Estadual de Ponta Grossa, Av. Carlos Cavalcanti 9.500, Uvaranas, 84010-919, Ponta Grossa, Paran a, Brazil.     

Middle Ordovician (Darriwilian) conodonts from the Goldwyer Formation of the Canning Basin,Western Australia

Zhen, Y.Y., Normore, L.S., Dent, L.M. & Percival, I.G., 11 July 2019. Middle Ordovician (Darriwilian) conodonts from the Goldwyer Formation of the Canning Basin, Western Australia. Alcheringa 44, 25–55. ISSN 0311-5518

Middle Ordovician conodonts attributed to 46 species were recovered from a stratigraphic interval spanning the Willara, Goldwyer and Nita formations in core sections from the Sally May-2 and Theia-1 petroleum exploration wells in the Canning Basin, Western Australia. The Histiodella serrata, Histiodella holodentata and Eoplacognathus pseudoplanus biozones are recognized in the lower and middle part of the Goldwyer Formation, indicative of an early–middle Darriwilian age. This revised conodont biostratigraphy enables more precise correlation with North America and North and South China. Several biogeographically distinctive conodont species, most likely of North Chinese origin, are recorded from the Goldwyer Formation. Their presence signals a strong palaeobiogeographic connection between the Sino-Korean Craton and the Canning Basin on the western margin of eastern Gondwana during the late Middle Ordovician.

Y.Y. Zhen* [yong-yi.zhen@planning.nsw.gov.au], W.B. Clarke Geoscience Centre, Geological Survey of New South Wales, 947–953 Londonderry Road, Londonderry NSW 2753, Australia; L.S. Normore [leon.normore@dmirs.wa.gov.au]; L.M. Dent [louisa.dent@dmirs.wa.gov.au], Department of Mines, Industry Regulation and Safety, Mineral House, Geological Survey of Western Australia, 100 Plain Street, East Perth, WA 6004, Australia; I.G. Percival [ian.percival@planning.nsw.gov.au], W.B. Clarke Geoscience Centre, Geological Survey of New South Wales, 947–953 Londonderry Road, Londonderry NSW 2753, Australia;     

Two new skeletons of the enigmatic,rail-like avian taxon Songzia Hou, 1990 (Songziidae) from the early Eocene of China

Sachs, S. &; Kear, B.P. November 2018. A rare new Pliensbachian plesiosaurian from the Amaltheenton Formation of Bielefeld in northwestern Germany. Alcheringa 42, 487-500. ISSN 0311-5518.

We describe a new plesiosaurian from the upper Pliensbachian Amaltheenton Formation of Bielefeld in northwestern Germany. The taxon is based upon an incomplete associated skeleton comprising part of the right mandibular ramus, several teeth, a series of cervical, pectoral, dorsal and caudal vertebrae, as well as ribs, limb girdle elements including a nearly complete right scapula, and various distal limb bones. A unique character state combination serves to distinguish the Amaltheenton Formation remains from other previously documented Early Jurassic plesiosaurians. The most important features are the presence of a longitudinal notch incising the posterior rim of the glenoid fossa and retroarticular process, and a pronounced ventrolateral shelf on the scapula, both of which constitute derived states otherwise shared with Early Cretaceous leptocleidians. However, phylogenetic analysis using a ‘total group’ Plesiosauria data-set that specifically accommodates for Pliensbachian taxa unanimously placed the Amaltheenton Formation plesiosaurian among Early–Middle Jurassic pliosaurids. This discovery is significant because it reveals unexpected homoplasy, but also because it establishes what is only the third formally named plesiosaurian taxon thus far documented from Pliensbachian strata worldwide.

Sven Sachs^* [sachs.pal@gmail.com], Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany and Im Hof 9, 51766 Engelskirchen, Germany; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden.     

A new,extraordinary ‘damselfly-like’ Odonatoptera from the Pennsylvanian of the Avion locality in Pas-de-Calais,France (Insecta: ‘Exopterygota’)

Nel, A., Roques, P., Prokop, J., & Garrouste, R., 11 September 2018. A new, extraordinary ‘damselfly-like’ Odonatoptera from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: ‘Exopterygota’). Alcheringa 43, 241–245. ISSN 0311-5518.

Enigmaptera magnifica gen. et sp. nov., type genus and species of the new odonatopteran family Enigmapteridae, is described from the Moscovian of Avion (northern France). It is the sister group of the major clade Neodonatoptera, placed together in the new clade Paneodonatoptera. Its wing venation has characters never found in other Odonatoptera. It is a further case of convergent wing petiolation in this superorder. Enigmaptera magnifica, like the protozygopteran Jacquesoudardia magnifica from the same outcrop, probably lived like the extant damselflies along the shores of lakes and rivers, hunting the small insects found in the same deposits. These discoveries show that very small insects were significant elements of the entomofaunal diversity and trophic chains of the Late Carboniferous ecosystems.

Romain Garrouste [garroust@mnhn.fr] Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, 57 rue Cuvier, CP 50, 75005 Paris, France; Patrick Roques [patrick.roques93@wanadoo.fr] Allée des Myosotis, Neuilly sur Marne, F-93330, France.; Jakub Prokop [jprokop@natur.cuni.cz] Charles University, Faculty of Science, Department of Zoology, Vini?ná 7, CZ-128 44, Praha 2, Czech Republic.     

Holocene Ostracods from the Hang Hau Formation in Lei Yue Mun,Hong Kong,and their palaeoenvironmental implications

Wang. H., Zhang, H., Cao, M. & Horne, D.J., October 2018. Holocene Ostracods from the Hang Hau Formation in Lei Yue Mun, Hong Kong, and their palaeoenvironmental implications. Alcheringa 43, 320–333. ISSN 0311-5518.

The Holocene Hang Hau Formation is the youngest part of the Quaternary succession in Hong Kong and yields abundant and diverse ostracods. This study of ostracod assemblages from two cores in Lei Yue Mun identifies eight genera and nine species of marine Ostracoda that were previously unreported from the Hang Hau Formation, increasing the known diversity from 67 to 76 species. Among these species, Neocyprideis timorensis (Fyan 1916) is reported for the first time in China. The recovery of abundant juvenile and adult specimens has facilitated illustration and discussion of an ontogenetic series for Neomonoceratina delicata Ishizaki & Kato, 1976, extending from the A-5 instar (fourth moult after hatching) to the sexually dimorphic A (adult) instar. Palaeoenvironmental analysis of the ostracod assemblages supports and strengthens previous interpretations indicative of a warm, shallow, nearshore-marine environment.

He Wang [wang0701@163.com] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Haichun Zhang [hczhang@nigpas.ac.cn] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; Meizhen Cao [mzcao@nigpas.ac.cn] Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, PR China; David J. Horne [d.j.horne@qmul.ac.uk] School of Geography, Queen Mary University of London, Mile End Road, London E1 4NS, UK. *Also affiliated with: University of Science and Technology of China, No. 96, JinZhai Road Baohe District, Hefei, Anhui 230026, PR China.